Tesnus tenellus, L. Benick, 1916: 244

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.4

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https://treatment.plazi.org/id/0F6987E5-FDEF-A0FC-FF1C-F7A9FDF2F934

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scientific name

Tesnus tenellus
status

 

tenellus L. Benick, 1916: 244 ( Stenus ; Type locality: Usambara).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1965: 493 ( Stenus ; synonym of utzungweanus ).

— Puthz, 1967i: 193 ( Stenus ; synonym of utzungweanus ).

kabetensis Cameron, 1950a: 183 ( Stenus ; subgenus Hypostenus ; Type locality: Kenya Colony: Kabete, 5800 feet).

— Puthz, 1967i: 193 ( Stenus ; synonym of utzungweanus ).

vacillator Sharp, 1876: 362 ( Stenus ; Type locality: Tapajos).

— Fauvel, 1901: 74 ( Stenus ; Colombia; Venezuela; Amazonian region; Brazil).

— Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— L. Benick, 1949b: 566 ( Stenus ; subgenus Tesnus ; characters).

- DISTRIBUTION: Amazonas, Colombia, Venezuela, Brazil.

vacuus Casey, 1884 , see: tahoensis Casey, 1884 .

vadoni Puthz, 1971: 251 ( Stenus ; subgenus Hypostenus ; Type locality: Madagascar: Fampanambo).

— Puthz, 1972: 8 ( Stenus ; characters; Madagascar).

- DISTRIBUTION: Madagascar.

vafellus Erichson, 1839 , see: cautus Erichson, 1839 .

vagans L. Benick, 1938c: 280 ( Stenus ; Type locality: Costa Rica).

— L. Benick, 1939: 635 ( Stenus ; subgenus Hypostenus ; cited as new species; characters;

Costa Rica: Ramal, Parismina Santa Clara).

- DISTRIBUTION: Costa Rica.

valens L. Benick, 1938: 20 ( Stenus ; subgenus Parastenus ; Type locality: Java: G. Tangkoeban Prahoe).

- DISTRIBUTION: Indonesia.

validithorax Petri, 1926 , see: guttula P. W. J. Müller, 1821 .

validus L. Benick, 1915 , see: coarcticollis Eppelsheim, 1890 .

vanderijsti Bernhauer, 1929, see: subopacus Fauvel, 1907 .

variabilis Puthz, 1968 , see: immarginatus Mäklin, 1853 .

variator Gistel, 1857: 23 ( Stenus ; [Note: Nomen dubium]; Type locality: Styriae).

- DISTRIBUTION: Austria.

variipennis Rougemont, 1983c: 35 ( Stenus ; subgenus “ Hypostenus ”; Type locality: Thailand: Chiang Rai Prov., Nam Tok Mae Yao).

— Rougemont, 1987: 708 ( Stenus ; subgenus “ Hypostenus ”; Thailand).

- DISTRIBUTION: Thailand.

varipes Casey, 1884: 187 ( Areus ; Type locality: Detroit , Michigan).

— Fauvel, 1901: 76 ( Stenus ; synonym of capucinus ).

— Bernhauer and Schubert, 1911: 173 ( Stenus ; subgenus Hypostenus ; catalog).

— Scheerpeltz, 1933: 1174 ( Stenus ; synonym of callosus ).

— Puthz, 1967q: 327 ( Stenus ; synonym of callosus ).

— Puthz, 1974d: 165 ( Stenus ; valid species; characters; notes).

— Campbell and Davies, 1991: 110 ( Stenus ; Canada).

— Downie and Arnett, 1996: 494 ( Stenus ; characters; USA).

- DISTRIBUTION: USA, Canada.

varius Puthz, 1982a: 140 ( Stenus ; subgenus Hypostenus ; Type locality: New Guinea:

Star Mountains, Gigabip, 5000 feet).

- DISTRIBUTION: New Guinea.

vasconicus Jarrige, 1963 , see: phyllobates Penecke, 1901 .

vastator Ryvkin, 1987c: 152 ( Stenus ; subgenus Nestus ; Type locality: USSR: Far East, vastus L. Benick, 1925b: 75 ( Stenus ; subspecies of humilis , for species misidentified as phyllobates by L. Benick, 1913a: 206; Type locality: Ostkarpaten: Schulergebirge; Rodnaergebirge; Hatzeger Gebirge; Fogeraser Gebirge; Roteturm-Pass; Karlsburg. Kronstadt; Bihargebirge; Banat [Herkulesbad]; Kroatien, Mons Zejac).

— L. Benick, 1913a: 206 ( Stenus ; [Note: Misidentification: Scheerpeltz, 1933: 1168, cited phyllobates Benick as a synonym of vastus ; Benick attributed the species to Penecke]; characters; Siebenbürgen).

— L. Benick, 1929a: 40 ( Stenus ; subgenus Nestus ; characters in key).

— Scheerpeltz, 1933: 1168 ( Stenus ; subgenus Stenus ; catalog).

— Szujecki, 1961: 30 ( Stenus ; subgenus Nestus ; characters; Poland).

— Lohse, 1964: 116 ( Stenus ; subgenus Nestus ; characters; central Europe).

— Smetana, 1964a: 50 ( Stenus ; subgenus Nestus ; characters).

— Puthz, 1967a: 19 ( Stenus ; synonymic notes).

— Szujecki, 1969a: 251 ( Stenus ; notes; Poland).

— Puthz, 1971v: 92 ( Stenus ; notes).

— Puthz, 1972s: 253 ( Stenus ; Yugoslavia).

— Lucht, 1987: 97 ( Stenus ; checklist; Poland; Czechoslovakia; Austria).

— Lohse and Lucht, 1989: 141, 143 ( Stenus ; subspecies of callidus ; Austria; Czech Republic).

