Arachnospila (Arachnospila) fumipennis (Zetterstedt 1838)

Lelej, Arkady S. & Loktionov, Valery M., 2011, Review of the nominotypical subgenus of Arachnospila Kincaid (Hymenoptera: Pompilidae) of Russia and neighbouring countries with the lectotypification of enigmatic Pompilus sogdianus Morawitz and description of new species, Zootaxa 2882, pp. 1-18 : 7-9

Arachnospila (Arachnospila) fumipennis (Zetterstedt 1838)

(Figs 12, 13, 30, 50–53)

Pompilus fumipennis Zetterstedt 1838: 438 , Ƥ [type locality "Lapponia Tornensis" ( Sweden)].

Psammochares fumipennis: Gussakovskij 1932: 44 , 3.

Pompilus (Arachnospila) fumipennis fumipennis: Wolf 1964a: 19 (Ƥ), 24 (3), 25; Krombein 1979: 1565.

Arachnospila (Arachnospila) fumipennis: Móczár 1968: 431 , Ƥ; Tobias 1978: 126 (Ƥ), 127 (3); Wahis 1986: 18; Oehlke & Wolf 1987: 350, Ƥ 3; Zonstein 1989: 41; Lelej 1995: 238 (Ƥ), 241 (3); 2005: 129, Ƥ; Loktionov et al. 2009: 228.

Arachnospila (Arachnospila) fumipennis fumipennis: Priesner 1968: 177 , 179, 180, Ƥ 3; Wolf 1972: 95 (Ƥ), 103 (3), 108.

Arachnospila fumipennis: Özbek et al. 2000: 118 , Ƥ 3; Zonstein 2002b: 136, Ƥ 3; Wolf 2004: 1144, Ƥ; 2005: 1744, Ƥ; Wiśniowski 2009: 184 (Ƥ), 186 (3), 197; Loktionov & Lelej 2010: 54.

Pompilus arrogans Tournier 1890: 217 , Ƥ (type locality " Helvetia "), nom. praeocc., nec Smith 1873. Junior subjective synonym of Pompilus fumipennis fumipennis Zetterstedt 1838 according to Wolf 1964a: 25.

Pompilus butaurus Tournier 1890: 216 , Ƥ (type locality " Helvetia "). Junior subjective synonym of Pompilus fumipennis fumipennis Zetterstedt 1838 according to Wolf 1964a: 25.

Pompilus arrogantulus de Dalla Torre 1897: 274. Replacement name for Pompilus arrogans Tournier 1890 nec Smith 1873. Junior subjective synonym of Arachnospila fumipennis Zetterstedt 1838 according to Wahis 1986: 18.

Pompilus nostras Kohl 1898: 362 , Ƥ (type locality "Arnsdf., 1808" [Arnsdorf, Germany]). Junior subjective synonym of Pompilus fumipennis fumipennis Zetterstedt 1838 according to Wolf 1964a: 25.

Pompilus borealis Aurivillius 1907: 23 , Ƥ 3 (type locality " Suecia borealis: Angermannia et Lapponia " [North Sweden]). Junior subjective synonym of Psammochares fumipennis Zetterstedt 1838 according to Haupt 1927: 195.

Psammochares lasiope Banks 1919: 81 , Ƥ [syntype: "Saranac Lake, 26 Aug.; Wilmington, 20–26 Aug.; New Russia, Essex Co., 18 Aug., (Bradley leg.)" ( USA)]. Junior subjective synonym of Pompilus (Arachnospila) fumipennis Zetterstedt 1838 according to Krombein 1979: 1565.

Psammochares malaisei Nordström & Forsius 1930: 1 , 3 nec Ƥ [holotype, 3, "Kamtchatka, Petropavlovsk, 2.–5.VII.1920 " (Kamchatka, Russia) [Stockholm Museum], examined by J. Abenius].

Pompilus (Arachnospila) fumipennis septimanus Wolf 1964a: 27 , Ƥ 3 [holotype, Ƥ, "Massif de la Massane, Garrigue, Pyr. Or., 2.VIII.1949 //? nostras Kohl Ƥ , J. de Beaumont det. 1950" ( France)]. Junior subjective synonym of Pompilus fumipennis Zetterstedt 1838 according to Wahis 1986: 18.

