Aleiodes (Arcaleiodes) siamensis, Quicke & Butcher, 2012

Butcher, Buntika Areekul, Smith, M. Alex, Sharkey, Mike J. & Quicke, Donald L. J., 2012, A turbo-taxonomic study of Thai Aleiodes (Aleiodes) and Aleiodes (Arcaleiodes) (Hymenoptera: Braconidae: Rogadinae) based largely on COI barcoded specimens, with rapid descriptions of 179 new species, Zootaxa 3457, pp. 1-232 : 18-32

publication ID

https://doi.org/ 10.11646/ZOOTAXA.3457.1.1

publication LSID

lsid:zoobank.org:pub:A8F8CF32-00EA-4877-A299-872C6B2081BA

persistent identifier

https://treatment.plazi.org/id/0D272257-9E74-E300-FF4D-FBD8DEA764A3

treatment provided by

Felipe

scientific name

Aleiodes (Arcaleiodes) siamensis
status

 

Key to the Aleiodes (Aleiodes) View in CoL View at ENA and Aleiodes (Arcaleiodes) species of Thailand

1 Hind trochantellus elongate, dorsal length at level where trochantellus most produced, more than 0.9 times maximum dorsal length of trochanter, and ventral (maximal) length more than 4 x posterior depth; hind coxa strongly transversely striate dorsally ( Fig. 3D View FIGURE 3 ); fore wing 2 nd submarginal cell always short, with vein 3-SR less than 1.2 x length of vein r; hind wing subbasal cell densely and evenly setose; males sometimes with large, setose, lozenge-shaped pits on metasomal tergites 2 and 3; females with large granular appearing gap between widely separated placode sensillae on medioventral part of antenna ( Fig. 4D View FIGURE 4 ); propodeum elongate and angular postero-laterally with posteriorly-directed tooth..................................................................................... Heterogamus View in CoL (not treated here)

- Hind trochantellus not elongate, dorsal (shortest) length shorter than 0.8 times dorsal length of trochanter, and ventral length less than 4 x posterior depth ( Fig. 182D View FIGURE 182 ); hind coxa usually without oblique striations basally ( Figs 181D View FIGURE 181 ); fore wing 2 nd submarginal cell variable; hind wing subbasal cell variable but often glabrous on at least anterior half; males never with large, setose, lozenge-shaped pits on metasomal tergites 2 and 3; females usually with placode sensillae evenly spaced around median flagellomeres ( Figs 4A,B View FIGURE 4 ) OR if with granular gap ( Fig. 4C View FIGURE 4 ), pronotum not elongate and deep pronope present ( Figs 75D View FIGURE 75 , 141D View FIGURE 141 ); propodeum seldom elongate seldom with postero-lateral angulation with posteriorly-directed tooth (e.g. Fig. 72E View FIGURE 72 ), though may have well-developed tooth on non-elongated propodeum ( Fig. 18E View FIGURE 18 )............... 2

2 (1) Hind wing vein 1-M moderately strongly arched ( Figs 192A View FIGURE 192 , 193A View FIGURE 193 ); fore wing veins 1-M and m-cu distinctly diverging posteriorly ( Fig. 193A View FIGURE 193 ); antennae with white annulus ( Fig. 193A View FIGURE 193 )............................................ 3

- Hind wing vein 1-M moderately more or less straight ( Figs 88G View FIGURE 88 , 97G View FIGURE 97 ); fore wing veins 1-M and m-cu usually distinctly converging posteriorly, rarely more or less parallel ( Fig. 97G View FIGURE 97 ) ( Aleiodes View in CoL s.s.) OR if diverging, (A. ( Hemigyroneuron )) distal part of fore wing subbasal cell strongly modified and with large glabrous area containing sclerome (see Figs 166D,E View FIGURE 166 ); antennae variable........................................................................... 6

3 (2) Distal part of fore wing subbasal cell strongly modified and often with large glabrous area containing sclerome (see Figs 166D,E View FIGURE 166 ); fore wing subdiscal cell glabrous (see Figs 166B,D,E View FIGURE 166 )............................................................. other S. E. Asian A. ( Hemigyroneuron ) species not yet recorded from Thailand but likely to occur there

- Distal part of fore wing subbasal not modified as above, setose, never with sclerome; fore wing subdiscal cell usually evenly setose [A. ( Arcaleiodes ) species]................................................................. 4

4 (3) Propodeum black ( Fig. 193E View FIGURE 193 ); metasoma striped ( Fig. 193F View FIGURE 193 ), tergite 1 with medial transverse black band, tergite 2 black, with posterior 0.3 white; apex of antenna entirely black; only apical segment of maxillary palp white, basal ones brown or black............................................................................................ 5

- Propodeum pale rufous ( Fig. 192E View FIGURE 192 ); metasoma not striped, tergite 1 orange except cream on apical 0.2, tergite 2 ivory white on basal 0.7, black apically; apex of antenna distinctly whitish ( Figs 192E,F View FIGURE 192 ); two apical segments of maxillary palp white ( Fig. 192C View FIGURE 192 )....................................................... A. (Arcaleiodes) siamensis sp. nov. 5 (4) Fore wing vein 1-CU1 0.6 x 2-CU1; subdiscal cell without longitudinal glabrous line subanteriorly; hind tibia white on basal 0.75, black apically (Fig. 194A); 3 rd metasomal tergite finely longituinally striate on basal 0.75 (Fig. 194F)........................................................................ A. (Arcaleiodes) vanachterbergi sp. nov.

- Fore wing vein 1-CU1 0.75 x 2-CU1; subdiscal cell with longitudinal glabrous line subanteriorly; hind tibia entirely orange ( Fig. 191A View FIGURE 191 ); 3 rd metasomal tergite more strongly longitudinally striate on basal 0.5 ( Fig. 191E View FIGURE 191 )....................................................................................... A. (Arcaleiodes) dresdeni sp. nov.

6 (2) Distal part of fore wing subbasal cell strongly modified and with large glabrous area containing sclerome ( Figs 166D,E View FIGURE 166 ); fore wing subdiscal cell glabrous ( Fig. 165E View FIGURE 165 ); males with glandular pore opening near the midposterior margin of metasomal tergites 4–6........................................................ A. (Hemigyroneron) roberti

- Distal part of fore wing subbasal usually unmodified, rarely largely glabrous BUT never with sclerome; fore wing subdiscal cell usually evenly setose; males [for all those species for which it is known] without gland pore on tergites 4–6 ................................................................................................. 7

7 (6) Fore femur very robust, less than 3.7 x longer from distal part of trochantellus to apex, than maximally deep ( Figs 78A View FIGURE 78 , 116A View FIGURE 116 , 145C View FIGURE 145 ); fore tarsus (excluding aroleum) very short, less than 0.75 x length of fore tibia, its median segment quadrate ( Fig. 145C View FIGURE 145 )....................................................................................... 8

- Fore femur never so robust, always more than 4 x longer from distal part of trochantellus to apex, than maximally deep ( Fig. 121A View FIGURE 121 ); fore tarsus (excluding aroleum) longer, more than 0.82 [minimally 0.85] x length of fore tibia ( Figs 15A View FIGURE 15 , 22A View FIGURE 22 ), its median segment approximately 2 x longer than wide or longer...................................... 11

8 (7) Hind basitarsus strongly laterally compressed and dorsally expanded medially in lateral view, inner face with dense long setae ( Fig. 3A View FIGURE 3 ); size variable but up to 1cm long................................................... Yelicones View in CoL 1

- Hind basitarsus normal, more or less cylindrical, approximately 4.5 x longer than deep and not dorally swollen medially in profile; small insects with body length approximately 3.5mm [ Aleiodes View in CoL part]................................... 9

9 (8) Fore femur extremely robust, less than 2.5 x longer from distal part of trochantellus to apex, than maximally deep ( Fig. 78A View FIGURE 78 )......................................................................... A. globifemurus sp. nov.

- Fore femur extremely robust, more than 3 x longer from distal part of trochantellus to apex, than maximally deep ( Figs 116A View FIGURE 116 , 146C View FIGURE 146 ).................................................................................... 10

10 (9) Mesoscutum (except medio-posteriorly) aciculate, rather shiny with some punctures at bases of setae ( Fig. 116C View FIGURE 116 ) and in profile flat-topped and steep anteriorly ( Fig. 116A View FIGURE 116 ); propodeum and metasomal tergites 1–3 largely black ( Figs 116E,F View FIGURE 116 ).................................................................................. A. nonicones sp. nov.

- Mesoscutum coarsely coriaceous ( Fig. 145D View FIGURE 145 ) and dorsally in lateral profile almost continuously curved ( Fig. 145E View FIGURE 145 ); body entirely yellow ( Fig. 145A View FIGURE 145 )........................................................ A. pseudicones sp. nov.

11 (7) Fore wing vein 1-1A giving rise to a distinct, strongly reclivous, anal cross vein sub-medially ( Fig. 165B View FIGURE 165 ); hind wing vein SR with a submedial angulation, usually giving rise to a distinct cross-vein r; fore wing 2 nd submarginal short and trapezoidal, vein 3-SR shorter than vein r ( Fig. 165A View FIGURE 165 )........................................... A. spurivena

- Fore wing vein 1-1A not giving rise to a distinct and strongly reclivous, anal vein medially ( Figs 106D View FIGURE 106 , 127E View FIGURE 127 ); hind wing vein SR without a submedial angulation and without cross-vein r and fore wing 2 nd submarginal more rectangular, vein 3- SR usually longer than vein r........................................................................ 12

12 (11) Distal part of fore wing subbasal cell glabrous ( Fig. 88G View FIGURE 88 ); fore wing subdiscal cell evenly setose ( Fig. 88G View FIGURE 88 ); distal part of fore wing vein M+CU and vein 1-CU1 strongly thickened near intersection with vein 1-M ( Fig. 88G View FIGURE 88 ); largely bright yellow............................................................................ A. kitchingi sp. nov.