— Ryvkin, 1990: 164 ( Stenus ; valid species; notes; Russia; Georgia).

— Hromádka, 1993: 44 ( Stenus ; subgenus Nestus ; Slovakia).

- DISTRIBUTION: Europe, Russia, Georgia.

vastus Bernhauer, 1934 , see: trepidus C. Waterhouse, 1877 .

vegetus Puthz, 1975: 193 ( Stenus ; subgenus Hypostenus ; Type locality: Bhutan: Wangidphodrang).

— Puthz, 1978: 127 ( Stenus ; India).

— Puthz, 1998: 393 ( Stenus ; subgenus Hypostenus ; Vietnam).

- DISTRIBUTION: India, Bhutan, Vietnam.

velebiticus L. Benick, 1915: 230 ( Stenus ; Type locality: Velebit, Ostaria).

— L. Benick, 1929a: 78 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1198 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1972s: 272 ( Stenus ; notes).

- DISTRIBUTION: Croatia.

velleris Ryvkin, 1990: 207 ( Stenus ; subgenus Parastenus ; Type locality: Abzhaziia, oz. Ritsa, 950–1100 m. Also cited from Russia).

- DISTRIBUTION: Russia, Georgia.

velocipes Fauvel, 1886b: 146 ( Stenus ; Type locality: Philippines).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1929: 255 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1969: 29 ( Stenus ; subgenus Hypostenus ; characters).

— Rougemont, 1984: 239 ( Stenus ; Sulawesi; Bali).

- DISTRIBUTION: Philippines, Indonesia.

velox Sharp, 1889: 333 ( Stenus ; Type locality: Nikko ; Tokio; Fukushima).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1968b: 45 ( Stenus ; characters).

— Hromádka, 979b: 66 ( Stenus ; aedeagus; note).

— Watanabe, 1995b: 250 ( Stenus ; Oki-no-shima).

- DISTRIBUTION: Japan.

venator Fauvel, 1895: 208 ( Stenus ; Type locality: Bírmanie, Carin Cheba , 900–1100 m; Carin Ghécu, 1300–1400 m).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Cameron, 1930: 328 ( Stenus ; characters; Burma).

— Scheerpeltz, 1933: 1168 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1980a: 37 ( Stenus ; characters).

— Rougemont, 1981a: 332 ( Stenus ; notes; Burma).

— Rougemont, 1983c: 22 ( Stenus ; Thailand).

— Puthz, 1998: 392 ( Stenus ; subgenus Stenus + Nestus ; Vietnam).

- DISTRIBUTION: Myanmar, Thailand, Vietnam.

veneriatus Normand, 1937 , see: capitatus Eppelsheim, 1878 .

venezuelanus Hromádka, 1983a: 131 ( Stenus ; subgenus Tesnus ; Type locality: Venezuela: Carretera, El Junquito Tover, D.F., 2200 m).

- DISTRIBUTION: Venezuela.

ventralis Sharp, 1876: 368 ( Stenus ; Type locality: Tapajos).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974f: 164 ( Stenus ; characters; Bolivia).

- DISTRIBUTION: Brazil, Bolivia.

ventricosus Fauvel, 1904 , see: bivulneratus Motschulsky, 1857 .

ventridens Puthz, 1987b: 404 ( Stenus ; subgenus Stenus ; Type locality: Kolumbien: 20 km E Silvia, Cauca, 11,000 feet).

- DISTRIBUTION: Colombia.

venustus Casey, 1884: 89 ( Stenus ; Type locality: Not cited).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Notman, 1920: 17 ( Stenus ; New York).

— Fall, 1926a: 61 ( Stenus ; Alaska).

— Scheerpeltz, 1933: 1168 ( Stenus ; subgenus Stenus ; catalog).

— Sanderson, 1957: 259 ( Stenus ; subgenus Stenus ; characters; British Columbia; Washington; Oregon).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

- DISTRIBUTION: Canada, USA.

veracruzmontis Puthz, 2000b: 113 ( Stenus ; subgenus Hemistenus ; Type locality: Mexico: Veracruz: 3. 2 km SW Las Vigas, Hwy 140, 2830 m).

- DISTRIBUTION: Mexico.

verecundus Sharp, 1874 , see: melanarius Stephens, 1833 .

vermicularis L. Benick, 1917b: 295 ( Stenus ; subgenus Stenus ; Type locality: Columbien: Las Tibayes, Terra templ.; [Note: See lectotype designation by Puthz, 1968]).

— Scheerpeltz, 1933: 1168 ( Stenus ; subgenus Stenus ; catalog).

— L. Benick, 1938c: 260 ( Stenus ; characters).

— Puthz, 1968: 1, 6 ( Stenus ; subgenus Stenus ; characters; lectotype designation).

— Puthz, 1995e: 202 ( Stenus ; notes; Ecuador; Colombia).

- DISTRIBUTION: Colombia, Ecuador.

quadriguttatus Puthz, 1971b: 2 ( Stenus ; subgenus Stenus ; Type locality: Colombia: Cundinamarca, road Facatativá–Anolaima, 2600 m).

— Puthz, 1995e: 202 ( Stenus ; synonym of vermicularis ).

vernaculus L. Benick, 1929, see: kuennemanni L. Benick, 1917 .

vernalis Naomi, 1997e: 750 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Shizuoka Pref.: Shimogamo, Minamiizu-cho).

- DISTRIBUTION: Japan.

versicolor L. Benick, 1929 , see: Dianous .

verticalis L. Benick, 1938: 27 ( Stenus ; subgenus Hypostenus ; Type locality: Java: G. Tongkoeban Prahoe, 4000–5000 Voet, Preanger).

