Xenophyllum dactylophyllum (Sch.Bip.) V.A.Funk, Novon 7(3): 239. 1997.

Calvo, Joel & Moreira-Munoz, Andres, 2020, Taxonomic revision of the Andean genus Xenophyllum (Compositae, Senecioneae), PhytoKeys 158, pp. 1-106 : 1

20. Xenophyllum dactylophyllum (Sch.Bip.) V.A.Funk, Novon 7(3): 239. 1997.

Werneria dactylophylla Sch.Bip., Bonplandia (Hannover) 4: 53, 55. 1856. Type. Peru. Puno: pr. Agapata [Ayapata] ad nives eternas, 4875 m, Jun 1854, W. Lechler 1807 (lectotype: G-00305680 (digital image!), designated by Funk (1997a: 239); isolectotypes: BR s.n.!, GH s.n.!, GOET s.n.!, K-000527745 (digital image!), LE s.n.!, LL-00374330 (digital image!), NY s.n.!, P-02088571 (digital image!), P-02088572 (digital image!), P-02088573 (digital image!), P-02088575 (digital image!), S-R-6522 (digital image!), W s.n.!).

Werneria dactylophylla f. glabriuscula Rockh., Bot. Jahrb. Syst. 70: 286. 1939. Type. Bolivia. San Francisco-Tal, Ancohuma, 5000 m, 3 Jul 1928, C. Troll 2076 (lectotype: M-0147065 (digital image!), designated here; isolectotype: B s.n.!, CONC-80182!).

Xenophyllum oscartovarii E.Linares, J.Campos, Nauray, Vicente Orell. & A. Galán, Arnaldoa 17(1): 104. 2010 [ “oscartovari”]. Type. Peru. Arequipa: Caylloma, sector Derivación, 19L 0179145 8636313 [erroneous, ca. 15°00'S, 72°11'W], 4740 m, 4 Feb 2009, E. Linares & A. Galán 2354 (holotype: USP n.v.; isotype: AQP n.v.), syn. nov.

Description.

Suffruticose plant, forming clumps of erect stems or cushions. Rhizomes 5-15 × 0.5-0.8 cm, horizontal to oblique, glabrous. Stems 11-24 cm tall, branched, glabrous, usually with leaves restricted to the upper part. Leaves imbricate, extending into a sheath-like base glabrous or with long silky trichomes; leaf laminas 5.7-9.9 × 2.4-4.1 mm, spatulate (finger-like at the apex), at least 9-divided at the apex, with the primary division extending deeper than the subsequent ones, entire, sometimes with some minute cilia, elliptical to terete upwards in cross section, glabrous or floccose-lanate, 1-nerved above (barely visible), 1-nerved beneath, somewhat fleshy, matte; primary division 2-3 mm long, 3 to 6 divided, rounded to truncate, sometimes with some scattered cilia on the margin. Capitula radiate, erect, sessile. Involucres 10.8-12.8 × 8.2-9.5 mm, cupuliform; involucral bracts 11 to 20, 6.6-7.6 × 1.9-2 mm, obtuse at the apex, greenish, sometimes purple-edged. Ray florets 17 to 22; corollas 9.2-11 × 1.3-1.6 mm, 4-veined, subentire to 3-toothed at the apex, conspicuously surpassing the involucre, white. Disc florets 37 to 50; corollas 5.3-6.3 mm long, yellowish; style branches truncate with a crown of sweeping trichomes or slightly penicillate, purplish. Achenes 3.1-4.8 × 0.9-1.3 mm, cylindrical, 7 to 9-ribbed, glabrous; pappus 8.1-8.9 mm long, barbellate, whitish. Chromosome number 2 n = 108( ± 4) ( Diers 1961). Fig. 25 .

Additional iconography.

Linares et al. (2010: 102, fig. 1, sub X. oscartovarii , as photo); Beltrán (2016: 359, fig. 3A, as photo).

Distribution and habitat.

Central Peru to central Bolivia. Bolivia (Cochabamba, La Paz, Potosí [n.v.]), Peru (Ancash, Apurímac, Arequipa, Ayacucho, Cusco, Huancavelica, Huánuco, Junín, Lima, Moquegua, Pasco, Puno). This species grows in rocky outcrops, scree slopes, cryoturbated soils, and exposed grasslands of the subhumid and humid puna ecoregions, between elevations of (2800-)3400-5500 m (Fig. 24 ).