- Distal part of fore wing subbasal cell usually evenly setose OR if glabrous THEN EITHER subdiscal cell largely glabrous (especially on anterior half) AND distal part of fore wing vein M+CU and vein 1-CU1 not strongly thickened near intersection with vein 1-M ( Fig. 96E View FIGURE 96 ); colour variable.................................................... 13

13 (12) Basal 3 metasomal tergites forming a carapace with tergite 3 strongly curved in profile concealing all more distal tergites or virtually so AND postero-laterally emarginate ( Figs 32G,H View FIGURE 32 , 59E,F View FIGURE 59 , 181E,F View FIGURE 181 ); mesosternum aciculate; hind coxa strongly transversely to obliquely sculptured basally; small wasps, body length <4.5 mm .............................. 14

- Basal 3 metasomal tergites not forming a carapace concealing all more distal tergites OR IF more or less forming a carapace concealing subsequent tergites ( Fig. 52A View FIGURE 52 ) THEN tergite 3 not posterolaterally emarginate; other characters variable......................................................................................... 16

14 (13) 3rd metasomal tergite approximately 1.2 x longer than 2 nd medially ( Fig. 32G View FIGURE 32 , 59E View FIGURE 59 ); metasoma tergites 2 and 3 at least cream-white laterally, and the 3 rd posteriorly ( Fig. 32H View FIGURE 32 ); 3rd metasomal tergite with postero-lateral corner strongly produced ventrally ( Figs 32H View FIGURE 32 , 59F View FIGURE 59 ); hind wing with vein m-cu distinct ( Fig. 32F View FIGURE 32 ).............................. 15

- 3rd metasomal tergite shorter than 2 nd medially ( Figs 181E View FIGURE 181 ); metasoma tergites 2 and 3 entirely black; 3rd metasomal tergite with posterolateral corner hardly produced ventrally ( Fig. 181F View FIGURE 181 ); hind wing without vein m-cu.... A. trisphaeropyx sp. nov.

15 (14) 1 st metasomal tergite dark posterolaterally, with paler area of variable extent posteromedially ( Figs 59E,F View FIGURE 59 ); 2 nd metasomal tergites largely cream, dark anterolaterally; fore wing vein 3-SR 1.8–1.9 x 2-SR; lateral margin of syntergite when viewed laterally gently curving near level of 2 nd suture ( Fig. 59F View FIGURE 59 ); occipital carina complete mediodorsally, more or less forming a point ( Fig. 59B View FIGURE 59 )................................................................. A. darlingtonia sp. nov.

- 1 st metasomal tergite dark on anterior 0.8 with poterior margin cream coloured all way to lateral corners, with paler area of variable extent posteromedially ( Fig. 32G View FIGURE 32 ); 2 nd metasomal tergites largely black except laterally; fore wing vein 3-SR 1.5 x 2-SR ( Fig. 32F View FIGURE 32 ); lateral margin of syntergite when viewed laterally sharply angled at level of 2 nd suture ( Fig. 32H View FIGURE 32 ); occipital arina broadly incomplete posteromedially ( Fig. 32C View FIGURE 32 )................................ A. butcheri sp. nov.

16 (13) Hind wing vein 2-SC+R transverse and inclivous ( Figs 29D View FIGURE 29 , 87E View FIGURE 87 ) AND relatively large insects with fore wing length> 6.5 mm AND distal part of fore wing subbasal cell largely glabrous ( Figs 29D View FIGURE 29 , 87E View FIGURE 87 ); fore wing uniformly yellow or with apical 0.7 dark, without clear apex; at least hind tarsus largely black ( Fig. 29A View FIGURE 29 )................................ 17

- Hind wing vein 2-SC+R interstitial ( Figs 34D View FIGURE 34 , 109C View FIGURE 109 ) to longitudinal ( Figs 64D View FIGURE 64 , 65E View FIGURE 65 , 82G View FIGURE 82 ) OR if transverse ( Figs 48A View FIGURE 48 , 179E View FIGURE 179 ) THEN smaller insects, less than 6.2 mm fore wing length OR apical part of fore wing subbasal cell largely setose and at most with a small glabrous oval shape surrounded by dense setosity; wing colour variable, sometimes with clear apex ( Fig. 159A View FIGURE 159 ).................................................................................. 18

17 (16) Fore wing venation largely yellow except parastigma and immediately adjoining vein parts black, wing membrane entirely pale yellow-hyaline ( Fig. 29A View FIGURE 29 ).................................................... A. bobwhartoni sp. nov.

- Fore wing venation in distal 0.7 of wing dark brown, wing membrane entirely pale yellow-hyaline on basal third, grey on distal 0.7 ( Fig. 87A View FIGURE 87 ).............................................................. A. jimwhitfieldi sp. nov.

18 (16) Pronotum largely cream white with deep medial pronope usually bordered by a pair of carinate or subcarinate diverging ridges ( Figs 75D View FIGURE 75 , 140C View FIGURE 140 , 141D View FIGURE 141 , 143A View FIGURE 143 ); metasomal tergite 3 strongly curved in dorsal profile, punctate ( Figs 130G View FIGURE 130 ); flagellum often with cream-white band, the flagellomeres in this part of the antenna with large, densely setose gap ventrally, devoid of longitudinal placode sensilla ( Figs. 4C View FIGURE 4 ) [ Aleiodes coronarius group 2]...................... 19

- Pronotum without deep pronope ( Fig. 48A View FIGURE 48 ); metasomal tergite 3 variable, often more or less straight in dorsal profile and often striate ( Figs 63A,F View FIGURE 63 ); flagellum if with cream-white band, not with large gap separating longitudinal placode sensilla in that region (see Figs 4A,B View FIGURE 4 )........................................................................ 45

19 (18) Barcoding sequence known, molecular characters only.................................................... 20 Molecular data unknown, partial morphological key..................................................... 34

20 (19) Bases 1819–1821 coding for V; male antenna with only 1 to 4 apical black flagellomeres beyond white stripe........ 21

- Bases 1819–1821 coding for I; male antenna with more than 8 apical black flagellomeres........................ 23

21 (20) Bases 1657–59 [TTA] coding for L; base 1686 C...................................... A. molecryptus sp. nov.

- Bases 1657–59 [GTA] coding for V; base 1686 T........................................................ 22

22 (21) Bases 1698 1731 1851 1935 2025 A G A T G....................................... A. penultimoluteus sp. nov.

- Bases 1698 1731 1851 1935 2025 G A G C A.......................................... A. apiconigrus sp. nov.

23 (20) Bases 1549–1551 TTG coding for L, 1882–84 GCT coding for A, 1888–90 TTA coding for L... A. concoronarius sp. nov.

- Bases 1549–1551 GTA or GTG coding for V, 1882–84 coding for I, T or V, 1888–90 ATA or ATG coding for M..... 24

24 (23) Bases 1549–51 ATA or ATG coding for M; bases 1873–75 GTA, GTG or GTT coding for V..................... 25

- Bases 1549–51 GTA or GTG coding for V; bases 1873–75 ATT or ATC coding for I............................ 28

25 (24) Bases 1603–05 AGG coding for S; bases 1816 TTA coding for L............................... A. deathi sp. nov.

- Bases 1603–05 GGG coding for G; bases 1816 GTA or GTT coding for V.................................... 26

26 (25) Bases 1807 –09ATT coding for I; bases 1894–96 GTA coding for V; bases 1972 ATA coding for M................................................................................................. A. angulodeum sp. nov.

- Bases 1807 –09GTT coding for V; bases 1894–96 TTA coding for L; bases 1972 TTA coding for L................ 27

27 (26) Bases 1537–39 ACA coding for T;.................................................. A. adorabelleae sp. nov.

- Bases 1537–39 ATA coding for M; ..................................................... A. gaspodei sp. nov.

28 (24) Bases 1946–48 ATT coding for I; bases 1969–71 GCA coding for A......................... A. pronopus sp. nov.

- Bases 1946–48 GTT or GTC coding for V; bases 1969–71 GTA or GCA coding for V or G respecively............. 29

29 (28) Bases 1732–34 TCT coding for S.................................................................... 30

- Bases 1732–34 GCT coding for A.................................................................... 32

30 (29) Bases 1807–09 ACT coding for T; bases 1884–86 ATT coding for I............................. A. cheni sp. nov.

- Bases 1807–09 ATT coding for I; bases 1884–86 GTT coding for V......................................... 31

31 (30) Bases 1591–93 GGA coding for G; bases 1744 GCT coding for A; bases 1816–18 ATA coding for M................................................................................................. A. chenduplus sp. nov.

- Bases 1591–93 AGA coding for S; bases 1744 ATT coding for I; bases 1816–18 GTA coding for V..... A. opus sp. nov.

32 (29) Bases 1816–18 GTA coding for V................................................ A. propodocarinus sp. nov.

- Bases 1816–18 ATA coding for M................................................................... 33

33 (32) Bases 1882–84 GTT coding for V................................................ A. procoronarius sp. nov.

- Bases 1882–84 ACT coding for T.................................................... A. coronopus sp. nov.