— Rougemont, 1981a: 333 ( Stenus ; Burma; China; Vietnam).

— Puthz, 1981a: 12 ( Stenus ; notes; Vietnam; China).

— Rougemont, 1984a: 352 ( Stenus ; China).

veselovae Ryvkin, 1987b: 273 ( Stenus ; subgenus Parastenus ; Type locality: USSR: S-Jamal, Sr.-Ščučje).

— Puthz, 1993c: 146 ( Stenus ; synonym of noctivagus ).

— Naomi, 1997a: 22 ( Stenus ; synonym of noctivagus ).

— Ryabukhin, 1999: 50 ( Stenus ; subgenus Parastenus ; cited as valid species; catalog; habitat; distribution; northeastern Russia).

— Ryvkin, 2000a: 355 ( Stenus ; subgenus Parastenus ; characters; cited as valid species;

Far Eastern Russia).

- DISTRIBUTION: Russia.

veselyi Rambousek, 1914 , see: erythrocnemus Eppelsheim, 1884 .

vespertinus Casey, 1884: 61 ( Stenus ; Type locality: Tenino, Washington Territory ).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Sanderson, 1957: 266 ( Stenus ; subgenus Stenus ; characters; British Columbia; Washington; Oregon).

— Puthz, 1967l: 48 ( Stenus ; cited as synonym of parallelopipedus ).

— Puthz, 1967l: 48 ( Stenus ; [Note: Misidentified as parallelopipedus according to Puthz, 1974d: 161]; notes).

— Puthz, 1974d: 161 ( Stenus ; notes).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

- DISTRIBUTION: Canada, USA.

incultus Casey, 1884: 71 ( Stenus ; Type locality: Arizona).

— Fall, 1901: 71 ( Stenus ; California).

— Bernhauer and Schubert, 1911: 160 ( Stenus ; catalog).

— L. Benick, 1938c: 266 ( Stenus ; characters).

— Puthz, 1974d: 161 ( Stenus ; synonym of vespertinus ).

luctuosus Casey, 1884: 81 ( Stenus ; Type locality: California; Lake Tahoe , Nevada;

Lake Quesnel, British Columbia).

— Bernhauer and Schubert, 1911: 161 ( Stenus ; catalog).

— L. Benick, 1938c: 264 ( Stenus ; characters).

— Puthz, 1974d: 161 ( Stenus ; synonym of vespertinus ).

vestalis Casey, 1884: 138 ( Stenus ; Type locality: Kern County, California).

— Fall, 1901: 71 ( Stenus ; California).

— Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— L. Benick, 1938c: 267 ( Stenus ; characters).

— Puthz, 1980k: 36 ( Stenus ; notes).

- DISTRIBUTION: USA.

exilis Casey, 1884: 143 ( Stenus ; Type locality: California).

— Bernhauer and Schubert, 1911: 174 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 268 ( Stenus ; characters).

— Puthz, 1980k: 36 ( Stenus ; synonym of vestalis ).

vestigialis Erichson, 1840: 732 ( Stenus ; Type locality: Mexico).

— Sharp, 1886: 653 ( Stenus ; Mexico).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 274 ( Stenus ; characters).

— Puthz, 1967q: 319 ( Stenus ; characters; notes).

— Puthz, 1971c: 17 ( Stenus ; characters).

- DISTRIBUTION: Mexico, Guatemala.

leucopus Sharp, 1886: 651 ( Stenus ; Type locality: Mexico, Jalapa, Cordova; Guatemala, San Joaquin, Volcan de Agua 8500 to 10,500 feet, Cerro Zunil, Dueñas , Purula , Sabo in Vera Paz ).

— Bernhauer and Schubert, 1911: 175 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 274 ( Stenus ; characters).

vexatus Casey, 1884: 51 ( Stenus ; Type locality: Thurston County, Washington Territory ).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Scheerpeltz, 1933: 1168 ( Stenus ; subgenus Stenus ; catalog).

— Sanderson, 1957: 268 ( Stenus ; possible synonym of austini ).

— Puthz, 1974d: 160 ( Stenus ; valid species; characters; notes).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

- DISTRIBUTION: USA, Canada.

vicarius Puthz, 1969 , see: malignus L. Benick, 1926 .

vicinoides Puthz, 1973 h: 200 ( Stenus ; Type locality: Canada: Saskatchewan: Maple Creek. Also cited from Manitoba, Alberta, British Columbia, North Dakota, Oregon , Wyoming , Nebraska , Colorado , Utah ).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

— Elias, 1991: 557 ( Stenus ; Quaternary fossil collection; Colorado).

- DISTRIBUTION: Canada, USA.

vicinus Casey, 1884: 57 ( Stenus ; Type locality: District Columbia; Pennsylvania; Selma, Alabama; Delaware Bay below Philadelphia; Cape May , New Jersey).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Puthz, 1972b: 12 ( Stenus ; characters; Michigan).

— Puthz, 1974d: 160 ( Stenus ; notes).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

— Downie and Arnett, 1996: 490 ( Stenus ; characters; USA).

- DISTRIBUTION: USA, Canada.

simplex Casey, 1884: 56 ( Stenus ; [preoccupied]; Type locality: Oregon).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Scheerpeltz, 1933: 1167 ( Stenus ; synonym of tricuspidatus ).

— Puthz, 1974d: 160 ( Stenus ; synonym of vicinus ).

tricuspidatus L. Benick, 1921c: 192 ( Stenus ; replacement name for simplex Casey ).

— Scheerpeltz, 1933: 1167 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1974d: 160 ( Stenus ; synonym of vicinus ).

vidua Assing, 2000a: 343 ( Stenus ; subgenus Parastenus ; Type locality: Portugal: Algarve, öslt. Lagos).

