Tesnus guatemalenus, Sharp, 1886: 665

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

guatemalenus Sharp, 1886: 665 ( Stenus ; Type locality: Guatemala, Cahabon 1000 feet).

— Bernhauer and Schubert, 1911: 175 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 276 ( Stenus ; characters).

— Puthz, 1971c: 16 ( Stenus ; synonym of sagax ).

chontalenus Sharp, 1886: 666 ( Stenus ; Type locality: Nicaragua, Chontales).

— Bernhauer and Schubert, 1911: 173 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 275 ( Stenus ; characters).

— Puthz, 1971c: 16 ( Stenus ; synonym of sagax ).

simplicifrons L. Benick, 1928d: 38 ( Stenus ; Type locality: Costa Rica: Turialba , 800 m).

— Scheerpeltz, 1933: 1182 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 275 ( Stenus ; characters).

— Puthz, 1971c: 16 ( Stenus ; synonym of sagax ).

augustinus Blackwelder, 1943: 216 ( Stenus ; Type locality: Trinidad: St. Augustine ).

— Puthz, 1973c: 53 ( Stenus ; synonym of sagax ).

bryanti Blackwelder, 1943: 219 ( Stenus ; Type locality: Trinidad: cocoa estate about 6 miles from La Brea, county of St. Patrick , ward of La Brea).

— Puthz, 1971c: 16 ( Stenus ; synonym of sagax ).

sagittalis Puthz, 1970 h: 84 ( Stenus ; subgenus Hypostenus ; Type locality: New Ireland: Bismarck Inseln).

— Puthz, 1972r: 479 ( Stenus ; characters; New Guinea).

- DISTRIBUTION: New Guinea.

sagittifer Puthz, 1988c: 89 ( Stenus ; subgenus Hypostenus ; Type locality: Peru: Loreto,

west of Iquitos).

- DISTRIBUTION: Peru.

sahlbergi Puthz, 1968e: 93 ( Stenus ; subgenus Stenus ; Type locality: U.d.S.S.R.: Issyk-kul).

- DISTRIBUTION: Kyrgyzstan.

saizi Puthz, 1972 g: 125 ( Stenus ; subgenus Hypostenus ; Type locality: Chile).

— Puthz, 1973g: 177 ( Stenus ; notes).

- DISTRIBUTION: Chile.

sakaii Naomi and Puthz, 1996: 154 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Ehime Pref.: Mt. Saragamine, 1100 m alt.).

— Naomi, 1997c: 6 ( Stenus ; characters; Japan).

- DISTRIBUTION: Japan.

sakana Hromádka, 1979d: 107 ( Stenus ; subgenus Parastenus ; Type locality: Japan:

Kinki Distr., Asuka).

— Naomi, 1988h: 68 ( Stenus ; characters; Japan).

- DISTRIBUTION: Japan.

sakura Hromádka, 1979f: 115 ( Stenus ; subgenus Nestus ; Type locality: Japan: Pref. Nara: Mt. Ohdaj).

— Naomi, 1989f: 41 ( Stenus ; subgenus Nestus ; characters).

- DISTRIBUTION: Japan.

salamandra Bondroit, 1912 , see: crassus Stephens, 1833 .

salebrosus L. Benick, 1942: 25 ( Stenus ; subgenus Parastenus ; Type locality: NO-Burma: Kambaiti, 2000 m).

— Puthz, 1998: 391 ( Stenus ; subgenus Parastenus ; Vietnam; China).

- DISTRIBUTION: India, Myanmar, Vietnam, China.

mangpuensis Cameron, 1943: 3 ( Stenus ; subgenus Mesostenus ; Type locality: Darjeeling district: Ghum; Mangpu, Rongdong Valley).

— Puthz, 1978: 130 ( Stenus ; India).

— Puthz, 1998: 391 ( Stenus ; synonym of salebrosus ).

— Fauvel, 1876a: 63 [= 1876: 248] ( Stenus ; Portugal; Corsica; Dauria).

— Fauvel, 1878: 103 [= 1878a: 23] ( Stenus ; Algeria).

— Rey, 1884: 369 [= 1884a: 217] ( Stenus ; subgenus Hemistenus ; characters; France).

— Fauvel, 1886: 30 [= 1886a: 22] ( Stenus ; Morocco; Algeria).

— Ganglbauer, 1895: 588 ( Stenus ; subgenus Hemistenus ; characters; western middle Europe; Spain; Algeria).

— Fauvel, 1902b: 77 ( Stenus ; Morocco; Algeria; Tunisia).

— Sainte-Claire Deville, 1906: 82 ( Stenus ; Corsica).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1917c: 57 ( Stenus ; notes).

— L. Benick, 1929a: 92 ( Stenus ; subgenus Hemistenus ; variety of binotatus ; characters in key). — Portevin, 1929: 401 ( Stenus ; characters; France).