Xenophyllum dactylophyllum has been reported from the Bolivian department of Potosí on the basis of the collection Steinbach s.n. ( Jørgensen 2014; Tropicos 2020). This material, which has not been located, was collected nearby Andacaba, a locality ca. 30 km south of the city of Potosí. If the identification is confirmed, it would represent the southern distribution limit of this species. The specimen US-00974139 most likely corresponds to a mislabeling because both the locality (San Carlos de Bariloche, Río Negro Province, Argentina) and the elevation (600 m) do not correspond to the known distribution area and altitudinal rank of this species.

Phenology.

Flowering nearly all year round.

Etymology.

The epithet dactylophyllum means finger-like leaves and it refers to the resemblance between the leaves of this plant and a hand.

Notes.

Xenophyllum dactylophyllum is characterized by having glabrous or floccose-lanate finger-like leaves, with at least 9 divisions at the apex. The divisions are usually arranged in multiples of three, the primary one extending deeper than the subsequent ones. The capitula are radiate, erect, with 17 to 22 ray florets with white corollas conspicuously surpassing the involucre.

Another species with finger-like leaves is X. staffordiae , which has occasionally been confused with X. dactylophyllum . They can be readily differentiated by the habit (11-24 cm tall in X. dactylophyllum vs. 30-60 cm tall in X. staffordiae ), leaf divisions (at least 9-divided, with the primary division extending deeper than the subsequent ones in X. dactylophyllum vs. at least 12-divided, with all divisions similar in length in X. staffordiae ), capitulum position (erect in X. dactylophyllum vs. rather nodding in X. staffordiae ), and length and color of the ray corollas (9.2-11 mm, surpassing the involucre, white in X. dactylophyllum vs. 6.4-8.3 mm, not surpassing the involucre, pale yellow in X. staffordiae ). Their distribution areas overlap from central to southern Peru. Other morphologically similar species are X. amblydactylum and X. lorochaqui (see comments under these species).

Our efforts in locating the type material of X. oscartovarii E.Linares, J.Campos, Nauray, Vicente Orell. & A. Galán were unfruitful. Due to the original description and the pictures included in the publication ( Linares et al. 2010), we believe that it fairly corresponds to X. dactylophyllum . Oddly, this latter species was not mentioned in the taxonomic discussion of the presumed new species. In addition, we studied several specimens of X. dactylophyllum collected close to the type locality of X. oscartovarii .

Additional specimens examined.