- DISTRIBUTION: Portugal.

vietnamensis Puthz, 1968 j: 464 ( Stenus ; subgenus Parastenus ; Type locality: Vietnam: Provinz Ninh-Binh, Res. Cuc-Phüöng: Dang, 200 m).

— Puthz, 1981a: 18 ( Stenus ; Vietnam).

— Puthz, 1981b: 146 ( Stenus ; characters).

- DISTRIBUTION: Vietnam.

viettei Jarrige, 1970: 55 ( Stenus ; subgenus Hypostenus ; Type locality: Mt. Tsaratanana, 1500 m).

— Puthz, 1972: 20 ( Stenus ; characters; Madagascar).

- DISTRIBUTION: Madagascar.

vilhenai Cameron, 1951a: 49 ( Stenus ; subgenus Hypostenus ; Type locality: Angola: Quanza, 12°S, 16°40′E, alt. 270 m).

— Puthz, 1965: 509 ( Stenus ; subgenus Hypostenus ; characters).

— Puthz, 1971: 251 ( Stenus ; subgenus Hypostenus ; characters; Angola).

- DISTRIBUTION: Angola.

vilis Kraatz, 1859 , see: fuscipes Gravenhorst, 1802 .

villiersi Cameron, 1949 , see: chyuluensis Cameron, 1942 .

villosiventris Lea, 1899: 543 ( Stenus ; Type locality: Windsor, N.S.W.).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1916: 241 ( Stenus ; subgenus Tesnus ; notes).

— Puthz, 1969e: 528 ( Stenus ; characters; Australia).

— Puthz, 1970e: 63 ( Stenus ; subgenus Tesnus ; characters; Australia).

— Puthz, 1972f: 109 ( Stenus ; Australia).

- DISTRIBUTION: Australia.

villosus Casey, 1884 , see: dolosus Casey, 1884 .

vinnulus Casey, 1884: 112 ( Stenus ; Type locality: Cambridge, Massachusetts; [Note: See lectotype designation by Puthz, 1972t]. Also cited from White Fish Point , Lake Superior ; Isle Royale , Lake Superior ; Marquette , Michigan).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Fall, 1926a: 61 ( Stenus ; Alaska, Yukon Territory).

Puthz, 1972t: 171 ( Stenus ; lectotype designation: Cambr. [Note: Cambridge , Massachusetts]) .

— Puthz, 1972e: 107 ( Stenus ; notes).

— Ryvkin, 1987c: 159 ( Stenus ; characters).

— Campbell and Davies, 1991: 112 ( Stenus ; USA; Canada).

— Downie and Arnett, 1996: 492 ( Stenus ; characters; USA).

— Ryabukhin, 1999: 46 ( Stenus ; subgenus Nestus ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Canada, USA, Finland.

confusoides Renkonen, 1935: 27 ( Stenus ; subgenus Nestus ; Type locality: Finnland: Lappland, Muonio).

— Palm, 1961a: 90 ( Stenus ; subgenus Nestus ; characters; Sweden; Norway; Finland).

— Puthz, 1967l: 49 ( Stenus ; notes; Transbaikal).

— Puthz, 1972t: 171 ( Stenus ; synonym of vinnulus ).

violaceus Bernhauer, 1906b: 194 ( Stenus ; Type locality: Brasilia).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974a: 69 ( Stenus ; notes).

- DISTRIBUTION: Brazil.

subviolaceus L. Benick, 1921b: 120 ( Stenus ; subgenus Hypostenus ; Type locality: Süd-Amerika: S. Catharina).

— Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974a: 69 ( Stenus ; synonym of violaceus ).

virginiae Casey, 1884: 87 ( Stenus ; Type locality: Pennington Gap , Virginia).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Puthz, 1974: 9 ( Stenus ; Illinois; Indiana; Missouri; Iowa; Maryland; Virginia; North Carolina; Tennessee; Texas).

— Puthz, 1974d: 163 ( Stenus ; notes).

— Puthz, 1988b: 141 ( Stenus ; characters; Kansas; Texas; Louisiana; Arkansas; Missouri; Iowa; Illinois; New York; Maryland; Virginia; West Virginia; North Carolina; Tennessee; Florida).

- DISTRIBUTION: USA.

texanus Casey, 1884: 88 ( Stenus ; Type locality: Columbus, Texas; western States ).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— L. Benick, 1938c: 266 ( Stenus ; characters).

— Puthz, 1974d: 163 ( Stenus ; synonym of virginiae ).

virgo Assing, 1996: 171 ( Stenus ; subgenus Parastenus ; Type locality: Spanien: Prov. Alicante, Sierra Bernia, 800 m).

- DISTRIBUTION: Spain.

virgula Fauvel, 1895: 210 ( Stenus ; Type locality: Bírmanie: Carin Ghécu , 1300–1400 m; [Note: See lectotype designation by Puthz, 1969]. Also cited from Bírmanie: Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m; Bengale, Kurseong; Sikkim, Darjeeling).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— Cameron, 1930: 394 ( Stenus ; subgenus Mesostenus ; characters; Burma; Bengal; Sikkim; India). — Puthz, 1969: 42 ( Stenus ; subgenus Parastenus ; lectotype designation: Carin Ghecù, 1300–

1400 m; characters; Burma; India).

— Puthz, 1971d: 23 ( Stenus ; characters; Taiwan).

— Puthz, 1971u: 532 ( Stenus ; characters; Taiwan).

— Puthz, 1976: 7 ( Stenus ; Nepal).

— Puthz, 1978: 130 ( Stenus ; India).

— Rougemont, 1981a: 339 ( Stenus ; Burma).

— Puthz, 1981b: 143 ( Stenus ; characters).

— Rougemont, 1983c: 44 ( Stenus ; Thailand).