— L. Benick, 1932b: 3 ( Stenus ; variety of binotatus ; notes; Morocco, Spain).

— Scheerpeltz, 1933: 1186 ( Stenus ; synonym of binotatus ).

— Scheerpeltz, 1960d: 134 ( Stenus ; subgenus Hemistenus ; variety of binotatus ; Portugal).

— Puthz, 1967: 7 ( Stenus ; subgenus Hemistenus ; Portugal).

— Puthz, 1971v: 103 ( Stenus ; valid species; notes; France; Spain; Portugal; North Africa ; Sardinia; Corsica) .

— Puthz, 1972d: 86 ( Stenus ; subgenus “ Hemistenus ”; characters).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

- DISTRIBUTION: France, Spain, Portugal, Italy, Morocco, Algeria, Tunisia. subconvexus Rey, 1884: 370 ( Stenus ; subgenus Hemistenus ; Type locality: Algérie).

— Fauvel, 1902b: 77 ( Stenus ; synonym of salinus ).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; synonym of salinus ).

— Scheerpeltz, 1933: 1186 ( Stenus ; synonym of binotatus ).

— Puthz, 1972d: 90 ( Stenus ; synonym of binotatus and salinus ).

— Puthz, 1973b: 29 ( Stenus ; synonym of salinus ).

salisburgensis Bernhauer, 1927, see: crassus Stephens, 1833 .

sallaei Sharp, 1886 , see: renifer LeConte, 1863 .

salticola Puthz, 1967 , see: leleupi Puthz, 1965 .

salvini Sharp, 1886: 659 ( Stenus ; Type locality: Panama, Volcan de Chiriqui 4000 to

6000 feet, Bugaba).

— Bernhauer and Schubert, 1911: 177 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 273 ( Stenus ; characters).

— L. Benick, 1939: 38 ( Stenus ; subgenus Hypostenus ; characters).

- DISTRIBUTION: Panama.

samarensis Bernhauer, 1926a: 128 ( Stenus ; subgenus Hypostenus ; Type locality:

Island Samar).

— L. Benick, 1929: 250 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1182 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974c: 158 ( Stenus ; Philippines).

- DISTRIBUTION: Philippines.

basilanus Puthz, 1974c: 158 ( Stenus ; subgenus Hypostenus ; subspecies of samarensis ; Type locality: Philippinen: Basilan).

sanctaecatharinae Bernhauer, 1916f: 174 ( Stenus ; subgenus Hypostenus ; Type locality: Brasilien: S. Catharina).

— L. Benick, 1921b: 121 ( Stenus ; characters).

— Scheerpeltz, 1933: 1182 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974a: 67 ( Stenus ; characters).

- DISTRIBUTION: Brazil.

sandakanensis Bernhauer, 1926a: 131 ( Stenus ; subgenus Hemistenus ; Type locality: Borneo: Sandakan).

— Rougemont, 1986: 282 ( Stenus ; Sarawak).

— Puthz, 1988: 661 ( Stenus ; Borneo).

- DISTRIBUTION: Borneo.

sandersoni L. Benick, 1941, see: capucinus Boheman, 1858 .

sandersonianus Puthz, 1972, see: pubescens Stephens, 1833 .

sandrae Cornell and Puthz, 1997: 132 ( Stenus ; subgenus Parastenus ; Type locality:

North Carolina: Swain County: Fork Ridge Trail near Clingman’s Dome).

- DISTRIBUTION: USA.

sanfilippoi Puthz, 1996a: 3 ( Stenus ; subgenus Hypostenus ; Type locality: Madagaskar: Umbebung von Antsirabe, ca. 1700 m).

- DISTRIBUTION: Madagascar.

sangensis Puthz, 1971: 98 ( Stenus ; Type locality: Congo: Kivu Terr. Uvira, Haute Sange , 2900 m, Ilot forestier).

- DISTRIBUTION: Congo.

sangha Puthz, 1986 h: 12 ( Stenus ; subgenus Parastenus ; Type locality: Nepal: Provinz Bagmati: Yangri Ridge, 4700–4800 m).

- DISTRIBUTION: Nepal.

sankisianus Bernhauer, 1915 m: 298 ( Stenus ; subgenus Hypostenus ; Type locality: Katanga: Sankisia).

— Scheerpeltz, 1933: 1182 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1967i: 225 ( Stenus ; characters).

— Puthz, 1971: 300 ( Stenus ; subgenus Hypostenus ; characters; Zaire).

- DISTRIBUTION: Congo.

sannator Puthz, 1991d: 2 ( Stenus ; Type locality: Indien: West Bengal, Darjeeling District: Mahanadi bei Kurseong , versant sud, 1200 m).