34 (19) Propodeum with well-developed posterolateral lobes, with indication of carina [irregular] up posterior edge of lobe ( Figs 15A View FIGURE 15 , 60C View FIGURE 60 ) AND lateral face of lobe coriaceous and distict from sculpture between it and spiracle; male antennae always with more than 8 black flagellomeres beyond white band.................................................. 35

- Propodeum more or less smoothly rounded in profile ( Figs 43A View FIGURE 43 , 46C View FIGURE 46 ) OR IF with some indication of posterolateral lobes

2. The A. coronarius group is one of the more challenging and while molecular data clearly sort them into putative species we have found no good characters to enable all species to be distinguished morphologically. Antennal coloration features appear useful but are sexually dimorphic in some species, and in at least two of the molecularly defined species, there some males show essentially the typical female pattern. Further, the great majority of specimens have one or both antennae broken at or around the white annellar region. Some of the damage looks like they have been bitten. The white zone also appears to have a glandular region of fine pores on one side in a large gap between the placoid sentillae. then lobe with coarse rugulose sculpure laterally continous with sculpture between lobe and spiracle ( Figs 11C View FIGURE 11 , 43E View FIGURE 43 ); male antenna variable, but sometimes with only 1–4 black flagellomeres beyond white band, or entirely black............ 36

35 (34) Propodeum dorsally almost straight with strongly squared posterolateral lobes ( Fig. 15A View FIGURE 15 )....... A. angulodeum sp. nov.

- Propodeum dorsally distinctly curved in profile anteriorly ( Fig. 60C View FIGURE 60 )............................ A. deathi sp. nov.

36 (34) Prododeal sculpture above lateral propodeal carina when viewed laterally, appearing diagonally striate; male antenna with pale stripe occupying only 2 or 3 flagellomeres and separated from apex by at least 15 flagellomeres [pale stripe almost absent in one specimen seen]; female with mesoscutum largely black ( Fig. 46A View FIGURE 46 ), male with dark marks on lateral lobes and anterior of middle lobe of mesoscutum ( Fig. 47D View FIGURE 47 ).................................... A. concoronarius sp. nov.

- Prododeal sculpture above lateral propodeal carina when viewed laterally, not appearing diagonally striate; male antenna with pale stripe variable but sometimes occupying more than 5 flagellomeres.................................. 37

37 (36) Hind coxa basolaterally rather coriaceous............................................... A. pronopus sp. nov.

- Hind coxa basolaterally rather polished with dense punctation............................................. 38

38 (37) Male [unknown for A. propodocarinus sp. nov.]......................................................... 39

- Female [unknown for A. gaspodei , A. molecryptus , A. adorabelleae spp. nov.]................................ 43

39 (38) Black antennal apex beyond white ring comprising only 1 or 2 flagellomeres ( Figs 108A View FIGURE 108 , 130A View FIGURE 130 ) [unknown for A. cheni sp. nov., A. chenduplus sp. nov.]............ A. molecryptus sp. nov., A. penultimoluteus sp. nov., A. apiconigrus sp. nov.

- Black antennal apex beyond white ring comprising more than 10 flagellomeres ( Figs 11D View FIGURE 11 , 143A View FIGURE 143 ) or antenna entirely black ( Fig. 75A View FIGURE 75 )....................................................................................... 40

40 (39) Mesoscutum extensively black anteriorly ( Figs 11D View FIGURE 11 , 75F View FIGURE 75 )................................................. 41

- Mesoscutum entirely rufous dorsally ( Figs 118F View FIGURE 118 , 140C View FIGURE 140 ) sometimes with black lateral margin ( Fig. 143E View FIGURE 143 )............ 42

41 (40) Antenna entirely black ( Fig. 75A View FIGURE 75 )..................................................... A. gaspodei sp. nov.

- Antenna with white ring ( Fig. 11A View FIGURE 11 )................................................. A. adorabelleae sp. nov.

42 (40) White antennal ring occupying more than 6 flagellomeres ( Figs 140A View FIGURE 140 , 143A View FIGURE 143 ) A. procoronarius sp. nov., A. cheni sp. nov.

- White antennal ring occupying fewer than 6 flagellomeres ( Fig. 118A View FIGURE 118 )..... A. opus sp. nov., A. propodocarinus sp. nov.

43 (38) With lateral margin of mesoscutum and anterior of notaulus nearly black, middle lobe of mesoscutum entirely orange ( Fig. 143E View FIGURE 143 ); midlongitudinal carina well developed on anterior 0.75 of propodeum ( Fig. 143E View FIGURE 143 ).... A. propodocarinus sp. nov.

- Mesoscutum entirely orange, or if anteriorly black then also including anterior of middle lobe; midlongitudinal carina well developed on less than anterior 0.65 of propodeum ( Fig. 43F View FIGURE 43 ).............................................. 44

44 (43) Anterior of mesoscutum largely black ( Fig. 43E View FIGURE 43 ); mediodorsal surface of propodeum rather irregularly transversely sculptured ( Fig. 43F View FIGURE 43 ); median area of metanotum rather large and coriaceous dorsally without midlongitudinal carina or deep pit ( Fig. 43F View FIGURE 43 )..................................................................... A. cheni sp. nov.

- Anterior of mesoscutum entirely orange OR with only small dark marks on anterior of each lobe......... other species

45 (18) Hind coxa with transverse to oblique striation on most of basal half, at least on medial side when viewed oblique dorsal angle ( Figs 3C View FIGURE 3 , 36F View FIGURE 36 , 67E View FIGURE 67 , 119B View FIGURE 119 , 157G View FIGURE 157 ) AND occipital carina complete ( Figs 36D View FIGURE 36 , 189D View FIGURE 189 ) and nearly always more or less forming a point mid-anteriorly ( Figs 67C View FIGURE 67 , 119D View FIGURE 119 , 173D View FIGURE 173 ) AND mesopleuron and mesosternum ventrally never smooth and shiny between punctures ( Figs 36E View FIGURE 36 )................................................................... 46

- Hind coxa more or less evenly coriaceous basally (basal to point of maximum width) especially on dorsal and dorsomedial face, not transversely to obliquely sculptured when viewed oblique dorsal angle ( Fig. 156A), though sometimes pimples of aciculation may form parallel oblique lines ( Fig. 3B View FIGURE 3 ), and posterior striation may extend along dorsal edge of dorso-lateral groove of coxa and sometimes more basally in some Chelonorogas group, but in these the mesopleuron and mesosternum are smooth and shiny between punctures; occipital carina variable............................... 66

46 (45) Antenna with a long, rather well-defined white stripe ( Figs 163A View FIGURE 163 , 189A View FIGURE 189 ); back of head black, the mark separated from eye dorsally by paler zone and produced medio-anteriorly to include stemmaticum ( Fig. 189D View FIGURE 189 ); apex of hind tibia with comb of setae (though these not as modified as in many species); hind tarsus conspicuously pale ( Fig. 189A View FIGURE 189 )............. 47

- Antenna without white stripe ( Figs 105A View FIGURE 105 , 157A View FIGURE 157 ); back of head not black; apex of hind tibia with or without comb of setae; hind tarsus dark or same colour as hind tibia............................................................ 49

47 (46) Mesosoma orange-red except pronotum darker submedially ( Fig. 189A View FIGURE 189 ); metasomal tergite 3 strongly arched dorsally in lateral view, and raised above profile of tergite 2 ( Fig. 189A View FIGURE 189 ); metasomal tergites 1–3 black except for pale cream-coloured mark medio-basally on tergite 2, posteriorly ( Fig. 189F View FIGURE 189 ); hind wing vein 2-1A absent............ A. yalaensis sp. nov. Mesosoma black ( Figs 125E, F View FIGURE 125 , 163C View FIGURE 163 ); metasomal tergites 2 and 3 both more or less flat dorsally in profile; metasomal tergites 1 and 2 entirely, and tergite 3 basally, black ( Fig. 125G View FIGURE 125 ), metasomal posteriorly red; hind wing vein 2-1A present as a tubular spur.................................................................................. 48

48 (47) Tarsal claws with 3 large, well-separated pecten spines; fore wing vein 2-CU1 2.2 times 1-CU1 ( Fig. 163F View FIGURE 163 ); 3 rd metasomal tergite with midlongitudinal carina present (weakly) on basal 0.4 ( Fig. 163G View FIGURE 163 ); antenna becoming pale between flagellomeres 16–20 ( Fig. 163A View FIGURE 163 )......................................................... A. s ongsi sp. nov.

- Tarsal claws with pecten formed of line of many adpressed setae (see Fig. 2C View FIGURE 2 ); fore wing vein 2-CU1 1.6 times 1-CU1; 3 rd metasomal tergite with without midlongitudinal carina ( Fig. 125G View FIGURE 125 ); antenna becoming pale between flagellomeres 23–27 ( Fig. 125A View FIGURE 125 )...................................................................... A. parasongsi sp. nov.

49 (46) Hind tibia with comb of especially modified adpressed setae medially and apically (see Fig. 2F View FIGURE 2 ).................. 50

- Hind tibia without comb of especially modified adpressed setae (see Fig. 2E View FIGURE 2 )................................. 52

50 (49) Claws with pecten of approximately 8 short, robust teeth that are longest near the middle of the line and diverge apically ( Fig. 114E View FIGURE 114 ); metasomal tergites almost entirely black except for small ivory spots near mid posterior margins of tergites 1 and 2 ( Fig. 114G View FIGURE 114 )..................................................................... A. niveni sp. nov. - Claws without conspicuous pecten; metasoma not coloured as above........................................ 51

51 (50) Fore wing vein r approximately 0.5 x length of 3-SR ( Fig. 64D View FIGURE 64 ); propodeum and metasomal tergites 1–3 cream medially, with bold black marks laterally ( Fig. 64E,F View FIGURE 64 ); mesopleuron less strongly sculptured posteriorly, more or less aciculate to fine coriaceous posteriorly.......................................................... A. divergerus sp. nov.

- Fore wing vein r approximately 0.8 x length of 3-SR ( Fig. 36A View FIGURE 36 ); propodeum and metasomal tergites 1–3 cream with large bold black marks medially ( Fig. 36E,F View FIGURE 36 ); mesopleuron coarsely sculptured all over, including right up to pleural suture ( Fig. 36E View FIGURE 36 ).............................................................................. A. calicis sp. nov.

