Goniographa naumanni, Varga & Ronkay, 2002

Varga, Z. & Ronkay, L., 2002, A Revision Of The Palaearctic Species Of The Eugraphe Hübner, [1821] 1816 Generic Complex. Part I. The Genera Eugraphe And Goniographa (Lepidoptera, Noctuidae), Acta Zoologica Academiae Scientiarum Hungaricae 48 (4), pp. 333-374 : 352-357

Goniographa naumanni sp. n.

( Figs 9–11 , 34, 35 , 60, 61 )

Holotype: male, “ Afghanistan, Badakhshan, Darrah-e-Kuf , 2480 m, 16.07.1972., leg. BRADE & NAUMANN ”; slide No. VZ4634 (coll. Z. VARGA).

Paratypes. Afghanistan: 1 male, 1 female, Prov. Kadaghan, Salang-Pass N-Seite, 2400 m, 11–12. VII.1971. leg. VARTIAN ; 1 male, Khurd-Kabul , 5.VII.1963. leg. KASY & VARTIAN , 1 male, Dasht-i- Nawar, Hokak , 2850 m, 7–9.IX.1963. leg. VARTIAN (coll. Z. VARGA) ; 1 male, Badakhshan, Darwaz, Shewa plateau, Darrah-e-Gulistan , 2800 m, 21.VIII.1973 , 1 male, Badakhshan, Darwaz, Shewa valley, Basindj , 1900–2000 m, 27.VIII.1973., leg. Naumann & Nauruz. (coll. NAUMANN) . Tadjikistan: 1 male, Pamir Mts, Shugnan , VII.1935, leg. WEIDINGER (coll. G. RONKAY) ; 9 males, 12 females, Pamir Mts, Chorog , July–August 1963 –65, leg. SHCHETKIN ; 1 female, from the same locality, 11.VIII.1992, leg. SHCHETKIN ; 1 female, from the same site, 29–31.VII.1999, leg. SHCHETKIN; 1 female, Pamir Mts, Shod , 2300 m, 20.X.1987, leg. NEKRASOV (coll. P. GYULAI and J. STUMPF) .

Slide Nos RL 7552m, RL 7555m, VZ3634; VZ3635, VZ3685, VZ4640, VZ7032, VZ7153 (males), RL7583f, RL7584f (females).

Diagnosis: The new species is almost equal in size with G. funkei , although the forewings are slightly narrower triangular with less pointed apex. Forewing colouration and pattern of the two species are highly similar, ground colour of G. naumanni is somewhat more ochreous grey, the postmedian line is less evenly curved and crenulate, the claviform stigma is slightly shorter and more obsolescent and the hindwing is a bit more suffused marginally. The satisfactory separation of the two species requires the study of the genitalia. This process is much easier in the females, as no dissection is needed to recognise the differences of the ostial plate.

The male genitalia of G. naumanni differ conspicuously from those of the two twin species by its considerably longer harpe, distally upcurved (“oxbow”-

Acta zool. hung. 48, 2002

Acta zool. hung. 48, 2002

shaped)pollex, the much stronger sclerotised carina bearing 2–3 large, acute teeth and the presence of a huge subbasal diverticulum of the not fully recurved vesica.

Comparing the female genitalia of G. naumanni with those of its two sister species, G. naumanni has conspicuously broader but shorter, trapezoidal, not triangular-infundibular ostium bursae and much stronger, heavily sclerotized, very broad, not tubulat but more or less cask-shaped ductus bursae.

Description. Wingspan 32–37 mm, length of forewing 14–17 mm. The main external features fit well with those of G. funkei , with only a bit more diffuse dark forewing markings. The holotype (male)and a typical female are illustrated in Figs 60 and 61 .

Male genitalia ( Figs 9–11 ). The specific features of the male genitalia are the long, slender harpe, the long, distally arched (“oxbow”-shaped)pollex, the nearly straight aedeagus with two or three large, pyramidal teeth on the ventral edge of the carina, the dorsally upturned but not fully recurved vesica with elliptical subbasal diverticulum and with short and strong submedial bulbed cornutus.