Bolivia. Cochabamba: im Geröll des Cerro Tunari, 17°17'S, 66°23'W, May 1911, T. Herzog 2094 (S n.v.; locality confirmed by M. Zárate in litt.); La Paz: Murillo, Huayna Potosí, 29 Apr 1977, S.G. Beck 1 (LPB); Bautista Saavedra, 1 km antes de la cumbre, donde hay desvíos por Charazani, Ulla Ulla y Amarete, 3 Aug 1985, S.G. Beck 11347 (LPB); Inquisivi, cordillera Tres Cruces, al pie de glaciar Atoroma, 16°55'S, 67°25'W, 23 Dec 1978, S.G. Beck 183 (LPB); Murillo, Milluni 11 km hacia Tuni, pie del glacial Huayna Potosí y María Lloco, 5 Jan 1980, S.G. Beck 2425 (LPB); Murillo, subiendo el valle Khallapa (hacia Hampaturi), entrando al valle lateral del río Palcoma y subiendo hacia la serranía Murillo, 16°25'S, 67°57'W, 6 May 2012, S.G. Beck 33940 (LPB); Murillo, fin del valle Kaluyo pasando el pueblo de Chacaltaya, Pampalarama, 150 m por debajo del glaciar, 16°19'S, 68°4'W, 14 Mar 2013, S.G. Beck, D. Ibáñez & C. Beck 34084 (LPB); Larecaja, cumbre, arriba del abra hacia mina Fabulosa, 16°3'S, 68°18'W, 10 Mar 2016, S.G. Beck 35111 (LPB); La Fabulosa, 16°3'S, 68°18'W, 26 Apr 1950, W.M. Brooke 6322 (CONC); Murillo, en las faldas del cerro Huayna Potosí, camino a Tuni Condoriri, 28 Jun 1991, V. Camacho & M. Rendon 1 (LPB); Loayza, sierra de Tres Cruces, 22 Feb 1979, A. Ceballos et al. 472 (COL); Bautista Saavedra, camino entre Escoma y Charazani, zona del ANMI Apolobamba, 15°15'S, 69°2'W, 4 Oct 2005, N. de la Barra et al. 1026 (BOLV, LPB); Humasani, 15 Jun 1974, Essel 23 (LPB); Bautista Saavedra, Pumazani, am Beginn der Strabe nach Amarete, 15°13'S, 68°59'W, 25 May 1980, T. Feuerer 4228 (LPB); Murillo, Zongo-Tal, am Lago Zongo, 16°17'S, 68°7'W, 1 Jun 1980, T. Feuerer 4335a (LPB); Los Andes, an der Strabe zur Mine Fabulosa, 10 Feb 1980, T. Feuerer 8472 (LPB); valle de Zongo, 1.5 km avant la cumbre, 16°16'S, 68°7'W, 24 Apr 1988, A. Fournet 815 (BOLV); Bautista Saavedra, Amarka, cerca del límite occidental del ANMI Apolobamba, 15°15'S, 69°2'W, 4 Sep 2004, A. Fuentes & C. Aldana 6721 (BOLV, LPB); Murillo, nev. Huayna Potosí, E slopes above rd., 16°17'S, 68°8'W, 12 Apr 1995, V.A. Funk & N. Bernal 11282 (LPB); Murillo, nev. Charquini, above aqueduct on N facing slope, 16°17'S, 68°6'W, 12 Apr 1995, V.A. Funk & N. Bernal 11287 (LPB); Los Andes, Hichu-Kkota valley, 20 km from base of lgn. Khara Kkota along rd. to mina Fabulosa, 16°10'S, 68°20'W, 25 Apr 1995, V.A. Funk & C. González-Quint 11371 (LPB, US); Los Andes, Hichu-Kkota valley, 21 km from base of lgn. Khara Kkota along rd. to mina Fabulosa, near the pass, 16°10'S, 68°20'W, 25 Apr 1995, V.A. Funk & C. González-Quint 11372 (LPB, US); Los Andes, above cumbre (pass) on rd. through Hichu-Kkota valley on rd. to mina La Fabulosa, 21 km from base of lag. Khara Kkota, 16°10'S, 68°20'W, 29 Apr 1995, V.A. Funk 11408 (LPB); Bautista Saavedra, Charazani, pampa de Ulla-Ulla, cerca de la estancia Medallani, 15°4'S, 69°6'W, 4 May 1993, P. Gutte & B. Herzog 702 (LPB); Franz Tamayo, Ulla Ulla, cordillera