52 (49) Fore wing discal cell extremely long and narrow, maximum length more than 2.9 x height at level of 1-M/1-SR+M junction; metasoma orange with tergite margins narrowly darker ( Fig. 67F View FIGURE 67 ); head with black mark behind eye ( Fig. 67D View FIGURE 67 ); maxillary palp smokey grey ( Fig. 67C View FIGURE 67 ); 1st metasomal tergite less than 1.1 x longer than maximally wide ( Fig. 67F View FIGURE 67 ); hind femora basally cream-white, distal 0.7 black or nearly so ( Fig. 67A View FIGURE 67 )......................... A. elongatus sp. nov.

- Fore wing discal cell of normal proportions, maximum length less than 2.7 x height at level of 1-M/1-SR+M junction ( Fig. 157H View FIGURE 157 ); metasomal tergites either strongly patterned black and cream or entirely yellow-brown; maxillary palp white; 1st metasomal tergite usually more than 1.2 x longer than maximally wide; hind femora variable..................... 53

53 (52) 3 rd metasomal tergite distinctly curved dorsally in lateral profile ( Fig. 126F View FIGURE 126 ), usually distinctly narrowing posteriorly from near midlength ( Fig. 126G View FIGURE 126 ), basally finely longitudinally striate becoming acuculate rugulose on posterior third...... 54

- 3 rd metasomal tergite more or less flat dorsally in lateral profile, not or hardly narrowing posteriorly from near midlength, longitudinal striation often coarser and extending nearly to posterior margin of tergite........................... 62

54 (53) Speculum large, smooth and shiny; dorsal part of mesopleuron with rugulose sculpture; metasoma black with broad cream stripe along full length of tergite 2.............................................. A. rectanguliguttatus sp. nov.

- Specular area entirely sculptured, aciculate; dorsal part of mesopleuron with strongly longitudinal pattern of rugae superimposed on coriaceous/rugulose sculpture; metasoma not as above, usually largely yellow brown with transverse brown bands on at least some tergites ( risaae group)...................................................... 55

55 (54) Baso-lateral lobes of 1st metasomal tergite not angularly produced and usually weakly convex posteriorly, not projecting far beyond the small swelling associated with the spiracle though sometimes extending to spiracle ( Fig. 126G View FIGURE 126 ).......................................................................................... A. parisaae sp. nov.

- Baso-lateral flange of 1st metasomal tergite angularly produced and distinctly concave posteriorly and posterior margin of lobe nearly transverse or even reclivous, projecting far beyond the small swelling associated with the spiracle ( Figs 153F View FIGURE 153 , 157I View FIGURE 157 )........................................................................................... 56

56 (55) Body largely orange-red, head largely black except orbits narrowly red, metasomal tergites 3–5 black ( Fig. 157A View FIGURE 157 ); CO1 without amino-acid deletion (codon at positions 2000–2002).............................. A. rugoscutus sp. nov.

- Body brown yellow, head not black, metasoma not patterned as above; CO1 with amino-acid deletion (codon at positions 2000–2002) [not known for A. orion sp. nov.].......................................................... 57

57 (56) Metasoma entirely brown-yellow ( Fig. 105 View FIGURE 105 )............................................. A. mellificus sp. nov.

- Metasoma distinctly patterned black or dark brown on brown-yellow, yellow or cream ( Figs 41A View FIGURE 41 , 91F View FIGURE 91 , 119C View FIGURE 119 , 153F, G View FIGURE 153 ). 58

58 (57) 2 nd metasomal tergite entirely cream white ( Fig. 91F View FIGURE 91 ); 3rd metasomal tergite basally cream white, apically black ( Fig. 91F View FIGURE 91 )................................................................................ A. lemniscus sp. nov.

- 2 nd metasomal tergite variously patterned with black; 3 rd tergite variable...................................... 59

59 (58) 2 nd metasomal tergite largely cream with posterior corners brown, the mark more or less meating at a point medially ( Figs 153F,G View FIGURE 153 ); 3rd metasomal tergite basally yellow-brown, apically black........................... A. risaae sp. nov.

- Metasomal tergite 2 not as above, EITHER black laterally with continuous cream-coloured central band OR at least with dark subtriangular marks extending to anterior corner of tergite; metasomal tergite 3 EITHER black laterally and posteriorly with cream median mark OR virtually entirely black............................................ 60

60 (59) Metasomal tergite 2 black laterally with continuous cream-coloured central band ( Fig. 41E View FIGURE 41 )........ A. centralis sp. nov.

- Metasomal tergite 2 with dark subtriangular lateral marks meeting posteriorly to enclose a lozenge-shaped cream mark ( Figs 119C View FIGURE 119 , 173G View FIGURE 173 )................................................................................. 61

61 (60) Pterostigma ivory-white; mesoscutum with distinct pale brown marks on middle and lateral lobes ( Figs 119B,D View FIGURE 119 ); propodeum largely foveate-rugose anteriorly............................................... A. orion sp. nov.

- Pterostigma dark grey ( Fig. 173F View FIGURE 173 ); mesoscutum uniformly brown-yellow ( Fig. 173E View FIGURE 173 ); propodeum more rugulose, especially antero-medially (Fig. 1734E)................................................. A. synorion sp. nov.

62 (53) Propodeum largely aciculate on either side of midlongitudinal carina, with transverse striae posteriorly ( Figs 10E View FIGURE 10 , 47C View FIGURE 47 ) 63

- Propodeum largely rugulose on either side of midlongitudinal carina ( Fig. 177F View FIGURE 177 )............................... 64

63 (62) Fore wing vein 3-SR 1.5 x 2-SR; 1 st metasomal tergite 1.2 x longer than apically wide ( Fig.47F View FIGURE 47 ); mesoscutum entirely orange ( Fig. 47E View FIGURE 47 ).................................................................... A. conina sp. nov.

- Fore wing vein 3-SR 2.0 x 2-SR; 1 st metasomal tergite 1.9 x longer than apically wide ( Fig. 10E View FIGURE 10 ); mesoscutum with black marks on lateral lobes ( Fig. 10C View FIGURE 10 ).................................................... A. aciculodeum sp. nov.

64 (62) 1st metasomal tergite without or with only weak basal flange that is not sharply angular (see Fig. 10E View FIGURE 10 ); pronotal collar not strongly producted into a neck ( Fig. 176D View FIGURE 176 )............................................................. 65

- 1st metasomal tergite with well developed flange (see Figs 86F View FIGURE 86 ); pronotal collar developed into a long neck ( Fig. 86E View FIGURE 86 ). 66

65 (64) Metasoma black with contiguous cream medial area from middle of tergite 1 to subposterior part of tergite 3 ( Fig. 61F View FIGURE 61 ); flagellum largely black............................................................... A. definus sp. nov.

- Metasomal pattern less clearly defined, pale area brown yellow; mesoscutum entirely pale ochreous yellow ( Fig. 176G View FIGURE 176 ); flagellum orange up to near apex..................................................... A. thirakupti sp. nov.

66 (45, 64) Fourth metasomal tergite strongly rugose to striate sculptured with at least some longitudinal striation ( Figs 23F View FIGURE 23 , 33F View FIGURE 33 , 42G View FIGURE 42 ), posterolaterally angled, in most females and some males with distinct to strong emargination postero-laterally ( Fig. 168F View FIGURE 168 ), always largely exposed and sometimes completely concealing subsequent tergites AND occipital carina broadly interrupted medially ( Figs 23C View FIGURE 23 , 33D View FIGURE 33 , 1910D) AND pronotum not produced into a neck ( Figs 23D View FIGURE 23 ) AND hind tibia without comb of specialised adpressed setae ( Fig. 2E View FIGURE 2 ); fore wing vein 1-CU1 at least 0.7 x length of 2-CU1 ( Figs 34B View FIGURE 34 , 42A View FIGURE 42 , 136A View FIGURE 136 ); 3rd metasomal tergite with complete, or virtually complete mid-longitudinal carina ( Figs 24E View FIGURE 24 , 33F View FIGURE 33 , 83D View FIGURE 83 , 131E View FIGURE 131 ) [ A. buzurae group that form suspended mummies].......................................................... 67

- Fourth metasomal tergite aciculate or smooth and shiny with more or less dense puncturation, often with microsculpture showing distinct transverse pattern, posterolaterally rounded and usually without emargination postero-laterally, OR if with well-developed rugose, rugulose or longitudinally striate sculpture at least on basal 0.5 ( Figs 12A View FIGURE 12 , 50E View FIGURE 50 , 86G View FIGURE 86 , 111F View FIGURE 111 ) THEN occipital carina complete ( Figs 50D View FIGURE 50 , 86C View FIGURE 86 , 111C View FIGURE 111 ) AND / OR pronotum produced into a neck ( Fig. 12C View FIGURE 12 ) OR apex of hind tibia with comb of specialised adpressed setae (see Fig. 2F View FIGURE 2 ) OR fore wing vein 1-CU1 <0.5 x length of vein 2-CU1; 3rd tergite sculpture variable......................................................................... 86

67 (66) Hind wing with vein m-cu distinct ( Fig. 34B View FIGURE 34 ); codon at positions 1795–1797 coding for F; codon at positions 1807–1809 coding for M.................................................................................... 68

- Hind wing without vein m-cu but vein 1-M usually distinctly bent near apex possibly indicating where vein m-cu could originate (Fig, 117A); codon at positions 1795–1797 coding for L; codon at positions 1807–1809 coding for T.......70

68 (67) Third metasomal tergite with dark, sub-circular marks laterally and tergite 4 with large dark mark mediobasally (see Fig. 443D)......................................................................................... 69

- Third and 4th metasomal tergites without dark marks (see Fig. 138E View FIGURE 138 )........................ A. probuzurae sp. nov.