Female genitalia ( Figs 34, 35 ): Ovipositor short, weak; posterior gonapophyses medium-long, slender, fine, anterior apophyses short. Ostium bursae relatively short but broad, trapezoidal, with strongly convex, medially slightly incised caudal margin. Sclerotizations of ostium bursae and ductus bursae almost completely fused with each others. Ductus bursae heavily sclerotized, medium-long, broad, rather cask-shaped, with strongly tapering proximal third. Apical part of bursa copulatrix finely scobinate, partly wrinkled-ribbed, partly gelatinous. Appendix bursae subconical, finely wrinkled and verrucose; corpus bursae elliptical-ovoid, signa absent.

Bionomics and distribution. The new species seems to be confined to the Hissar Mts, the western Pamir Mts (Shugnan range, Chorog)and the north-eastern territories of Afghanistan (Prov. Badakhshan, Kadaghan and Darwaz Mts). In the north-west, there is a narrow overlap between the

Acta zool. hung. 48, 2002

areas of G. funkei and G. naumanni ; the south-eastern boundary of the distribution of the new species is the Paghman Mts near Kabul (Khurd-Kabul).

Etymology. The new species is dedicated to one of its collectors, the prominent entomologist,

Prof.Dr. Clas M. NAUMANN, who has made several important entomological discoveries in Afghanistan.

Goniographa marcida ( CHRISTOPH, 1893) comb. n.

( Figs 12 , 36, 37 , 62, 63 )

Agrotis marcida CHRISTOPH, 1893 , Dt. ent. Z. Iris 6: 90. Type locality: Askhabad ( Turkmenistan: Ashgabat).

Type material examined: 1 male, 1 female syntypes, Turkmenistan, Askhabad ( ZIN St. Petersburg) .

Additional material examined: Turkmenistan: 2 males, 2 females, Kopet-Dagh Mts, Firyuza , 400–600 m, 25.IX.1991, No. L 27, leg. A. PODLUSSÁNY, L. RONKAY & Z. VARGA ; 4 males, 12 females, Kopet-Dagh Mts, Karayalchi , 1600 m, 5.X.1991, No. L 36, leg. A. PODLUSSÁNY, L. RONKAY & Z. VARGA (coll. HNHM Budapest, B. HERCZIG, P. GYULAI, G. RONKAY and Z. VARGA) . Iran: 1 male, 1 female, Prov. Khorassan, Kopet-Dagh Mts , 2300 m, 10 km N Jevenly, 27.VIII.2000, leg. A. GARAI & P. GYULAI ; 5 males, 5 females, Prov. Khorassan, Kopet-Dagh Mts, Jozak NP, 2 km W Jozak, 1350 m, 28.VIII.2000, leg. A. GARAI & P. GYULAI (coll. P. GYULAI) .

Slide Nos RL 7476m, 7030 VZ (males), RL 7477f, RL 7569f, RL 7574f (females).

Acta zool. hung. 48, 2002

Diagnosis: Goniographa marcida differs from its sister species, G. gyulaipeteri by its generally more unicolorous brownish forewings with less intense paler irroration in the median area and by the rather concolorous brown hindwings of both sexes (the inner area of the hindwing is much paler, often prominently whitish in G. gyulaipeteri ).

Comparing the male genitalia of G. marcida and G. gyulaipeteri , the distal part of valva of G. marcida has longer processus, with more arcuate pollex originating rather far from “pseudopollex”, these processi are shorter in G. gyulaipeteri and the “pseudopollex” is situated almost at the “halfway between valval apex and pollex. The harpe of G. marcida is somewhat longer, S-shaped, the juxta is narrower than in its sister species. The configuration of the vesica also shows easily recognisable differences as the vesica of G. marcida is upturned dorsally but not recurved as in G. gyulaipeteri (see Figs 12–14 ), and the cornutus is longer, narrower with smaller basal bulb.