— Ryvkin, 1992: 42 ( Stenus ; notes; Vietnam).

- DISTRIBUTION: India, Nepal, Bangladesh, Myanmar, Vietnam, China, Taiwan. kwantungensis Cameron, 1949b: 463 ( Stenus ; subgenus Hemistenus ; Type locality:

South China: eastern Kwantung, Yim Na San).

— Puthz, 1971u: 532 ( Stenus ; synonym of virgula ).

viridanoides Puthz, 1970: 18 ( Stenus ; subgenus Parastenus ; Type locality: Indien: Darjeeling). - DISTRIBUTION: India.

viridans Baudi, 1857: 109 ( Stenus ; Type locality: inundante fluvius Padus).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— L. Benick, 1917c: 57 ( Stenus ; notes).

— L. Benick, 1929a: 56 ( Stenus ; subgenus Hypostenus ; characters in key).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1970p: 175 ( Stenus ; notes).

- DISTRIBUTION: Italy.

viridans Champion, 1925 , see: viridanus Champion, 1925 .

viridanus Champion, 1925: 169 ( Stenus ; replacement name for viridans Champion ).

— Cameron, 1930: 398 ( Stenus ; subgenus Mesostenus ; characters; India).

— Scheerpeltz, 1933: 1198 ( Stenus ; subgenus Hemistenus ; catalog).

— Abdullah and Qadri, 1968a: 306 ( Stenus ; characters; Pakistan).

— Puthz, 1975: 196 ( Stenus ; Bhutan).

— Zheng, 1992b: 168 ( Stenus ; subgenus Parastenus ; characters; China).

— Zheng, 1993: 225, 227 ( Stenus ; subgenus Parastenus ; key characters; China; India).

- DISTRIBUTION: Pakistan, India, Bhutan, China.

viridans Champion, 1925: 106 ( Stenus ; subgenus Mesostenus ; [preoccupied]; Type locality: W. Almora Division of Kumaon and Nainital, Deopatta Peak, alt. 8000 ft.).

— Scheerpeltz, 1933: 1198 ( Stenus ; synonym of viridanus ).

viridescens Cameron, 1930: 340 ( Stenus ; Type locality: Chakrata district: Manjgaon; Chulli and Khedar Khuds, alt. 6500 to 7500 feet).

— Scheerpeltz, 1933: 1168 ( Stenus ; subgenus Stenus ; catalog).

— Abdullah and Qadri, 1968a: 305 ( Stenus ; characters; Pakistan).

— Puthz, 1980a: 29 ( Stenus ; characters).

- DISTRIBUTION: Pakistan, India.

virideus Cameron, 1939b: 140 ( Stenus ; subgenus Hypostenus ; Type locality: New Guinea:

Mt. Tafa, altitude 8550 feet).

— Puthz, 1972r: 478, 498 ( Stenus ; characters; Papua).

- DISTRIBUTION: New Guinea.

viridiaeneus W. J. MacLeay, 1873: 149 ( Stenus ; Type locality: Gayndah).

— Fauvel, 1877: 209 ( Stenus ; characters; Australia).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Puthz, 1970e: 62 ( Stenus ; characters; Australia).

- DISTRIBUTION: Australia.

viridimicans Puthz, 1975: 196 ( Stenus ; subgenus Parastenus ; Type locality: Bhutan: Nobding,

41 km nach Wangdiphodrang auf der Strasse nach Tongsa, 2800 m).

viridis Bernhauer, 1926a: 129 ( Stenus ; subgenus Hypostenus ; Type locality: Mindanao: Surigao).

— L. Benick, 1929: 271 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

- DISTRIBUTION: Philippines.

viriditinctus Champion, 1920 , see: Dianous .

visendus Last, 1970: 347 ( Stenus ; subgenus Hypostenus ; Type locality: New Guinea: Buana, Sarawaged Range, 3000 ft., 20–30 miles north of Lae).

— Puthz, 1972r: 480, 505 ( Stenus ; characters; New Guinea; Papua).

— Puthz, 1982a: 135 ( Stenus ; New Guinea).

- DISTRIBUTION: New Guinea.

vista Sanderson, 1946 , see: niveus Fauvel, 1865 .

vitabundus Puthz, 1988: 657 ( Stenus ; subgenus Hypostenus ; Type locality: North Borneo: Sabah: Tawan District, Kalabakan).

- DISTRIBUTION: Borneo.

vitalei Bernhauer, 1935a: 41 ( Stenus ; variety of brunnipes ; Type locality: Sizilien: Ciccia; Castanea; Calamara).

— Normand, 1935: 364 ( Stenus ; cited as brunnipes ; [Note: Misidentification of vitalei according to Puthz, 1968g]; Algeria; Tunisia).

— Korge, 1962: 151 ( Stenus ; subgenus Tesnus ; valid species; Sicily; Algeria).

— Puthz, 1967: 6 ( Stenus ; subgenus Nestus ; subspecies of brunnipes ).

— Puthz, 1968g: 205, 214 ( Stenus ; subgenus Tesnus ; valid species; characters; Italy; Dalmatia; Tunisia; Spain).

— Puthz, 1972s: 261 ( Stenus ; Yugoslavia; Portugal; Tunisia; Sicily; Italy; Dalmatia; Spain).

— Bordoni, 1975h: 7 ( Stenus ; distribution in Italy).

— Sparacio, 1995: 149 ( Stenus ; characters; notes; Sicily).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

- DISTRIBUTION: Morocco, Algeria, Tunisia, Libya, Portugal, Spain, Italy, Croatia.

yakourensis Fagel, 1957e: 325 ( Stenus ; subgenus Tesnus ; Type locality: Grande Kabylie: Yakouren, forêt de Beni-Ghobri, Bois Sacré, 750 m).