Arachnospila (Arachnospila) fumipennis paladeae Wolf & Móczár 1972: 244 , 245 (key), 261, Ƥ 3 (holotype, Ƥ, Mongolia: Bulgan aimak, 7 km NW von Somon Chanžargalant, 1350 m, 16.VI.1968). Junior subjective synonym of Pompilus fumipennis Zetterstedt 1838 according to Wahis 1986: 18.

Diagnosis of male. Hypopygium basally without sharp differences between sclerotized and unsclerotized parts (Figs 12, 13); median carina of hypopygium preapically distinctly concave (lateral view) (Fig. 13); genitalia as in Fig. 30 ; fore and hind wings as in Figs 52, 53; flagellomere 8 length 2.0–2.4 × its width; metasoma with ferruginous-red basal segments.

Diagnosis of female. Protarsomere 1 with three spines of tarsal comb; fore and hind wings as in Figs 50, 51; flagellomere 7 length 3.3 × its width; flagellomere 8 length 3.6–3.8 × its width; T1, T2 usually ferruginous-red.

Material examined. Russia. Primorskiy Terr.: 1 3, "Kedrovaya Pad" Reserve, 23.IX.1976; 2 3, Anisimovka, 1.VI.1994, 15. VI.1997; 1 3, Kievka, 14.V.1979; 1 Ƥ, Lazovsky Reserve, 17.VII.2007; 1 3, Preobrazhenie, 24.VIII.1979; 2 Ƥ 4 3, Barabash-Levada, 17.VII.1962, 2.IX.1978, 4–6.VI.1980; 1 Ƥ, 15 km SE Barabash-Levada, 5.VI.1986; 1 Ƥ, 20 km NW Melnichnoe, 2.VIII.1986; 1 3, Chuguevka, 20.VIII.2008; 3 3, Vysokogorsk, 12.VI, 28.VII.1986; 1 3, Nesterovka, 5.VII.1986 [ IBSS]; 1 3, Samarga River, 1– 4.VIII.[19]25. Khabarovsk Terr.: 1 Ƥ 2 3, Khabarovsk, 2.IX. [19]25 [ ZIN]; 1 Ƥ, mouth of Suluk-Makit River, 7.VII.1988; 1 Ƥ, Khummi, 25.VIII.1972; 2 Ƥ, mouth of Gorin River, 5.IX.1984, 20.IX.1985; 1 3, Amut Lake, 21.VII.1991; 4 3, Gornyi, 29.VI.2005. Amurskaya Prov.: 1 3, Arkhara, 9.VI.1987; 1 Ƥ 1 3, Svobodnoe, 1.VIII.2003; 5 Ƥ, Svobodnyi Trud, 11.VIII.1982; 2 Ƥ, Saskal, 12, 13.VIII.1982; 1 Ƥ, Gluchari, 15.VIII.1982; 1 Ƥ, Dim, 16.VII.1975; 1 Ƥ, Novospassk, 8.VIII.1982. Sakhalin: 1 Ƥ, 60 km N Okha, forest, 31.VII.1981 [ IBSS]. Kamchatka: 1 3, Elizovo, VII.1928; 1 3, the same place, 23.VII. [19]29 [ ZIN]; 1 Ƥ, Lazo, 15.VII.1958; 1 Ƥ 1 3, Kozyrevsk, 20.VI.1975, 16.VII.1985; 1 3, pass between volcanoes Koryakskiy and Avachinskiy, 1000 m, 26.VII.1985; 1 3, Vestnik Bay, 27.VII.1999. Magadan Prov.: 1 Ƥ 6 3, Seimchan, 24.VII, 4.VIII.1975; 2 Ƥ, upper part of Kolyma River, mouth of Detrin River, 10– 14.VII.1992; 1 3, 50 km N Magadan, 2.VII.1975; 1 3, 150 km NW Magadan, 13.VII.1975; 1 3, 20 km E Kulu, 25.VII.1983; 1 3, lower part of Karkaron River, 5.VII.1982. Chukotka: 2 Ƥ, Omolon River, 180 km N Omolon, 17, 28.VII.1976; 1 Ƥ, upper part of Bolshaya Osinovaya River, 24.VII.1991. Yakutia: 1 3, Pokrovsk, 20.VII.1986. Chita Prov.: 1 Ƥ 3 3, 20 km SSE Krasnokamensk, 4, 5.VIII.2007. Buryatia: 1 3, Khoronkhoi, 22.V.1980; 3 Ƥ, Troitzkosavsk [Kyakhta], 13, 20.VII. [19]26; 1 3, the same place, 23.VI.1928; 1 Ƥ, Baraty, 26.VII.2007; 1 Ƥ, Kyahta, 29.V.2008; 1 3, Ust-Kiran, Chikoi River, 28.V.2008. Irkutsk Prov.: 1 Ƥ 7 3, Baikalsk, 20.VI, 29– 30.VIII.1983, 16–17.VII.1988, 14.VIII.1994; 1 Ƥ 1 3, Angarsk, 8–9.VIII.1994; 3 3, 15 km E Ust-Orda, 31.VII.1994, 2–3.VIII.1994; 1 3, Bratsk, 6.VI.1995 [ IBSS]. Krasnoyarsk Terr.: 1 Ƥ 1 3, Krasnoyarsk. Khakasia: 1 Ƥ, "Osnatjenn." [Sayanogorsk]. Komi: 1 Ƥ, Ust-Tzilma, 15.VII–8.VIII. [19]08. Murmansk Prov.: 1 3, Khibiny, 4.VII. [1]928. Stavropol Terr.: 1 Ƥ, Kislovodsk. Karachaevo-Tcherkesia: 1 Ƥ, Krasnogorskaya, 11.VI.1925 [ ZIN]. Kazakhstan. East-Kazakhstan Prov.: 1 3, 40 km S Tarbagatay, 4.VII.1986; 1 3, 33 km NE Tarbagatai, 8.VII.1986. Akmolinsk Prov.: 1 3, 20 km E Kokchetav, 17.VII.1977 [ IBSS]. Pavlodar Prov.: 1 3, Pavlodar, 8.VII.1929. Aktjubinsk Prov.: 3 Ƥ, Shalkar, Barsuki sands [ ZIN].