The female genitalia of the two closely related species, G. marcida and G. gyulaipeteri , differ conspicuously by their appendix bursae ( Figs 36–40 )which is membranous-scobinate in G. marcida while G. gyulaipeteri has a large, strongly sclerotized dorso-medial fold. The ostium bursae of G. marcida is broader, shorter, without caudal incision, the proximal part of the ductus bursae is finely curved laterad, having short but strong medio-lateral crest and rounded, verrucose ventral plate while the ductus bursae of G. gyulaipeteri is straight, having more or less parallel margins, without proximo-lateral crest and verrucose plate.

Acta zool. hung. 48, 2002

Description. Wingspan 27–35 mm, length of forewing 12–16 mm. The characterisation of the external features is rather laconic in the original (Latin)description and the measures are somewhat larger (“length of forewing 14–17 mm). The colour pictures of a typical male (from Iran, Khorassan) and a typical female (from the Kopet-Dagh, Turkmenistan)are given in Figs 62 and 63 .

Male genitalia ( Fig. 12 ): Uncus strong, pointed; valva straight, elongate with nearly parallel margins; harpe arcuate and pointed; pollex spine-shaped, slightly curved; juxta shield-shaped, narrower apically; aedeagus strong, dentate ribbon with fine teeth, recurved terminally, vesica broad, saccate, only moderately recurved, with a short, acute bulbed cornutus in submedial position.

Female genitalia ( Figs 36, 37 ): Ovipositor rather short, weak; gonapophyses slender, fine. Ostium bursae sclerotized, its ventral plate broadly triangular-calyculate, caudal margin convex, evenly arcuate, without incision. Ductus bursae, long, broad, flattened, strongly, granulously sclerotized, anterior end with short but strong medio-lateral crest and a large, rounded, verrucose proximo-lateral plate. Ventral surface of ductus bursae with narrow longitudinal medial suture. Apical part of bursa copulatrix membranous, wrinkled-ribbed; appendix bursae large, subconical, with membranous apex and wrinkled-ribbed, scobinate basal two-thirds. Corpus bursae small, elliptical-ovoid, weakly membranous; signa absent.

Bionomics and distribution. Goniographa marcida seems to be confined to the Kopet-Dagh mountain system, both in Turkmenistan and Iran (Khorassan region).

Figures

Fig. 9. Goniographa naumanni sp. n., holotype, slide No. VZ4634

Fig. 10. Goniographa naumanni sp.n., paratypes, slide Nos a = VZ3634, b = VZ4640, c = VZ3635

Fig. 11. Goniographa naumanni sp. n., paratypes, slide Nos a = VZ3634, b = VZ3635

Figs 30–35. 30–31 = Goniographa funkei (PÜNGELER), slide No. RL7554: 30 = female genitalia, 31 = posterior part in larger magnification;32–33 =G. metafunkei sp.n.,holotype, slide No.RL7528: 32 = female genitalia, 33 = posterior part in larger magnification; 34–35 = G. naumanni sp. n.,

Figs 58–65. 58–59 = Goniographa metafunkei sp. n.: 58 = holotype, 59 = paratype male, Kirghisia; 60–61 = G. naumanni sp. n.: 60 = holotype, 61 = paratype female, Afghanistan; 62–63 = G. marcida (CHRISTOPH): 62 = male, Turkmenistan, 63 = female, NE Iran; 64–65 = G. gyulaipeteri sp. n.: 64 = holotype, 65 = paratype male, Tadjikistan

Fig. 12. Goniographa marcida (CHRISTOPH), slide Nos a-b = RL7476, c = VZ7030

Figs 36–42. 36–37 = Goniographa marcida (CHRISTOPH), slide No. RL7477: 36 = female genitalia, 37 = posterior part in larger magnification; 38–40 = G. gyulaipeteri sp. n., paratypes: 38 = female genitalia, slide No. RL7501, 39–40 = posterior part in larger magnification (slide Nos RL7501, RL7502); 41 = Xestia (s. l.) ornata (STAUDINGER), slide No. RL7482, posterior part in larger magni-