- DISTRIBUTION: India.

sannio Puthz, 1980d: 187 ( Stenus ; subgenus Stenus ; Type locality: Indien: Darjeeling Distrikt: Singmari-Bharapatea Bung).

— Puthz, 1991d: 3 ( Stenus ; characters; India; Nepal).

- DISTRIBUTION: India, Nepal.

santira Naomi, 1988 i: 78 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Miyagi Pref.: Mt. Zaô).

— Naomi and Takeda, 1991: 12 ( Stenus ; Japan).

- DISTRIBUTION: Japan.

sarawakensis Puthz, 1973d: 79 ( Stenus ; subgenus Hypostenus ; Type locality: Borneo: Sarawak: Mt. Dulit, R. Koyan, 2500 feet).

— Puthz, 1988: 659 ( Stenus ; characters; Borneo).

- DISTRIBUTION: Borneo.

sardous Kraatz, 1857 , see: ossium Stephens, 1833 .

sasajii Naomi, 1988 , see: puberulus Sharp, 1874 .

satoi Puthz, 1991d: 20 ( Stenus ; subgenus Hypostenus ; Type locality: Philippinen: Palawan: Iwahin River Valley).

— Puthz, 1998h: 122 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

- DISTRIBUTION: Philippines.

satsuki Naomi, 1987: 1 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Ehime Pref.:

Mt. Ishizuchi).

— Naomi, 1988h: 70 ( Stenus ; characters; Japan).

- DISTRIBUTION: Japan.

satushin Naomi and Puthz, 1994: 212 ( Stenus ; subgenus Nestus ; Type locality: Japan:

Ehime Pref.: Mt. Takanawa, Hôjô-shi).

sauteri Bernhauer, 1907: 379 ( Stenus ; Type locality: Kanagawa).

— Bernhauer and Schubert, 1911: 166 ( Stenus ; catalog).

— Naomi, 1988g: 62 ( Stenus ; subgenus Stenus ; characters; Japan).

— Naomi and Takeda, 1991: 9 ( Stenus ; Japan).

— Naomi, Kuranishi, Saito, and Maruyama, 2000: 108 ( Stenus ; subgenus Stenus ; Kamchatka). - DISTRIBUTION: Japan.

sauterianus Bernhauer, 1943a: 173 ( Stenus ; Type locality: Formosa: Koroton ).

— Puthz, 1968b: 41 ( Stenus ; subgenus Stenus ; characters; Taiwan).

— Puthz, 1971d: 21 ( Stenus ; characters; Taiwan).

— Puthz, 1971u: 532 ( Stenus ; characters; Taiwan).

— Puthz, 1981d: 118 ( Stenus ; characters).

- DISTRIBUTION: Taiwan.

sawadai Hromádka, 1979d: 108 ( Stenus ; subgenus Parastenus ; Type locality: Japan:

Pref. Hyogo: Sasayama).

— Naomi, 1988h: 70 ( Stenus ; characters; Japan).

- DISTRIBUTION: Japan.

sawadaianus Hromádka, 1979c: 100 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Pref. Kyoto: Mt. Hiei, Yamashiro).

— Naomi, 1988h: 70 ( Stenus ; characters; Japan).

— Puthz, 1993a: 160 ( Stenus ; subgenus Parastenus ; characters; Japan).

— Naomi, 1997c: 2, 6 ( Stenus ; subgenus Parastenus ; characters; Japan).

- DISTRIBUTION: Japan.

sawadaiellus Naomi and Puthz, 1994b: 299 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Nagano Pref.: Mt. Ontake).

- DISTRIBUTION: Japan.

saxatilis Gistel, 1857: 66 ( Stenus ; [Note: Nomen dubium]; Type locality: Helvetia).

- DISTRIBUTION: Switzerland.

saxicola Puthz, 1974 , see: Dianous .

sayi Casey, 1884: 164 ( Areus ; Type locality: California).

— Fall, 1901: 71 ( Stenus ; California).

— Bernhauer and Schubert, 1911: 177 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 274 ( Stenus ; characters).

— Sanderson, 1957: 253 ( Stenus ; subgenus Hypostenus ; characters; British Columbia; Washington; Oregon; Idaho).

— Puthz, 1966f: 120 ( Stenus ; subgenus Hypostenus ; notes).

— Puthz, 1971i: 1751 ( Stenus ; aedeagus).

- DISTRIBUTION: Canada, USA.

scaber Fauvel, 1871: 20 ( Stenus ; Type locality: Nice; [Note: See lectotype designation by Puthz , 1969]. Also cited from Corse, Ajaccio ; Piémont; Bône).

— Fauvel, 1873: 127 [= 1873a: 285] ( Stenus ; characters; France; Italy; Corsica; Algeria).

— Fauvel, 1878: 104 [= 1878a: 24] ( Stenus ; Algeria).

— Rey, 1884: 346 [= 1884a: 194] ( Stenus ; subgenus Mesostenus ; characters; France).