69 (68) Bases 1552–54 CTT coding for L; bases 1570–72 ATT coding for I; bases 1714–16 ATT coding for I; bases 1804–06 ATA coding for M; bases 1948–50 GCA coding for A.................................................. A. buzurae 3

- Bases 1552–54 ATT coding for I; bases 1570–72 GTT coding for V; bases 1714–16 GTA coding for V; bases 1804–06 TTA coding for L; bases 1948–50 TCA coding for S................................ A. buzuriquadruplus sp. nov.

70 (67) First metasomal tergite in lateral view with dorsal carinae uniting near top of tergite, and dorsal profile distinctly depressed medially ( Fig. 175A,C View FIGURE 175 ); flagellum with fewer than 35 (34) segments; pterostigma more or less entirely pale ( Fig. 175A View FIGURE 175 ); posterior margin of 2 nd metasomal tergite more or less evenly curving near midline ( Fig. 175E View FIGURE 175 ). A. tetrarugulosus sp. nov.

- First metasomal tergite in lateral view with dorsal carinae uniting well below top of tergite, and dorsal profile not depressed medially ( Figs 124A View FIGURE 124 , 131A View FIGURE 131 ); flagellum with more than 35 segments; pterostigma with apical 0.3 to 0.5 dark ( Fig. 23A View FIGURE 23 ); posterior margin of 2 nd metasomal tergite more or less straight near midline ( Figs 23F, H View FIGURE 23 ).............. 71

71 (70) Females with posterior margin of 4th metasomal tergite produced into a pair of points sublaterally (at inner border of posterolateral emargination) ( Fig. 23F View FIGURE 23 , 24A,E View FIGURE 24 ) [typical Vietorogas syn. nov. species key here 4]................... 72

- Females with posterior margin of 4 th metasomal tergite not produced into pair of points ( Figs 131E View FIGURE 131 , 168F View FIGURE 168 )........... 75

72 (71) Fore wing vein 1-CU1 1.6 x m-cu.......................................... A. bachma Long & van Achterberg

- Fore wing vein 1-CU1 <1.5 [1.1–1.4] x m-cu.......................................................... 73

73 (72) Antenna pale yellow basally............................................. A. abditivus Long & van Achterberg

- Antenna virtually entirely dark brown, only marginally paler basally......................................... 74

74 (73) Posterior margin of 3rd metasomal tergite rather sharply angled sublaterally; tooth on 4 th tergite sharper ( Fig. 23F View FIGURE 23 )................................................................................... A. bachmaduplus sp. nov.

- Posterior margin of 3rd metasomal tergite not distinctly angled sublaterally; tooth on 4 th tergite more rounded ( Fig. 24E View FIGURE 24 ).............................................................................. A. bachmatriplus sp. nov.

75 (71) Head posteriorly, with region on either side of occipital carina black-brown ( Figs 42C View FIGURE 42 , 131D View FIGURE 131 , 136C View FIGURE 136 ); mesoscutum boldly marked with black ( Figs 42C View FIGURE 42 , 136E View FIGURE 136 )................................................................. 76

- Head posteriorly uniformly pale ( Fig. 80C View FIGURE 80 ); mesoscutum entirely pale cream-white to brown-yellow ( Figs 35D View FIGURE 35 , 80C View FIGURE 80 , 83A View FIGURE 83 )............................................................................................ 81

76 (75) Mesoscutum dark laterally and along line of anterior part of notauli ( Figs 42C View FIGURE 42 , 131C View FIGURE 131 , 168D View FIGURE 168 ); mesopleuron almost entirely dark brown to black; pterostigma entirely black......................................................... 77

- Mesoscutum with dark marks away from edge of each lateral lobe and a pair of sub-medial longitudinal dark marks on the middle lobe ( Figs 98C View FIGURE 98 , 117C View FIGURE 117 ); mesopleuron dark only anterodorsally; pterostigma basally pale ( Figs 98A View FIGURE 98 , 117A View FIGURE 117 )..... 79

77 (76) Face largely ( Fig. 42B View FIGURE 42 ), lateral lobes of mesoscutum largely ( Fig. 42C View FIGURE 42 ) and mesopleuron to below precoxal suture brown to black ( Fig. 42E View FIGURE 42 ); base 1830 T; base 1569 A............................................. A. chamba sp. nov.

- Face largely, lateral lobes of mesoscutum largely and mesopleuron to below precoxal suture pale brown yellow to pale yellow; base 1830 G; base 1569 G.................................................................... 78

78 (77) Bases: 1577 C; 1795 T;1854 T; 1923 T............................................. A. phantasmatis sp. nov.

- Bases: 1577 T; 1795 C; 1854 C; 1923 C............................................ A. subemarginatus sp. nov.

79 (76) Face with submedial brown marks ( Fig. 137B View FIGURE 137 ); tergite 4 with moderately developed semicircular emargination posterolaterally; 1547 A; 1456 A; 1577 T............................................ A. postmaculus sp. nov.

- Face with continuous brown zone across middle; ( Figs 98B View FIGURE 98 , 117B View FIGURE 117 ) tergite 4 hardly emarginate posterolaterally; 1547 not A; 1456 not A; 1577 C............................................................................. 80

80 (79) Bases: 1547 T; 1556 T; 1854 T; 2050 C............................................. A. occimaculatus sp. nov.

- Bases: 1547 G; 1556 C; 1854 C; 2050 T............................................... A. maculiput sp. nov.

81 (75) Pterostigma unicolourous black ( Fig. 35A View FIGURE 35 )........................................... A. buzuritriplus sp. nov.

- Pterostigma bicolorous or largely white ( Figs 80A View FIGURE 80 )...................................................... 82

82 (81) Fore wing vein 1-CU1 as long as or longer than (1.1x) vein 2-CU1.......................................... 83

- Fore wing vein 1-CU1 shorter than vein 2-CU1 ( Fig. 190A View FIGURE 190 )............................................... 84

83 (82) Metasomal tergites 1 and 4 with large nearly black marks, and tergites 2 and 3 with black lateral circular marks ( Fig. 83B–D View FIGURE 83 ); base 1608 C; base 1632 A; base 1736 G................................. A. helenhippersonsae sp. nov.

- Metasoma almost entirely whitish with anterior half of 1 st tergite, and small lateral spots on tergites 2 and 3 dark grey ( Fig. 80D View FIGURE 80 ); base 1608 T; base 1632 G; base 1736 A........................................ A. griseimaculus sp. nov.

84 (82) 1 st tergite 1.2 x wider posteriorly than long; metasoma white with only dark marking on 1 st tergite ( Fig. 124E View FIGURE 124 )........................................................................................ A. parabuzurae sp. nov.

- 1 st tergite less than 1.1 x wider posteriorly than long ( Fig. 190E View FIGURE 190 ); metasoma variable........................... 85

85 (84) Body largely orange-yellow to brown-yellow with extensive black marks on propodeum and metasoma ( Fig. 33 View FIGURE 33 ), dark mark on tergite 1 enclosing a medial pale area ( Fig. 33F View FIGURE 33 )................................ A. buzuriduplus sp. nov.

- Body largely cream-white with small darker marks antero laterally on 1 st metasomal tergite ( Fig. 190 View FIGURE 190 ) A. zuburae sp. nov.

86 (66) Metasomal tergite 4 exposed, with at least basal half strongly sculptured, rugose to longitudinally striate ( Figs 86G View FIGURE 86 ) AND occipital carina complete ( Fig. 86C View FIGURE 86 ) or at most with very narrow medial break AND hind tibia without comb of adpressed setae apico-medially (see Fig. 2E View FIGURE 2 )................................................................... 87

- Metasomal tergite 4 variably exposed but not coarsely sculptured, usually with fine aciculation ( Figs 19E View FIGURE 19 , 38F View FIGURE 38 , 122F View FIGURE 122 , 128E View FIGURE 128 ), and sometimes with these forming a sub-transverse pattern basally ( Figs 61F View FIGURE 61 , 65A View FIGURE 65 ) OR if with some rugulose to striate sculpture basally THEN occipital carina either medially absent ( Figs 30D View FIGURE 30 , 55D View FIGURE 55 ) OR hind tibia with a comb of adpressed setae apicomedially (see Fig, 2F)........................................................... 106

87 (86) Pronotal collar elongate with broad midlongitudinal ridge that is rather clearly demarked posteriorly ( Fig. 140B View FIGURE 140 ); antennae with approximately 15 white segments preceding approximately 4 apical dark ones ( Fig. 140A View FIGURE 140 ). A. procarinatus sp. nov.