Apolobamba cerca al pueblo de Pelechuco 17 km NE, 14°55'S, 69°10'W, 22 Aug 1986, P. Holt 15A (LPB); José M. Camacho, 60 km al E de Ulla Ulla, arriba del paso Osipal, 29 Jul 1993, P. Holt 2 (LPB); José M. Camacho, 60 km al E de Ulla Ulla, arriba de Osipal, 29 Jul 1993, P. Holt 3 (LPB); Murillo, cerro Charquini, a 1h 20min del centro de La Paz, 16°17'S, 68°6'W, Jun 2015, A. Huon & R.I. Meneses 1 (LPB); Murillo, Huayna Potosí, 29 Apr 1977, E. Jordan 1 (LPB); Bautista Saavedra, división camino Amarete-Charazani-Ulla Ulla, 15°14'S, 69°3'W, 14 Jun 1997, M. Kessler et al. 10144 (LPB); Murillo, Zongo pass, slope up towards Huayna Potosí, 16°14'S, 68°8'W, 6 Aug 1991, M. Kessler 2840 (LPB); Humasani, 17 Jun 1975, R. Lara 1499 (LPB); Larecaja, viciniis Sorata, prope Anilaya, ad lacum Yuriguana, 15°45'S, 68°33'W, Mar/May 1858, G. Mandon 101 (LE, RB, W); Bautista Saavedra, abra Pumasani, 15°15'S, 69°2'W, 20 Apr 1982, X. Menhofer 1108 (LPB); Murillo, nevado Huayna Potosí, 14 Jul 1982, X. Menhofer 1420 (LPB); Franz Tamayo, lado E de la cordillera de Apolobamba, camino de Pelechuco a Sorata, cerca de la mina abandonada de Sunchuli, 15°1'S, 69°0'W, 25 May 1984, X. Menhofer 2344 (LPB); Franz Tamayo, lado E de la cordillera de Apolobamba, camino de Pelechuco a Sorata, cerca de la mina abandonada de Sunchuli, 15°1'S, 69°0'W, 25 May 1984, X. Menhofer 2345 (LPB); Omasuyos, Hichu Cota, subiendo a la cumbre, 16°7'S, 68°21'W, 28 Apr 1985, M. Moraes 150 (LPB); Los Andes, Hichu-Kkota, 26 Jan 1984, C. Ostria 119 (LPB); Los Andes, valle de Hichu Kkota, 17 Nov 1983, C. Ostria 48 (LPB); Inquisivi, cordillera Tres Cruces, 13 Sep 2001, B.J. Ruthsatz 10533 (LPB); Murillo, 4.5 km N of Milluni on road to Zongo pass (ca. 3 km S of pass), 16°18'S, 68°8'W, 7 Feb 1985, J.C. Solomon 13210 (LPB); Murillo, pass at the head of valle del Zongo, 16°17'S, 68°8'W, 6 Mar 1983, J.C. Solomon 9785 (LPB); Larecaja, Sorata, laguna glacial nevado Illampu, 15°49'S, 68°33'W, 20 Jul 1996, J.R.I. Wood 11255 (BOLV, HSB, LPB); Bautista Saavedra, on road between Escoma and Charazani, ca. 3 km before descent to Charazani, 15°16'S, 69°1'W, 10 Jun 2000, J.R.I. Wood & C. Wendleburger 16400 (LPB); Larecaja, Sorata, zona de la laguna glacial nevado Illampu, 15°50'S, 68°36'W, 7 Apr 2004, J.R.I. Wood 20663 (LPB); Murillo, cordillera Huayna Potosí, base de la cordillera de Huayna Potosí, 16°16'S, 68°7'W, 27 Mar 2007, M. Zárate 2440 (BOLV, LPB); Franz Tamayo, Pelechuco, al N en línea recta a 3.8 km del campamento Chocollo, 14°43'S, 69°13'W, 25 Nov 2017, F. Zenteno, D. Villalba & L. Mamani 21277 (LPB). Peru. Ancash: abra Yanashalla, 9°51'S, 77°4'W, 25 May 2013, C. Aedo & J. Molina 20440 (MA); Huaylas, Macate, cerro al norte de la laguna Capalo, 21 May 2000, A. Cano, A. Ramírez & C. Cáceres 10549 (USM); Asunción, Chacas, debajo de la punta Olímpica, entre las lagunas, 9°8'S, 77°30'W, 24 Apr 2004, A. Cano, M.I. La Torre & W. Mendoza 14536 (USM); Asunción, Chacas, alrededores de la laguna Lebrón, 9°12'S, 