— Fauvel, 1886: 31 [= 1886a: 23] ( Stenus ; Algeria).

— Ganglbauer, 1895: 596 ( Stenus ; subgenus Mesostenus ; characters; France; Italy; Corsica; Algeria).

— Fauvel, 1902b: 79 ( Stenus ; Algeria).

— Sainte-Claire Deville, 1906: 84 ( Stenus ; Corsica).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1929a: 67 ( Stenus ; subgenus Parastenus ; characters in key).

— Portevin, 1929: 404 ( Stenus ; characters; France).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Wüsthoff, 1934a: Tafel III ( Stenus ; illustration of aedeagus).

— Puthz, 1970p: 176 ( Stenus ; synonymic notes).

— Puthz, 1970m: 223 ( Stenus ; notes).

— Puthz, 1971v: 87 ( Stenus ; Italy) .

— Puthz, 1972s: 269 ( Stenus ; notes).

— Tikhomirova, 1973a: 174 ( Stenus ; subgenus Parastenus ; Sredizemonomor’e).

— Bordoni, 1974b: 2 ( Stenus ; subgenus Hemistenus ; Italy).

— Bordoni, 1975c: 181 ( Stenus ; Italy; Sardinia; Sicily; Capri) .

— Bordoni, 1975h: 20 ( Stenus ; Italy; Corsica; Sardinia; Sicily; Capri).

— Puthz, 1980e: 360 ( Stenus ; Croatia).

— Angelini, 1991: 194 ( Stenus ; Italy).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

— Herman, 2001: 34 ( Stenus ; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).

- DISTRIBUTION: Algeria, Italy, France, Croatia.

bituberculatus Motschulsky, 1857a: 511 ( Stenus ; [Note: This name is older than scaber ]; Type locality: Marseille).

— Fauvel, 1873: 127 [= 1873a: 285] ( Stenus ; synonym of scaber ).

— Ganglbauer, 1895: 596 ( Stenus ; synonym of scaber ).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; synonym of scaber ).

— Herman, 2001: 34 ( Stenus ; synonym of scaber ).

italicus Baudi, 1870: 397 ( Stenus ; [Note: This name is older than scaber ]; Type locality: Pedemontium; [Note: See lectotype designation by Puthz, 1970p]. Also cited from Ravenna).

— Fauvel, 1873: 127 [= 1873a: 285] ( Stenus ; synonym of scaber ).

— Rey, 1884: 347 [= 1884a: 195] ( Stenus ; synonym of scaber ).

— Ganglbauer, 1895: 596 ( Stenus ; synonym of scaber ).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; synonym of scaber ).

— Puthz, 1969: 39 ( Stenus ; synonym of scaber ).

— Puthz, 1970p: 176 ( Stenus ; lectotype designation; synonym of scaber ).

— Herman, 2001: 32 ( Stenus ; synonym of scaber ).

scabiosus Casey, 1884: 60 ( Stenus ; Type locality: Marquette , Lake Superior).

— Blatchley, 1910: 407 ( Stenus ; characters; Indiana).

— Bernhauer and Schubert, 1911: 166 ( Stenus ; catalog).

— Puthz, 1973h: 203 ( Stenus ; characters; Manitoba; Yukon Territory; Alaska).

— Campbell and Davies, 1991: 112 ( Stenus ; USA; Canada).

— Downie and Arnett, 1996: 490 ( Stenus ; characters; USA).

- DISTRIBUTION: Canada, USA.

scabriculus J. Sahlberg, 1876: 53 ( Stenus ; Type locality: Pyhäjärvi i Yläne, vid Helsingfors; Ruovesi i norra Tavastland, vid Svir, Justjärvi och Petrosavodsk i Ryska Karelen, vid Kirjavalaks, i Ladoga Karelen, vid Haapajärvi i Nurmis på Lysarholmen nära Jakobstad samt nordligast vid Kuusamo kyrkoby).

— Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— L. Benick, 1929a: 13 ( Stenus ; subgenus Tesnus ; characters in key; Fennoscandia; Russia).

— Scheerpeltz, 1933: 1172 ( Stenus ; subgenus Tesnus ; catalog).

— Palm, 1961a: 100 ( Stenus ; subgenus Tesnus ; characters; Sweden; Norway; Finland).

— Puthz, 1968g: 205 ( Stenus ; subgenus Tesnus ; characters).

- DISTRIBUTION: Norway, Finland, Russia.

scabrior Stephens, 1833 , see: cicindeloides Schaller, 1783 .

scabripennis Sharp, 1886: 649 ( Stenus ; Type locality: Mexico, Jalapa, Cordova; Guatemala, near the city, San Geronimo, Capetillo , Panima in Vera Paz , Cubilguitz ; Panama, Bugaba ).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1938c: 280 ( Stenus ; characters).