Distribution. Russia (Primorskiy Terr., Khabarovsk Terr., Amurskaya Prov., North Sakhalin, Kamchatka, Magadan Prov., Chukotka, Yakutiya, Buryatia, Irkutsk Prov., Krasnoyarsk Terr., *Khakasia, center and south of European part) ( Lelej 2005; Tobias 1978); Albania, Austria, Belgium, Bosnia and Herzegovina, Czech Republic, Finland, France, Germany, Greece, Italy, Norway, Romania, Slovakia, the Netherlands ( Wahis 2010), Poland ( WiŠniowski 2009), Turkey ( Özbek et al. 2000), Israel ( Wolf 1998a), Near East ( Wahis 2010), Mongolia ( Móczár 1968), Kazakhstan, Kyrgyzstan ( Zonstein 2002b), Tajikistan ( Wolf 2004). In North America occurs from Alaska, Yukon and Northwest Territory to Labrador south to New York, North Dakota, Alberta and Washington ( Krombein 1979, Finnamore 1997).

Remarks. J. Abenius (Swedish Environmental Protection Agency, Nynäshamn) kindly informed us (per. comm.) that males (genitalia of holotype have been examined by him) of Psammochares malaisei Nordström & Forsius 1930 belong to A. fumipennis (Zetterstedt 1838) while the females of malaisei belong to sogdiana -group. Unfortunately we have not seen the slide of male genitalia of P. malaisei and postponed the decision about the synonymy of this species.

Figures

FIGURES 28 – 35. Male genitalia, ventral view. 28. Arachnospila sogdianoides, Chita Prov. 29. A. sogdiana, lectotype. 30. A. fumipennis, Irkutsk Prov. 31. A. eisukei, Iturup. 32. A. amurensis, Primorkiy Terr. 33. A. clericalis Irkutsk Prov. 34. A. rufa, Omsk. 35. A. gussakovskii sp. nov., holotype.