- Pronotal collar not elongate OR if elongate ( Figs 72C View FIGURE 72 , 111D View FIGURE 111 ) THEN more or less flat dorsally, without median ridge ( Figs 74D View FIGURE 74 , 111C View FIGURE 111 ); antennae without white stripe............................................................. 88

88 (87) Pronotum elongated with median length> 0.4 x lateral length of mesoscutum measured back to level of posterior corner of pronotum ( Figs 72C View FIGURE 72 , 111D View FIGURE 111 ); hind wing without trace of tubular vein 2-1A; veins 1-1A and cu-a forming a nearly continuous curve ( Fig. 72E View FIGURE 72 ); back of head strongly narrows such that maximum width across occipital carina <0.62 x maximum width of head ( Figs 12E View FIGURE 12 , 68C View FIGURE 68 ) [ Aleiodes narangae group]....................................... 89

- Pronotum not so elongated, with median length <0.4 x lateral length of mesoscutum measured back to level of posterior corner of pronotum ( Figs 100C View FIGURE 100 , 121C View FIGURE 121 , 149E View FIGURE 149 , 182B View FIGURE 182 ); hind wing usually with small but distinct tubular vein 2-1A; veins 1–1A and cu-a forming more of an angle ( Fig. 30G View FIGURE 30 ); back of head variable.................................. 95

89 (88) Body brown extensively marked with black including along side of mesoscutum, propodeum, tergites 1 and 2 basally and laterally, tergite 3 ( Figs 86F View FIGURE 86 ); 3rd and 4th metasomal tergites distinctly emarginate posterolaterally ( Fig. 185B View FIGURE 185 ); codon at bases 1594–96 coding for T......................................................................... 90

- Body almost entirely honey-brown, at most with small darker brown smudge on lateral part of pronotum, small marks medially on 1 st tergite ( A. narangae ) and small submedian brown marks on 2 nd tergite; 3rd and 4th metasomal tergites virtually straight posterolaterally ( Fig. 185A View FIGURE 185 )); codon at bases 88–90 coding for S (unknown for narangae ).........91

90 (89) Metasoma variable, but marked with black and cream-yellow, at least lateral parts of tergites 3 and 4 cream-yellow ( Fig. 86G View FIGURE 86 ); base 1551 A; 1554 T; base1654 T; base 1626 C; base 1731 A; base 1977 T................. A. incisus sp. nov.

- Metasoma orange yellow with dark brown marks ( Fig. 72F View FIGURE 72 ); base 1551 G; base 1554 A; base 1654 C; base 1626 T; base 1731 G; base 1977 C.............................................................. A. fovodeum sp. nov.

91 (89) 1 st metasomal tergite with distinct hour-glass shaped brown mark anteriorly ( Fig. 111E View FIGURE 111 ); 2 nd tergite with small brown lines submedially on anterior half ( Fig. 111E View FIGURE 111 ); midlongitudinal carina of 3 rd tergite well differentiated, rest of tergum rugulose with longitudinal tendency ( Fig. 111F View FIGURE 111 )......................................................... A. narangae

- 1 st and 2 nd metasomal tergites entirely honey-brown ( Figs 68G View FIGURE 68 , 74E View FIGURE 74 ); midlongitudinal carina of 3 rd tergite not or hardly differentiated ( Figs 68G View FIGURE 68 , 74E View FIGURE 74 ), rest of tergum largely with longitudinal striate sculpture......................... 92

92 (91) Codon at postion 1783–1785 coding for V............................................................. 93

- Codon at postion 1783–1785 coding for M............................................................. 94

93 (92) Base 1592 A; base 1679 A; base 1719 T; base 1730 G; base 2082 C; base 2091 G................ A. valinus sp. nov.

- Base 1592 G; base 1679 G; base 1719 C; base 1730 A; base 2082 T; base 2091 A.............. A. etvalinus sp. nov.

94 (92) Base 1592 A; base 1679 G; base 1730 A; base 1836 G; base 1910 G; base 1993 T; base 2008 G; base 2151 A............................................................................................. A. agagg sp. nov.

- Base 1592 G; base 1679 A; base 1730 G; base 1836 A; base 1910A; base 1993 C; base 2008 A; base 2151 T............................................................................................... A. gaga sp. nov.

95 (98) Hind femur short and deep, approximately 4 x longer (measured from narrowing at apex of trochantellus) than maximally deep ( Fig. 121E View FIGURE 121 ); 4th metasomal tergite distinctly though weakly emarginate posterolaterally; metasoma black with posterior of tergite 1 and whole of tergite 2 cream ( Fig. 121E View FIGURE 121 )............................ A. pallimedius sp. nov.

- Hind femur not so swollen, more than 4.2 x longer (measured from narrowing at apex of trochantellus) than maximally deep ( Figs 149A View FIGURE 149 ); 4th metasomal tergite not emarginate posterolaterally; metasoma not coloured as above.......... 96

96 (95) Metasoma entirely black, with striate-rugose sculpture ( Figs 149F View FIGURE 149 , 182F View FIGURE 182 )..................................... 97

- Metasoma not entirely black ( Fig. 100D View FIGURE 100 ), either brown-yellow ( Fig. 38H View FIGURE 38 ), with pale mid-longitudinal area or with pale mid-section; metasomal sculpture variable............................................................. 98

97 (96) 1 st metasomal tergite with more or less straight sides behind distinct protruding basal lobes ( Fig. 149F View FIGURE 149 ); all legs black ( Fig. 149A View FIGURE 149 ); head black, except orbits dorsally narrowly rufous ( Fig. 149D View FIGURE 149 ); body length> 5 mm ... A. reticulisoma sp. nov.

- 1 st metasomal tergite with inwardly-curving sides from base ( Fig. 182F View FIGURE 182 ), without protruding basal lobes ( Fig. 182D View FIGURE 182 ); all legs yellow ( Fig. 182A View FIGURE 182 ); head red except for stemmaticum ( Fig. 182B View FIGURE 182 ); body length approximately 4 mm ... A. tulipus sp. nov.

98 (96) Malar space extremely long such that distance from lowest part of hypostomal carina (viewed laterally) to bottom of eye greater than 0.7 x height of eye in frontal aspect ( Fig. 100B View FIGURE 100 ); fore wing vein 1-M strongly curved ( Fig. 100A View FIGURE 100 ); mesosoma and metasomal tergites 1 and 2 orange, a least tergites 4–6 black ( Figs 100D View FIGURE 100 )................... A. malarius sp. nov.

- Malar space shorter ( Figs 10B View FIGURE 10 , 26B View FIGURE 26 , 30B View FIGURE 30 , 66B View FIGURE 66 ); coloration not as above; other characters variable................ 99

99 (98) Body entirely brown yellow ( Figs 26A View FIGURE 26 , 30A View FIGURE 30 , 66A View FIGURE 66 ).................................................... 100

- Body with prominent black pattern, metasomal tergite 2 black laterally, cream-white medially ( Figs 112F View FIGURE 112 , 188A View FIGURE 188 ).... 102

100 (99) Hind wing vein M+CU 2 x length of 1-M ( Fig. 30G View FIGURE 30 ); fore wing vein r longer than 3-SR ( Fig. 30G View FIGURE 30 ); basal half of metasomal tergite 4 longitudinally striate; apex of fore wing subbasal cell with lozenge-shaped glabrous area ( Fig. 30G View FIGURE 30 ) ................................................................................. A. brevimedius sp. nov.

- Hind wing vein M+CU less than 1.5 [1.1–1.18] x length of 1-M; fore wing vein 3-SR longer than r ( Fig. 26E View FIGURE 26 ); basal half of metasomal tergite 4 variable; apex of fore wing subbasal cell densely setose.................................. 101

101 (100) Dorsal carinae of 1 st tergite forming a sharp triangular point; 1 st tergite approximately 1.6 x longer than wide posteriorly................................................................................ A. basistriatus sp. nov.

- Dorsal carinae of 1 st tergite trilobed, almost square posteriorly; 1 st tergite approximately 1.15 x wider posteriorly than long ( Fig. 66F View FIGURE 66 )........................................................................... A. eadyi sp. nov.

102 (99) Occiput with strong transverse rugulose sculpture ( Fig. 188D View FIGURE 188 ); occipital carina rounded, broken at extreme centre......................................................................................... A. vietuput sp. nov.

- Occiput coriaceous; occipital carina variable......................................................... 103

103 (102) Occipital carina forming an anteriorly directed point medially ( Fig. 112E View FIGURE 112 )..................... A. nathismus sp. nov.

- Occipital carina not forming a point, evenly rounded or forming rounded lobe ( Fig. 21D View FIGURE 21 )....................... 104

104 (103) Propodeum coarsely rugose, short; 1 st metasomal tergite with strong irreglar longitudinal striation ( Fig. 21A View FIGURE 21 )........................................................................................... A. asperum sp. nov.

- Propodeum finely aciculate with only faint rugosity, elongate ( Fig. 10E View FIGURE 10 ); 1 st metasomal tergite strongly aciculate without strong longitudinal striation ( Fig. 10E View FIGURE 10 )............................................................... 105

105 (104) Propodeum evenly aciculate ( Fig. 10E View FIGURE 10 ); lateral lobes of mesoscutum with dark marks ( Fig. 10C View FIGURE 10 ); occipital carina mediodorsally with a distinct posteriorly directed dip........................................ A. aciculodeum sp. nov.

- Propodeum with distinct (though weak) foveate-rugose sculpture on aciculate background (especially when viewed from posterodorsal angle); mesoscutum without dark marks; occipital carina forming a large anterior lobe A. propodealis sp. nov.

106 (86) Mesosternum, mesopleuron and metapleuron densely deeply punctate and or rugose but smooth and shiny between punctures ( Figs 16E View FIGURE 16 , 52C View FIGURE 52 , 76D View FIGURE 76 , 164C View FIGURE 164 ), though occasionally punctures so dense as to be abutting ( Figs 58C View FIGURE 58 , 96C View FIGURE 96 ); typically large insects, fore wing length> 4.5mm; hind wing subbasal cell largely glabrous on anterior half ( Figs 40D View FIGURE 40 , 164C View FIGURE 164 , 171C View FIGURE 171 ); head black, 2 nd and 3rd metasomal tergites never dark laterally and pale medially [species fall in to concept of Chelonorhogas of European authors]................................................................ 107

- Mesosternum, mesopleuron and metapleuron densely aciculate, coriaceous or granular and not shiny between punctures ( Fig. 113D View FIGURE 113 ); often smaller insects; hind wing subbasal cell variable but usually densely and evenly setose ( Fig. 92C View FIGURE 92 ); coloration variable [species fall in to concept of subgenus Aleiodes of European authors]....................... 114

107 (106) Antescutal depression very well-developed, medially occupying more than half length of pronotum in dorsal view ( Fig. 16B View FIGURE 16 ); coxae red, hind tibia basally white, hind tarsus black ( Fig. 16A View FIGURE 16 ); 1 st metasomal tergite short and broad, approximately 0.8 times as long as maximally wide, dorsope normal, with carinae running posteriorly from fused dorsal carinae markedly diverging posteriorly ( Fig. 16D View FIGURE 16 ); propodeum foveate rugose to rugulose anteriorly; fore wing vein 1- CU1 less than 0.5 x length of 2-CU1 ( Fig. 16A View FIGURE 16 )........................................... antescutum sp. nov.