77°29'W, 20 May 2009, A. Cano et al. 19378 (USM); Huari, Rajupampa, laderas hacia el nevado Santa Rosa, 9°29'S, 77°17'W, 7 Oct 2012, A. Cano et al. 21319 (USM); Huaylas, Riurín, alturas de Pueblo Libre y alrededores, cerro Huachoq Qocha y cerro Gallu Huaqanan, 9°12'S, 77°47'W, 19 May 1999, A. Cano et al. 9252 (USM); Huaylas, Ocshapampa (Oxapampa), 9°11'S, 77°51'W, 11 Oct 1999, A. Cano et al. 9733 (USM); Bolognesi, Pariac Punta, pampa de Lampas, Chiquián, 10°11'S, 77°12'W, 4 May 1952, E. Cerrate 1513 (USM); Bolognesi, Ticllos, paso de Chonta, 10°15'S, 77°15'W, 29 Apr 1956, E. Cerrate 2651 (USM); Bolognesi, Chilcas, laguna de Quitcka, 25 Nov 1981, E. Cerrate, B. León & J. Campos 8400 (USM); near top of divide over cordillera Blanca, upper slopes of Huascarán, above lagunas Llanganuco, 9°1'S, 77°35'W, 10 Jul 1982, A. Gentry et al. 37431 (USM); Asunción-Chacas, laguna Librón, 9°12'S, 77°29'W, 13 Aug 2008, B. Roca Ramos 1 (USM); Asunción-Chacas, laguna Librón, 9°12'S, 77°29'W, 13 Aug 2008, B. Roca Ramos 2 (USM); Asunción-Chacas, laguna Atlanta, a 2 km de la laguna Atlanta o Azulcocha, 9°3'S, 77°33'W, 12 Aug 2008, B. Roca Ramos 3 (USM); Huari, camino Olleros a Chavín, desde el abra hasta el lado oriental de la cordillera Blanca, 9°36'S, 77°17'W, 21 Oct 1999, J. Roque & K. Young 1221 (USM); Recuay, Huascarán N.P., pass between nevado Pasto Ruri and nevado Raria, río Pachacoto drainage, 9°52'S, 77°11'W, 31 Mar 1985, D.N. Smith & F. Escalona 10207 (USM); Yungay, Huascarán N.P., Llanganuco sector, quebrada Demanda, W of Chacraraju base camp, 9°1'S, 77°36'W, 13 Apr 1985, D.N. Smith & V. Cautivo 10290 (USM); Recuay, Huascarán N.P., quebrada Quenua Ragra, 9°58'S, 77°13'W, 10 May 1985, D.N. Smith, R. Valencia & A. Gonzales 10700 (LPB, USM); Recuay, Huascarán N.P., camino de herradura passing over Cahuish tunnel, 9°41'S, 77°15'W, 10 Jul 1985, D.N. Smith & M. Buddensiek 11126 (USM); Carhuaz, Huascarán N.P., quebrada Ishinca, side valley to laguna Ishinca, 9°23'S, 77°25'W, 16 Jul 1985, D.N. Smith & M. Buddensiek 11208 (USM); Huari, Huascarán N.P., just crossing the Ulta pass, 9°7'S, 77°30'W, 28 Jul 1985, D.N. Smith 11304 (USM); Carhuaz, Huascarán N.P., quebrada Honda, from pass towards valley bottom, N-side of valley, 9°18'S, 77°24'W, 3 Oct 1985, D.N. Smith, M. Buddensiek & R. Valencia 11644 (USM); Recuay, Huascarán N.P., lateral valley of quebrada Queshque, toward río Pachacoto drainage, 9°50'S, 77°18'W, 19 Mar 1986, D.N. Smith, R. Valencia & M. Torres 11900 (USM); Yungay, Huscarán N.P., between lake Llanganuco and Portachuelo, 9°3'S, 77°36'W, 16 Aug 1984, D.N. Smith 8258 (USM); Huaylas, Huascarán N.P., pass between quebrada Los Cedros and Hatuncocha, 8°51'S, 77°45'W, 12 Mar 1985, D.N. Smith & R. Valencia 9963 (USM); Huaylas, sector denominado Huiripunta, 6 Sep 1994, G. Yarupaitán & E. Salas 1482 (USM); Apurímac: Abancay, Ampay, 13°34'S, 72°53'W, Aug 1937, C. Vargas & Santander 514 (CUZ); Arequipa: pr. Chivay, ladera S del nevado Huarancante, 