Nyctimus Thorell, 1877

Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan & Drescher, Jochen, 2025, Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia, ZooKeys 1255, pp. 95-125 : 95-125

Nyctimus Thorell, 1877

Type species.

Nyctimus bistriatus Thorell, 1877 .

Diagnosis.

Small thomisids, 2–5 mm in body length. Prosoma cuboid, as long as wide, with granulated surface and bearing long macrosetae (Fig. 1 ). Body colouration dark with white / yellow markings, mostly in the form of spots and bars. Opisthosoma with five large sigillae arranged in a triangular formation Embolus is weakly curved and possesses a thick base. Tegulum with a “ hood ” (Fig. 3 A ). Tibial apophyses are well developed, VTA present, either free (Fig. 3 A ) or hook-shaped and attached to RTA (Fig. 7 A ). Epigynum highly variable, spermatheca rounded or oval, single pair (Fig. 3 C, D ).

Species composition.

Nyctimus bistriatus Thorell, 1877 ( ♂ ♀), Nyctimus falcatus Benjamin & Dhiya’ulhaq , sp. nov. ( ♂), Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq , sp. nov. ( ♂ ♀), Nyctimus mutilloides Benjamin & Dhiya’ulhaq , sp. nov. ( ♂), Nyctimus quadripunctatus Dhiya’ulhaq & Benjamin , sp. nov. ( ♂ ♀), Nyctimus rendang Dhiya’ulhaq & Benjamin , sp. nov. ( ♂ ♀), Nyctimus saksang Dhiya’ulhaq & Benjamin , sp. nov. ( ♂), Nyctimus trimeni ( Simon, 1895) .

Figures

Figure 1. Nyctimus bistriatus Thorell, 1877 from Borneo; habitus, dorsal view. A. Male (RMNH. ARA. 17873); B. Female (RMNH. ARA. 17875). Scale bars: 0.5 mm.

Figure 3. Nyctimus bistriatus Thorell, 1877 from Borneo. A. Male (RMNH. ARA. 17873), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17875), epigynum; C. ibid., ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.

Figure 7. Nyctimus kinabaluensis Benjamin & Dhiya’ulhaq, sp. nov., from Borneo. A. Male (RMNH. ARA. 17859), left palp, ventral view; B. ibid., retrolateral view; C. Female (RMNH. ARA. 17862), epigynum, ventral view; D. ibid., dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; E = embolus; RTA = retrolateral tibial apophysis; S = spermatheca; TH = tegular hood; VTA = ventral tibial apophysis. Scale bars: 0.2 mm.