Ptychidia sarmatica ( Papp, 1954 ) Harzhauser & Landau, 2019

Harzhauser, Mathias & Landau, Bernard, 2019, Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera, Zootaxa 4681 (1), pp. 1-136 : 71-73

Ptychidia sarmatica ( Papp, 1954) new comb.

Figs 6X , 19G, 19H, 19I, 19J

Turritella (Haustator) sarmatica n. sp. — Papp, 1948: 111 [nomen nudum].

* Turritella (Haustator) sarmatica n. sp. — Papp, 1954: 37, pl. 9, figs 20–22.

Turritella (Haustator) sarmatica Papp — Sieber, 1960: 241.

Turritella (Haustator) sarmatica Papp — Schultz, 1998: 130, pl. 59, fig. 5.

Turritella badensis sarmatica— Roetzel, 2003: 399.

Turritella eryna sarmatica Papp — Piller & Harzhauser, 2005: 452.

Turritella eryna sarmatica ( Papp, 1954) — Mandic et al., 2008: 350, fig. 7g.

non Turritella sarmatica Papp, 1954 — Poljak, et al. 2016: 144, pl. 3, fig. 9 [probably an Archimediella ].

Type material. Holotype NHMW 2018/0281/0003, illustrated in Papp (1954, pl. 9, fig. 20), paratype NHMW 2018/0281/0004, illustrated in Papp ( Papp, 1954: pl. 9, figs 21), paratype NHMW 2018/0281/0005, illustrated in Papp ( Papp, 1954: pl. 9, figs 22), 3 paratypes NHMW 2018/0281/0006; all Sarmatian, middle Miocene, Hollabrunn- Aspersdorf ( Austria).

Studied material. Holotype NHMW 2018/0281/0003, 3 paratypes NHMW 2018/0281/0006; 1 spec. NHMW 2018/0281/0001 illustrated in Mandic et al. (2008, fig. 7g), 1 spec. NHMW 2018/0281/0002, 1 spec. NHMW 2013/0284/0014; all Hollabrunn-Aspersdorf ( Austria).

Illustrated material. Fig.19G : holotype NHMW 2018/0281/0003, SL: 25.2 mm, MD: 7.7 mm, illustrated in Papp (1954, pl. 9, fig. 20), Fig. 19H : NHMW 2018/0281/0001, SL: 20.9 mm, MD: 7.4 mm, Fig. 19I : NHMW 2013/0284/0014, SL: 7.7 mm, MD: 3.0 mm, Fig. 19J : NHMW 2018/0281/0002, SL: 17.7 mm, MD: 7.2 mm.

Description. Shell small, slender to moderately slender. Protoconch and early teleoconch whorls unknown. First preserved neanic whorls high, convex, tricostate with incised suture. B spiral cord forming weak mid-whorl angulation, shifting to lower half of whorl during ontogeny. A, B and C spiral cords remain weak throughout ontogeny, being separated by broad, flat interspaces, each with one weak secondary spiral cord. Two secondary spiral cords appear on sutural ramp, usually adjoined by tertiary spiral threads. Another spiral thread between C spiral cord and faint peribasal cord. Lateral sinus with moderately steep angle; moderately deep, with faint adapical inflection point close to suture and faint abapical inflection point; abapical LS p. Aperture and base not preserved. No information on inner lirae due to preservation.

Shell measurements and ratios. No complete specimen is known; fragments suggest a maximum height of about 40 mm and 7.9 mm width. AA: ~17°, PA; ~14°; lateral sinus (n = 5): LS angle = 19.4° (σ = 2.2°), LS p = 1.2 (σ = 0.1), LS d = 5.2 (σ = 0.9).

Discussion. This rare species, which is the last turritellid in the Miocene of the Central Paratethys, was treated as subspecies of Ptychidia erynella (‘ turris , eryna ’) or P. vindobonensis ( Handmann, 1882) (‘ badensis ’), but it differs from both species in its smaller size and the higher and less convex early spire whorls. Its spiral sculpture is less prominent and the lateral sinus angle is less steep. The morphology of the lateral sinus is clearly closer to P. vindobonensis , whereas the lateral sinus of P. erynella is deeper. Ptychidia erynella is restricted to the early Badenian of the Paratethys, whereas P. vindobonensis is very frequent during the middle and late Badenian. Thus, it is likely that P. sarmatica represents an offshoot of P. vindobonensis , which survived the Badenian-Sarmatian Extinction Event (Harzhauser & Piller 2007).

The ecological requirements of P. sarmatica , as mud-flat dwelling species, differ from that of P. vindobonensis , which is only known from sublittoral settings. Consequently, the stable isotope signatures differ from those of P. vindobonensis and P. erynella (see Harzhauser et al. 2007), excluding also reworking from Badenian strata.

Paleoenvironment. The locality Hollabrunn-Aspersdorf represents tidal-flat deposits of an incised-valley probably under polyhaline salinities (Harzhauser et al. 2007; Mandic et al. 2008; Gebhardt et al. 2009).

Distribution. Central Paratethys. Early Sarmatian (middle Miocene): North Alpine -Carpathian Foreland Basin: Hollabrunn-Aspersdorf ( Austria) ( Papp 1954). Despite the vast number of Sarmatian outcrops, this species is known so far exclusively from Hollabrunn-Aspersdorf in Austria, where it occurs in the lower Sarmatian Mohrensternia Zone ( Mandic et al. 2008).

Figures

FIGURE 6. Growth line traces of Turritellidae species based on own data; bold: type species of genera; black: Paratethyan species; asterisk: non-Paratethyan species (unknown basal sinuses are left blank).

FIGURE 19. A–F. Ptychidia partschi (Rolle, 1856). A1–A2. Pöls (Austria), NHMW 1861/0001/0270. B1–B2. Pöls (Austria), NHMW 2018/0275/0001, C1–C2. Holotype of Turritella (Haustator) partschi quadricarinata Sieber, 1960, Wetzelsdorf- Kreuzschaller (Austria), UMJG&P 60.658. D1–D2. Wetzelsdorf-Grössl (Austria), UMJG&P 60.632. E. Ptychidia partschi (Rolle, 1856), Wetzelsdorf-Wenzelsteffi (Austria), UMJG&P 60.847. F. Pöls (Austria), NHMW 1861/0001/0270. G–J. Ptychidia sarmatica (Papp, 1954), Hollabrunn-Aspersdorf (Austria). G. Holotype, NHMW 2018/0281/0003. H. NHMW 2018/0281/0001. I. NHMW 2013/0284/0014. J. NHMW 2018/0281/0002. K–L. Ptychidia austrorotundata new nom., Maria Dreieichen (Austria). K. KM F/1152. L. NHMW 1963/0555/0000. M–N. Ptychidia ernesti (Handmann, 1882), Steinebrunn (Austria). M1–M2. NHMW 1963/0547/0000. N. Steinebrunn (Austria), NHMW 2018/0194/0005.