15°45'S, 71°32'W, 1 Apr 2005, C. Aedo & A. Galán 11037 (MA, USM); Caylloma, pasando la cumbre Chucura, 15°45'S, 71°33'W, 16 Jan 1999, S.G. Beck 26468 (LPB); Castilla, Orcopampa, minas de Poracota, cerca a quebrada Faculla, 15°14'S, 72°32'W, 20 Apr 2011, H. Beltrán 7115 (USM); Ramón Castilla, Orcopampa, alrededores de Cia. Minera Ares, 15°16'S, 72°17'W, 31 Mar 2000, A. Cano & N. Valencia 10106 (USM); La Unión, Huaynacotas, Sarajorepampa, 15°1'S, 72°47'W, 18 Mar 2011, D. Montesinos 2951 (HSP, MOL); Castilla, Tapay, cerro Blanco, Apacheta, 17 Sep 2011, N. Vega 1782 (USM); Condesuyos, nevado de Coropuna, 15°34'S, 72°42'W, Jun 1976, M. Weibel s.n. (USM); Ayacucho: Cangallo, Paras, abra Apacheta, 13°21'S, 74°44'W, 25 Jun 2001, J. Roque & C. Arana 3289 (USM); Páucar del Sara Sara, Oyolo, a 15 km al NO de Pampamarca, camino a Sayla, 15°5'S, 73°2'W, 14 Sep 2013, C. Tejada 233 (HSP); Cusco: Urubamba, Machupijchu, 13°10'S, 72°32'W, Nov 1982, B. Peyton 97 (CUZ); Espinar, al oeste del poblado de Condoroma, a 1 km de la carretera, 15°16'S, 71°10'W, 30 Sep 2009, W. Ramírez 585 (USM); Urubamba, Pumahuanca, 13°11'S, 72°8'W, 8 Jun 1991, A. Tupayachi 1515 (CUZ); Huancavelica: Huaytará, alrededores de abra Apacheta, 13°18'S, 74°41'W, 3 Sep 2013, P. Gonzáles et al. 2780 (USM); Huachocolpa, alrededores de la unidad minera Caudalosa, 13°4'S, 75°0'W, 23 Mar 2015, P. Gonzáles 3528 (USM); paso de Chonta, 12°38'S, 74°26'W, 8 May 1958, O. Tovar 2948 (USM); Huánuco: Lauricocha, Cauri, alrededores de las lagunas Tinkicocha, Lauricocha, Tauricocha y Patarcocha, 10°27'S, 76°44'W, 8 Sep 2012, P. Gonzáles & N. Valencia 1921 (USM); Dos de Mayo, entre Lauricocha y Otuto, [without date], A. Raimondi 1589 (USM); Lauricocha, San Miguel de Cauri, campamento Raura, laguna artificial antes de ingresar a Niñococha bajo, 10°27'S, 76°45'W, 7 May 2004, F. Salvador, S. Ríos & E. Arias 735 (USM); Lauricocha, San Miguel de Cauri, a 100 m de puesto de control Santa Rosa (campamento Raura), ladera abajo, 11°35'S, 76°11'W, 13 May 2004, F. Salvador, S. Ríos & E. Arias 944 (USM); Junín: Huancayo, Pucará, Raquina, 12°10'S, 75°7'W, [without date], D. Barrón 57 (USM); Yauli, paso de Anticona, cumbre más alta de la carretera central, 11°35'S, 76°11'W, 17 Dec 1951, E. Cerrate 998 (USM); Anticona pass, ca. 140 km E of Lima on Hwy. to La Oroya, 11°35'S, 76°11'W, 16 Dec 1978, M. Dillon & B.L. Turner 1300 (USM); cercanías de Morococha, 11°35'S, 76°8'W, 13 Nov 1863, J. Isern 404 (MA); La Oroya, 11°31'S, 75°53'W, 15 Jun 1915, K. Maisch 150 (USM); Huaytapallana, 11°57'S, 75°2'W, 4 May 1961, O. Tovar 3386 (USM); Yauli, Ticlio, 11°35'S, 76°11'W, Feb 1974, O. Tovar 7187 (USM); Yauli, Milloc, camino del desvío de Casapalca, 20 Mar 1983, M.A. Vargas 77 (USM); Jauja, al pie del nevado de Tullujuto, Aug 1920, A. Weberbauer s.n. (USM); Huancayo, Quilcas, Tunacorral, 11°51'S, 75°9'W, 5 Apr 1994, G. Yarupaitán, E. Olivera & R. Canto 1355 (USM); Lima: cordillera Raura, pr. mina Raura, 10°29'S, 76°44'W, 