- Antescutal depression narrow and not occupying more than half mid-longitudinal length of pronotum ( Fig. 71D View FIGURE 71 ); IF hind tibia basally white, THEN hind tarsus white ( Fig. 180A View FIGURE 180 ) OR coxae black; 1 st metasomal tergite longer, at least 0.9 times as long as maximally wide ( Fig. 180F View FIGURE 180 ), dorsope variable; other characters variable............................... 108

108 (107) Mid and lower part of mesopleuron and mesosternum smooth and shiny, sparsely punctate, precoxal sulcus virtually absent, indicated only be a very shallow crease dorsally ( Fig. 52C View FIGURE 52 ); mesosoma red, head and metasoma black ( Fig. 52A View FIGURE 52 )......................................................................................... A. convexus

- Mid and lower part of mesopleuron and mesosternum more densely punctate, precoxal sulcus usually deep and narrow but impressed ( Fig. 71C View FIGURE 71 ) OR if less well defined ( Fig. 58C View FIGURE 58 ), then usually indicated by a broad depressed area; not coloured as above ( Figs 58A View FIGURE 58 , 71A View FIGURE 71 , 76A View FIGURE 76 , 96A View FIGURE 96 , 164A View FIGURE 164 , 180A View FIGURE 180 )........................................................ 109

109 (108) Precoxal sulcus not well-defined, whole lower side of mesopleuron very weakly depressed and similarly densely deeply sculptured as adjacent areas with only slight increase in size of punctures in depressed area ( Fig. 58C View FIGURE 58 ); all coxae black ( Fig. 58C View FIGURE 58 ).......................................................................... A. damus sp. nov. - Precoxal sulcus more or less well-defined, usually narrow, depressed and with differentiated, transversely crenulate, coarser sculpture than adjacent areas ( Figs 56D View FIGURE 56 , 71C View FIGURE 71 , 76D View FIGURE 76 , 164C View FIGURE 164 ); all coxae red or yellow ( Figs 56D View FIGURE 56 , 71A View FIGURE 71 , 76D View FIGURE 76 , 164C View FIGURE 164 )....... 110

110 (109) Metasomal tergites 1 and 2 cream-yellow anteriorly, posteriorly black ( Fig. 71D View FIGURE 71 )............ A. flavostriatus sp. nov.

- Metasomal tergites 1 and 2 not striped, either all black or 1st tergite red and sometimes medial part of 2 nd red...... 111

111 (110) Antenna with sharply-defined white anellus (dark grey in one very small male) ( Fig. 76A View FIGURE 76 )........ A. georgiae sp. nov. Antenna entirely black ( Figs 96A View FIGURE 96 , 164A View FIGURE 164 ) or with diffuse paler (yellow-brown) region over broad media zone ( Fig. 180A View FIGURE 180 ).............................................................................................. 112

112 (111) Propodeum and metasomal tergites entirely black ( Fig. 164A View FIGURE 164 ); fore and mid legs and hind femora largely pale yellow...................................................................................... A. sophieae sp. nov.

- Propodeum and at least 1st metasomal tergite red ( Figs 96F View FIGURE 96 , 180F View FIGURE 180 ); fore and mid legs and hind femora orange-red, red or black.......................................................................................... 113

113 (112) Fore wing vein 1-CU1 less than 0.5 x length of 2-CU1 ( Fig. 180E View FIGURE 180 ); flagellum with pale coloured segments over broad medial part ( Fig. 180A View FIGURE 180 ); hind tarsus cream-coloured ( Fig. 180A View FIGURE 180 ); fore wing distinctly darker subapically ( Fig. 180E View FIGURE 180 ); metasomal tergite 3 more convex dorsally in profile ( Fig. 180A View FIGURE 180 )........................... A. tricoloripes sp. nov. Fore wing vein 1-CU1 longer than 2-CU1 ( Fig. 96E View FIGURE 96 ); flagellum unicolorous black ( Fig. 96A View FIGURE 96 ); hind tarsus black; wings entirely hyaline ( Figs 96A,E View FIGURE 96 ); metasomal tergite 3 less convex dorsally in lateral view ( Fig. 96A View FIGURE 96 ). A. longivena sp. nov.

114 (106) Apex of hind tibia without comb of specialised adpressed setae (see Figs 2E View FIGURE 2 , 29E View FIGURE 29 )............................ 115

- Apex of hind tibia with comb of especially modified and adpressed setae on inner side (and usually also ventrally between hind tibial spurs) (see Fig. 2F View FIGURE 2 )...................................................................... 160

115 (114) Occipital carina complete medio-dorsally though sometimes irregular and if so, then [usually] giving rise to at least a faint midlongitdinal carina running to stemmaticum ( Fig. 100C View FIGURE 100 ) [N.B. complete in A. lobocarinus sp. nov. but easily perceived to be medially absent if head not viewed from anterior-dorsal aspect]; size variable........................... 116

- Occipital carina broadly absent medio-dorsally ( Figs 13D View FIGURE 13 , 56D View FIGURE 56 , 103D View FIGURE 103 ); usually small wasps, body length <6 mm .... 147

116 (115) Ocelli enormous almost touching eyes ( Figs 97B View FIGURE 97 , 104D View FIGURE 104 ); claws with strong pecten of dark, discrete teeth.......... 117

- Ocelli of normal size or conspicuously small ( Figs 18C View FIGURE 18 , 100C View FIGURE 100 , 157D View FIGURE 157 ); claws variable but seldom with such strong pecten............................................................................................... 118

117 (116) Hind wing vein 2-SC+R longitudinal ( Fig. 104A View FIGURE 104 ); fore wing vein 3-SR approximately 1.7 x 2-SR ( Fig. 104A View FIGURE 104 ); antennae with slightly less than basal half black, distal part white to tip ( Fig. 104A View FIGURE 104 ); malar space longer ( Fig. 104B View FIGURE 104 )....................................................................................... A. megaophthalmos sp. nov.

- Hind wing vein 2-SC+R quadrate to sub-transverse ( Fig. 97G View FIGURE 97 ); fore wing vein 3-SR less than 1.4 [1.35] x 2-SR ( Fig. 97G View FIGURE 97 ); antennae with black apex ( Fig. 97A View FIGURE 97 ); malar space shorter ( Fig. 97D View FIGURE 97 )................... A. macroophthalmus sp. nov.

118 (116) Malar space extremely long such that distance from lowest part of hypostomal carina (viewed laterally) to bottom of eye greater than 0.7 x height of eye ( Fig. 100B View FIGURE 100 ); fore wing vein 1-M strongly curved ( Fig. 100A View FIGURE 100 ); mandible twisted such that lower tooth directly behind upper one in frontal aspect ( Fig. 100B View FIGURE 100 ); clypeus protruding; mesopleuron with large speculum................................................................................. A. malarius sp. nov.

- Malar space less than 0.65 x height of eye ( Figs 18B View FIGURE 18 , 49B View FIGURE 49 ); other characters variable........................... 119

119 (118) Antenna with a large part, at least from the middle to near apex, pale though not abruptly white-striped [not known for certain in A. conpectenus sp. nov. but probably so] ( Figs 18A View FIGURE 18 , 171A View FIGURE 171 ) AND 2 nd metasomal tergite shorter than posteriorly wide ( Figs 18G View FIGURE 18 , 49F View FIGURE 49 , 171E View FIGURE 171 ); hind tarsus conspicuously cream or white in females ( Figs 18A View FIGURE 18 , 49A View FIGURE 49 ); telotarsi conspicuously dark compared with other tarsal segments ( Fig. 18A View FIGURE 18 ); midlongitudinal propodeal carina complete, running to to semicircular carina just in front of apodeme, though often somewhat irregular on posterior part of propodeum ( Fig. 18F View FIGURE 18 ); ocelli large, distance across ocellar triangle> 4 x shortest distance between posterior ocellus and eye ( Figs 18C View FIGURE 18 , 49D View FIGURE 49 ); moderately large insects, body length> 5 mm .......................................................... 120

- Antenna entirely black ( Figs 155A View FIGURE 155 , 157A View FIGURE 157 ) OR basally red-brown become black towards apex; metasomal tergite 2 proportions, telotarsus colour, hind tarsus colour and propodeal carina variable; ocelli usually smaller, distance across ocellar triangle <4 x shortest distance between posterior ocellus and eye ( Figs 155D View FIGURE 155 , 157D View FIGURE 157 ); size variable......... 122

120 (119) Claws with pecten of approximately 8 long, reddish, clearly separate teeth along whole length.................. 121

- Claws with pecten of approximately 12 adpressed pale teeth along whole length of basal lobe appearing like a silvery or golden lamella at low magnification (see Fig. 2C View FIGURE 2 ) [Highly sexually dimorphic ( Fig. 171A View FIGURE 171 )].. A. subson sp. nov. [females]

121 (120) Mesosoma black with extensive pattern of cream-white markings on mesoscutum, mesopleuron, metapleuron, propodeum ( Fig. 18D E View FIGURE 18 ); metasomal tergites 1 and 2 cream with black marks ( Fig. 18F View FIGURE 18 )................. A. archicolorus sp. nov.