— Fauvel, 1902b: 76 ( Stenus ; cited as brunneipes; [Note: Misidentification of yakourensis according to Puthz, 1966f: 120]; Algeria; Tunisia).

— Korge, 1962: 151 ( Stenus ; synonym of vitalei ).

— Puthz, 1966f: 120 ( Stenus ; note; not synonym of brunnipes vitalei ).

— Puthz, 1967: 6 ( Stenus ; subgenus Nestus ; subspecies of brunnipes ).

— Puthz, 1967f: 273 ( Stenus ; subspecies of brunnipes ; Algeria; Tripoli; Malta Morocco).

— Puthz, 1968g: 214 ( Stenus ; synonym of vitalei ).

lusoensis Fagel, 1960c: 224 ( Stenus ; subgenus Tesnus ; Type locality: Portugal: concelho de Mealhada, Luso, route de Pampilhosa).

— Puthz, 1967: 6 ( Stenus ; subgenus Nestus ; subspecies of brunnipes ; Portugal).

— Puthz, 1968g: 214 ( Stenus ; synonym of vitalei ).

viti Puthz, 1977a: 109 ( Stenus ; Type locality: Iran: Guilan, Nav’s Valley , ca. 10 km Luftlinie vom Kaspischen Meer entfernt am Weg von Heroabad zum Meer, 1500 m).

— Ryvkin, 1990: 161 ( Stenus ; notes).

- DISTRIBUTION: Iran.

volkeri Naomi, 1998: 385 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Osaka-fu:

Mt. Mikusa).

- DISTRIBUTION: Japan.

voluptabilis Puthz, 1998: 383 ( Stenus ; subgenus Hypostenus ; Type locality: Vietnam: Hoa Binh). - DISTRIBUTION: Vietnam.

volvulus Fauvel, 1904e: 312 ( Stenus ; Type locality: Tamatave).

— Puthz, 1972: 9 ( Stenus ; characters; Madagascar).

— Puthz, 1992a: 15 ( Stenus ; Madagascar).

- DISTRIBUTION: Madagascar.

voraginosus L. Benick, 1938: 22 ( Stenus ; subgenus Parastenus ; Type locality: Java: G. Tangkoeban Prahoe, 4000–5000 Voet, Preanger; Malabar, Preanger, Kawah Wayang).

— Puthz, 1970: 22 ( Stenus ; subgenus Parastenus ; Java).

- DISTRIBUTION: Indonesia.

vorticosus Cameron, 1943, see: sikkimensis Cameron, 1928 .

vulcanus Rougemont, 1984: 236 ( Stenus ; subgenus Parastenus ; Type locality: C. Java:

Mt. Merapi, S. slope, ca. 1500 m).

- DISTRIBUTION: Indonesia.

vulgaris Puthz, 1984: 126 ( Stenus ; subgenus Hypostenus ; Type locality: Guatemala:

Los Amatos. Also cited from Honduras and Nicaragua).

- DISTRIBUTION: Guatemala, Honduras, Nicaragua.

vulturnus L. Benick, 1951, see: picipes Stephens, 1833 .

wadihalfaensis Scheerpeltz, 1963, see: cameratus L. Benick, 1921 .

wagneri L. Benick, 1917 , see: ampliventris J. Sahlberg, 1890 .

wagnerianus Puthz, 1999: 18 ( Stenus ; subgenus Hypostenus ; Type locality: Rwanda:

Forêt de Nyungwe, Kamiranzovu, 2000 m).

- DISTRIBUTION: Rwanda.

walkeri Bernhauer, 1931 , see: melanarius Stephens, 1833 .

wallacei Puthz, 1968 , see: nothus L. Benick, 1929 .

wallisi Puthz, 1996: 322 ( Stenus ; subgenus Hypostenus ; Type locality: Brasilien: Nova Teutonia, 27°11′S, 52°23′W).

- DISTRIBUTION: Brazil.

wanglangus Zheng, 1993c: 311 ( Stenus ; subgenus Parastenus ; cited as wanglangus in the abstract and wanglandus at the description [Note: The intended spelling is wanglangus ]; Type locality: China: Sichuan: Ping-Wu County, Wanglang).

- DISTRIBUTION: China.

wankai Fleischer, 1909 [Note: Nomen nudum], see: gracilipes Kraatz, 1857 .

warabi Naomi and Nomura, 1990: 48 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Hyogo Pref.: Hyonosen).

- DISTRIBUTION: Japan.

wasabi Hromádka, 1982: 133 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Kyoto:

Mt. Daimoji).

— Naomi, 1990a: 1 ( Stenus ; Japan).

— Naomi, 1997d: 601, 603 ( Stenus ; subgenus Hypostenus ; characters; discussion of internal structure of aedeagus; Japan).

- DISTRIBUTION: Japan.

wasmanni Fauvel, 1895: 214 ( Stenus ; Type locality: Bírmanie: Carin Cheba , 900–1100 m; [Note: See lectotype designation by Puthz, 1969]. Also cited from Bírmanie: Carin Ghécu, 1300–1400 m; Sikkim, Darjeeling).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Cameron, 1930: 376 ( Stenus ; subgenus Hypostenus ; characters; Burma; Sikkim; India; Assam). — Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

— Abdullah and Qadri, 1968a: 308 ( Stenus ; characters; East Pakistan).

— Puthz, 1969: 19, 31 ( Stenus ; subgenus Hypostenus ; lectotype designation: Carin Cheba; characters; Burma).

— Puthz, 1975: 189 ( Stenus ; Bhutan).

— Rougemont, 1980: 181 ( Stenus ; Nepal). — Rougemont, 1981a: 337 ( Stenus ; Burma).