30 May 2013, C. Aedo & J. Molina 20538 (MA); Huarochirí, paso de Anticona, 11°35'S, 76°11'W, 1 May 1999, H. Beltrán & E. Li 3307 (USM); Yauyos, Laraos, pampas de Quiray cerca a la laguna de Huinso, 12°23'S, 75°48'W, 4 Feb 2000, H. Beltrán 3415 (USM); Yauyos, Laraos, estancia de Quiray, 6 horas de camino al pueblo, 12°24'S, 75°45'W, 11 May 2001, H. Beltrán 4059 (USM); Yauyos, Laraos, camino de Jalcacha a Palca, 12°24'S, 75°45'W, 4 Nov 1992, H. Beltrán 406 (USM); Yauyos, Laraos, Malpaso, 12°27'S, 75°40'W, 12 May 2001, H. Beltrán 4204 (USM); Oyón, compañía minera Iscaycruz, 10°47'S, 76°43'W, 30 May 2008, H. Beltrán 6509 (USM); Huarochirí, desvío de carretera central hacia Chinchan y Marcapomacocha, 11°34'S, 76°15'W, 23 Sep 2014, H. Beltrán & W. Aparco 7737 (USM); Huarochirí, desvío de carretera central hacia Chinchan y Marcapomacocha, 11°34'S, 76°15'W, 23 Sep 2014, H. Beltrán & W. Aparco 7749 (USM); Oyón, encima de mina Uchuchacua, parte más alta por carretera, 10°36'S, 76°40'W, 5 Apr 2015, H. Beltrán & W. Aparco 7771 (USM); Huarochirí, Chicla, abra Anticona (Ticlio), 11°36'S, 76°11'W, 29 Apr 2017, H. Beltrán, S. Castillo & M. Arakaki 7965 (USM); límite con dpto. Junín, entre Casapalca y Ticlio, 11°35'S, 76°11'W, 1 Dec 1977, S. Castroviejo, M. Costa & E. Valdés-Bermejo 1118 (MA); Huarochirí, arriba de Chumpicocha, 11°57'S, 76°9'W, 28 May 1953, E. Cerrate 2011 (USM); Anticona nevado, 11°35'S, 76°10'W, 8 Aug 1987, M. Chanco, O. Tovar & A. Galán 1220 (USM); Huarochirí, lago Aguascocha, near mina Caprichosa, above Casapalca, 11°38'S, 76°13'W, 1 Mar 1964, P.C. Hutchison & O. Tovar 4270 (USM); subida al puerto de Piedra Parada, camino de Lima a Tarma, 26 Sep 1863, J. Isern 434 (MA); Canta, La Viuda, 11°21'S, 76°26'W, 7 Aug 1964, I. Meza 228 (USM); Morococha, 11°35'S, 76°8'W, [without date], A. Raimondi 8389 (USM); Cajatambo, Raura, 10°26'S, 76°47'W, 15 Apr 1988, S. Rivas et al. s.n. (USM); Canta, Cullhuay, alrededores de la laguna de Chuchún, 11°22'S, 76°26'W, 17 May 2003, G. Vilcapoma 6070 (USM); Moquegua: General Sánchez Cerro, Yunga, Sura-Perusa, 16°11'S, 70°38'W, 13 Apr 2012, D. Montesinos 3789 (HSP, USM); General Sánchez Cerro, Yunga, Sura, 16°11'S, 70°38'W, 3 Mar 2018, D. Montesinos & J. Calvo 5938 (HSP); Pasco: abra Ucchuchacua, 10°36'S, 76°40'W, 31 May 2013, C. Aedo & J. Molina 20584 (MA); Pasco, Huaron, 11°2'S, 76°27'W, Nov 1956, H. Macedo s.n. (USM); Puno: Carabaya, Corani, Minaspata, 14°2'S, 70°43'W, 15 Oct 2016, P. Gonzáles 3829 (USM).

Figures

Figure 24. Distribution map of Xenophyllum decorum (left hand, closed triangle), X. amblydactylum (left hand, open circle), and X. dactylophyllum (right hand).

Figure 25. Xenophyllum dactylophyllum A habit B stem apical part and capitulum C leaf D ray corolla and style E disc corolla and style branches F disc corolla and stamens (vertically sectioned, style removed) G style H achene with pappus. All details drawn from Funk & Gonzalez-Quint 11372 (US). Illustration by Alice Tangerini.