— Rougemont, 1981b: 378 ( Stenus ; Thailand).

- DISTRIBUTION: India, Nepal, Bhutan, Bangladesh, Myanmar, Thailand.

watanabei Puthz, 1968 , see: coronatus L. Benick, 1928 .

watrousi Puthz, 1998 h: 142 ( Stenus ; subgenus Parastenus ; Type locality: Philippinen: Luzon: Mt. Makiling, 4 km SE Los Banos).

- DISTRIBUTION: Philippines.

weiratheri L. Benick, 1939, see: callidus Baudi, 1848 .

weisei Bernhauer, 1912 , see: sedatus Sharp, 1889 .

weiseri Bernhauer, 1927 , see: colonus Erichson, 1840 .

wendeleri L. Benick, 1929: 269 ( Stenus ; subgenus Hypostenus ; Type locality: Binaluan, Palawan).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

- DISTRIBUTION: Philippines.

tropicus Wendeler, 1926: 72 ( Stenus ; [preoccupied]; Type locality: Nord-Palawan [ Philippinen], Binaluan).

— Scheerpeltz, 1933: 1184 ( Stenus ; synonym of wendeleri ).

wenzeli Puthz, 1967 , see: sibiricus J. Sahlberg, 1880 .

werneri Puthz, 1974c: 166 ( Stenus ; subgenus Parastenus ; Type locality: Philippinen: Mindanao: Cotabato Province, Burungkot, Upi, 1500 feet).

- DISTRIBUTION: Philippines.

wilhelmmontis Puthz, 1982a: 125 ( Stenus ; subgenus Stenus ; Type locality: New Guinea:

NE: Mt. Wilhelm).

- DISTRIBUTION: New Guinea.

wittei Cameron, 1950 g: 22 ( Stenus ; subgenus Hypostenus ; Type locality: Belgian Congo: Ruhengeri, sources Kirii).

— Puthz, 1967j: 44 ( Stenus ; characters; Zaire; Ruanda).

— Puthz, 1971: 262 ( Stenus ; subgenus Hypostenus ; characters; Rwanda).

- DISTRIBUTION: Congo, Rwanda.

wittmeri Puthz, 1972 h: 123 ( Stenus ; subgenus Parastenus ; Type locality: Iran: Gole Gole Lovae, via Minoo-dasht, Strasse Gorgan-Meshed, 750–1400 m).

— Ryvkin, 1990: 202 ( Stenus ; notes).

- DISTRIBUTION: Iran.

wollastoni Gemminger and Harold, 1868: 641 ( Stenus ; replacement for fulvescens ).

— Fauvel, 1902b: 76 ( Stenus ; synonym of heeri ).

— Bernhauer and Schubert, 1911: 170 ( Stenus ; synonym of heeri ).

— Smetana, 1963b: 33 ( Stenus ; subgenus Tesnus ; valid species; characters; Madeira).

— Puthz, 1966g: 132 ( Stenus ; characters).

— Puthz, 1968g: 205 ( Stenus ; subgenus Tesnus ; characters).

— Assing and Wunderle, 1995b: 2 ( Stenus ; characters; Madeira).

- DISTRIBUTION: Madeira.

fulvescens Wollaston, 1857: 198 ( Stenus ; [preoccupied]; Type locality: Cruzinhas, Madeira Is.; [Note: See lectotype designation by Puthz, 1966g]. Originally cited from mountains of Madeira, at high elevations).

— Wollaston, 1865: 513 ( Stenus ; habitat; Madeira Islands).

— Wollaston, 1871: 309 ( Stenus ; notes; synonym of wollastoni ).

— Fauvel, 1902b: 76 ( Stenus ; variety of heeri ).

— Bernhauer and Schubert, 1911: 170 ( Stenus ; synonym of heeri ).

— Puthz, 1966g: 132 ( Stenus ; lectotype designation: Cruzinhas, Madeira Is.; synonym of wollastoni ).

wrangeli Puthz, 1973 h: 196 ( Stenus ; Type locality: USA: California: Sonoma Co. , Sebastopol).

- DISTRIBUTION: USA.

wuesthoffi L. Benick, 1941e: 135 ( Stenus ; subgenus Hemistenus ; Type locality: Kiev; Sarepta).

— Szujecki, 1961: 49 ( Stenus ; subgenus Hemistenus ; characters; Poland).

— Puthz, 1967l: 50 ( Stenus ; notes).

— Puthz, 1972d: 86 ( Stenus ; subgenus “ Hemistenus ”; characters).

— Ryvkin, 1990: 176 ( Stenus ; notes; Ukraine).

- DISTRIBUTION: Poland, Ukraine.

wuorentausi L. Benick, 1921 , see: hyperboreus J. Sahlberg, 1876 .

wusthoffianus Machulka, 1947 , see: tarsalis Ljungh, 1810 .

xenia Hromádka, 1983: 120 ( Stenus ; subgenus Parastenus ; Type locality: Brazil: 18 km NE Orixima).

- DISTRIBUTION: Brazil.

yakourensis Fagel, 1957, see: vitalei Bernhauer, 1935 .

yamajii Naomi and Puthz, 1994b: 305 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Okayama Pref.: Mt. Mumou, Shinshou-mura).

- DISTRIBUTION: Japan.

yamato Naomi, 1988 g: 57, 61 ( Stenus ; subgenus Stenus ; Type locality: Japan: Fukui Pref.: Karikomi Pond, Koike).

- DISTRIBUTION: Japan.

yanoi Sawada and Nakane, 1954 , see: anthracinus Sharp, 1889 .

yanoianus Puthz, 1980a: 33 ( Stenus ; Type locality: Taiwan but see Puthz , 1981d: 119 for specific locality).

— Puthz, 1981d: 119 ( Stenus ; subgenus Stenus ; [Note: The species is cited as new here but was cited the previous year in a key]; characters; type locality is: Formosa: Kodama, Arisan).

- DISTRIBUTION: Taiwan.

yashiro Naomi and Puthz, 1994a: 257 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Saga Pref.: Ryumon Valley).

— Naomi, 1988h: 65, 68 ( Stenus ; subgenus Parastenus ; cited as hirtellus [Note: Misidentification of yashiro according to Naomi and Puthz, 1994a: 257]; characters; Japan).

— Naomi, 1997c: 6 ( Stenus ; characters; Japan).

- DISTRIBUTION: Japan.

yasutoshii Naomi, 1997b: 137 ( Stenus ; subgenus Nestus ; Type locality: Japan: Yamanashi Pref.: near Akazawa Spa, Masuho).

- DISTRIBUTION: Japan.

yoshidai Naomi, 1997e: 757 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Tokushima Pref.: Nakatani, Shishikui, Kaifu).

- DISTRIBUTION: Japan.

yunnanensis Cameron, 1946a: 683 ( Stenus ; subgenus Hypostenus ; Type locality: Yunanfou). — Puthz, 1981a: 7 ( Stenus ; notes; China).

- DISTRIBUTION: China.

zaishin Naomi, 1990: 102 ( Stenus ; subgenus Nestus ; Type locality: Japan: Tokushima Pref.: Mt. Tsurugi).

- DISTRIBUTION: Japan.

ohbayashii Naomi, 1997b: 141 ( Stenus ; subgenus Nestus ; subspecies of zaishin ; Type locality: Japan: Ehime Pref.: Hontani, ca. 950 m, Odamiyama).

zdenae Hromádka, 1990: 127 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Gumma

Pr.: Mt. Kurofu, 2100 m).

— Puthz, 1993a: 166 ( Stenus ; subgenus Parastenus ; characters; Japan).

- DISTRIBUTION: Japan.

bunraku Hromádka, 1990: 128 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Gumma Pr.: near Mt. Shirane, 2000 m).

— Puthz, 1993a: 166 ( Stenus ; synonym of zdenae ).

zerchei Puthz, 1993b: 115 ( Stenus ; subgenus Parastenus ; Type locality: Süd-Karpaten: Fagaras: Bulea-See, 2050 m).

- DISTRIBUTION: Romania.

zetteli Puthz, 1998 h: 138 ( Stenus ; subgenus Hypostenus ; Type locality: Philippinen: Mindoro or.: W. Puerto Galera, Tamaraw Beach, Talipanan River).

- DISTRIBUTION: Philippines.

ziegleri Puthz, 1998 i: 217 ( Stenus ; subgenus Tesnus ; Type locality: Vietnam: Ha Tinh,

Ky Thuong, 18°00′ N, 106°06′ E, 125 m).

- DISTRIBUTION: Vietnam.

zimmermanni Puthz, 1968b: 43 ( Stenus ; subgenus Nestus ; Type locality: Japan: Mt. Takao, Tokyo-sub).

— Naomi, 1989f: 41 ( Stenus ; subgenus Nestus ; characters).

— Naomi, 1990: 101 ( Stenus ; characters; Japan).

— Naomi and Takeda, 1991: 10 ( Stenus ; notes; Japan).

- DISTRIBUTION: Japan.

curvaticellus Naomi, 1997b: 142 ( Stenus ; subgenus Nestus ; subspecies of zimmermanni ; Type locality: Japan: Ishikawa Pref.: Nabetani, Tatsunokuchi-cho).

zipanguensis Puthz, 1968 , see: coronatus L. Benick, 1928 .

zoufali Fleischer, 1909 , see: intricatus Erichson, 1840 .

zukwalanus Bernhauer, 1931 , see: aethiopicus Bernhauer, 1915 .

zuluanus Bernhauer, 1934, see: nairobiensis Fauvel, 1907 .

zumpti Bernhauer, 1939 i: 254 ( Stenus ; subgenus Hypostenus ; Type locality: Kamerun: Umg. Kamerunberg: Tiko; Bombe, Mungofluss).

Puthz, 1967i: 198 ( Stenus ; characters; Guinea) .

— Puthz, 1971: 219 ( Stenus ; subgenus Hypostenus ; characters; Cameroon; Guinea).

— Puthz, 1999: 17 ( Stenus ; aedeagus).

- DISTRIBUTION: Cameroon, Guinea.

zunicoides Puthz, 1974 g: 203 ( Stenus ; Type locality: Mexico: Xochimilco ).

- DISTRIBUTION: Mexico.

zunicus Casey, 1884: 32 ( Stenus ; Type locality: Arizona, Morrison ).

— Fall, 1901: 71 ( Stenus ; California).

— Bernhauer and Schubert, 1911: 169 ( Stenus ; catalog).

— L. Benick, 1938c: 264 ( Stenus ; characters).

- DISTRIBUTION: USA, Mexico.

abscondens Sharp, 1887: 798 ( Stenus ; Type locality: Mexico, Chihuahua city).

— Bernhauer and Schubert, 1911: 152 ( Stenus ; catalog).

— L. Benick, 1939: 19 ( Stenus ; subgenus Nestus ; characters).

— Puthz, 1974g: 205 ( Stenus ; synonym of zunicus Casey ).

zunilensis Sharp, 1886: 645 ( Stenus ; Type locality: Guatemala, Cerro Zunil 4000 to

5000 feet).

— Bernhauer and Schubert, 1911: 169 ( Stenus ; catalog).

— L. Benick, 1938c: 263 ( Stenus ; characters).

— L. Benick, 1939: 18 ( Stenus ; characters).

- DISTRIBUTION: Guatemala.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Tesnus

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