- Mesosoma entirely black (except pale mark on pronotum) ( Fig. 49C, D View FIGURE 49 ); metasomal tergites 1 and 2 entirely red ( Fig. 49F View FIGURE 49 ).............................................................................. A. conpectenus sp. nov.

122 (119) Hind coxa very coarsely sculpured basally both dorsally and medially ( Fig. 157C View FIGURE 157 ); mesoscutum very coarsely sculptured on lateral and anterior of medial lobe ( Fig. 157E View FIGURE 157 )........................................ A. rugoscutus sp. nov

- Hind coxa smooth, aciculate or fine coriaceous basally both dorsally and medially (see Fig. 3B View FIGURE 3 ); mesoscutum not coarsely sculptured on lateral and anterior of medial lobe, generally coriaceous wth more or less dense small punctures at bases of setae ( Figs 14D View FIGURE 14 , 155D View FIGURE 155 )........................................................................... 123

123 (122) Fore wing vein 1-CU1 very short, approximately 0.27 x length of 2-CU1 ( Fig. 155E View FIGURE 155 ); ocelli very small, maximum diameter of posterior ocellus less than 0.75 x shortest distance between posterior ocellus and eye ( Fig. 155D View FIGURE 155 ); antennae very long, approximately 1.5 x wing length ( Fig. 155A View FIGURE 155 ); mesosoma largely black ( Fig. 156A); metasoma red basally, black apically ( Fig. 156B).............................................................. A. rufomedius sp. nov.

- Fore wing vein 1-CU1 longer, at least 0.35 x length of 2-CU1 ( Fig. 133E View FIGURE 133 ); ocelli usually larger, maximum diameter of posterior ocellus more than 0.8 x shortest distance between posterior ocellus and eye ( Fig. 28E View FIGURE 28 ) OR if ocelli small ( Fig. 133D View FIGURE 133 ) THEN antenna shorter OR body entirely yellow ( Fig. 133A View FIGURE 133 )...................................... 124

124 (123) Hind tarsus conspicuously cream or white ( Fig. 28A View FIGURE 28 ); telotarsi conspicuously dark compared with other tarsal segments ( Fig. 28A View FIGURE 28 ); mesoscutum medio-posteriorly cream coloured, otherwise black ( Fig. 28E View FIGURE 28 ); body longer than 5 [6] mm....................................................................................... A. biscutus sp. nov.

- Hind tarsus not conspicuously white, either same colour as hind tibia or yellow or red ( Figs 14A View FIGURE 14 , 40A View FIGURE 40 , 113A View FIGURE 113 ); telotarsi not conspicuously darker than other tarsal articles ( Fig. 40A View FIGURE 40 ); mesoscutum variable; body length variable.............125

125 (124) Hind wing vein M+CU more than 2 [2.15–2.45] x 1-M ( Fig. 133E View FIGURE 133 ); 2 nd submarginal cell very rectangular, vein 3-SR less than 1.5 x 2-SR; basal lobes of tergite 1 small but angularly produced ( Fig. 133F View FIGURE 133 ); metasomal tergites 1 and 2 very strongly and coarsely longitudinally striate ( Figs 14G View FIGURE 14 , 133F View FIGURE 133 )..................................................... 126

- Hind wing vein M+CU less than 2 [1.1–1.8] x 1-M; 2 nd submarginal cell more elongate; basal lobes of tergite 1 and sculpture of 2 nd and 3 rd tergites variable.............................................................. 127

126 (125) Body largely reddish with metasomal tergites 3–6 black ( Fig. 14A,G View FIGURE 14 ); propodeum very coarsely sculptured though without conspicuous mediolateral protuberances ( Fig. 14F View FIGURE 14 )........................................ A. anguaae sp. nov.

- Body entirely pale yellow ( Fig. 133A View FIGURE 133 ); propodeum less coarsely sculptured and with distinct, though weak, posterolateral angulations ( Fig. 133C,D View FIGURE 133 )............................................................. A. placidus sp. nov.

127 (125) Head usually largely black dorsally except orbits more or less narrowly orange red to cream ( Figs 25C View FIGURE 25 , 40E View FIGURE 40 , 113E View FIGURE 113 ) (though with wider orbit mark in A. protocastaneus sp. nov.; Fig. 144D View FIGURE 144 ); metasoma boldly patterned red or orange and black, tergite 2 unicolorous red ( Figs 25E View FIGURE 25 , 40F View FIGURE 40 , 113A View FIGURE 113 ); occipital carina giving rise to at least a faint midlongitudinal ridge running to stemmaticum ( Figs 144D View FIGURE 144 )....................................................................... 128

- Head yellow to red-brown dorsally except for black stemmaticum OR if somewhat darkened medioposteriorly and/or near occipital carina THEN metasoma not boldly patterned red or orange and black, tergite 2 black laterally and pale medially; occipital carina variable........................................................................... 131

128 (127) Midlongitudinal propodeal carina absent on posterior third; female antenna with fewer than 42 [38–40] flagellomeres.129

- Midlongitudinal propodeal carina complete; female antenna with more than 43 [45–47] flagellomeres.............. 130

129 (128) Metasomal tergite 1 entirely red ( Fig. 113F View FIGURE 113 ); metasomal tergites 4–7 black ( Fig. 113F View FIGURE 113 ); back and top of head largely black dorsally with sharply defined brown-yellow orbits ( Fig. 113E View FIGURE 113 ).............................. A. necsubson sp. nov.

- Metasomal tergite 1 black mark anteriorly enclosing yellow semicircle posteriorly ( Fig. 144E View FIGURE 144 ); metasomal tergites 4–7 entirely orange yellow ( Fig. 144E View FIGURE 144 ); back and centre of top of head dark brown black, yellowish orbits wider and less clearly defined ( Fig. 144D View FIGURE 144 )...................................................... A. protocastaneus sp. nov.

130 (128) Second metasomal tergite 1.1 x wider posteriorly than long, and 1.1 x wider han anteriorly wide ( Fig. 40F View FIGURE 40 ); codon at bases 1594–96 coding for S; codon at bases 1597–99 coding for G; codon at bases 1816–18 coding for V; codon at bases 1990–93 coding for M............................................................. A. castaneus sp. nov.

- Second metasomal tergite parallel-sided and longer than wide ( Fig. 25E View FIGURE 25 ); codon at bases 1594–96 coding for T; codon at bases 1597–99 coding for S; codon at bases 1816–18 coding for M; codon at bases 1990–93 coding for T.......................................................................................... A. basipunctatus sp. nov.

131 (127) Metasoma entirely yellow to brown yellow, at most slightly darkened anteriorly, without obvious median pale line ( Figs 26A View FIGURE 26 , 66A View FIGURE 66 )...................................................................................... 132

- Metasoma dark with a pale median line at least on tergite 2 ( Figs 57F View FIGURE 57 , 148E View FIGURE 148 , 178E View FIGURE 178 )............................ 137

132 (131) Propodeum with strong mediolateral protuberances ( Figs 65C,F View FIGURE 65 ); setosity of distal part of fore wing membrane approximately twice as dense as that of basal cell ( Fig. 65E View FIGURE 65 )................................. A. downeyi sp. nov.

- Propodeum without strong mediolateral protuberances ( Figs 26C View FIGURE 26 , 50C View FIGURE 50 , 54E View FIGURE 54 ); setosity of distal part of fore wing membrane approximately same density as that of basal cell........................................................ 133

133 (132) Body more than 5.5 [6.0-6.6] mm long; 1 st metasomal tergite approximately 1.5 x longer than posteriorly wide; basolateral lobes of 1 st metasomal tergite distinctly protruding, rather angular ( Fig. 50E View FIGURE 50 )................................ 134

- Smaller, body less than 5.5 [3.8-5] mm long; 1 st metasomal tergite approximately 1.1 x wider posteriorly than long; basolateral lobes of 1 st metasomal tergite variable, hardly ( Fig. 120E View FIGURE 120 ) to moderately produced, rounded and posteriorly convex.......................................................................................................................................................................................... 135

134 (133) Occiput evenly aciculate without transverse sculpture ( Fig. 26D View FIGURE 26 ); antennae brown yellow basally becoming black from near the middle ( Fig. 26A View FIGURE 26 )......................................................... A. basistriatus sp. nov.

- Occiput with distinct transverse corruscation ( Fig. 50D View FIGURE 50 ); antennae brown yellow on basal 0.75 becoming black apically ( Fig. 50A View FIGURE 50 )...................................................................... A. constriatum sp. nov.

135 (133) Occiput rather coarsely sculptured with distinct transverse striation superimposed on coriaceous-punctate sculpture ( Fig. 54D View FIGURE 54 ); body almost entirely brown-orange, metanotum darker ( Fig. 54A View FIGURE 54 ); wing membrane brownish and conspicuously darker near veins; 2 nd metasomal tergite 1.2 x wider posteriorly than long ( Fig. 54F View FIGURE 54 ); antenna basally red-brown become black towards apex ( Fig. 54A View FIGURE 54 ); antenna with approximately 38 flagellomeres................. A. corrusciput sp. nov.

- Occiput coreacous with hardly any indication of transverse sculpture ( Figs 110B View FIGURE 110 , 120B View FIGURE 120 ); body pale yellow, metanotum not conspicously darker ( Fig. 110D View FIGURE 110 ); wing membrane clear to pale grey ( Fig. 110A View FIGURE 110 ); antenna with approximately 41 or more flagellomeres [not known for certain in A. mutilus sp. nov.].............................................. 136

136 (135) Occipital carina not or hardly complete mediodorsally where it is weak and rather pointed and confused with adjacent sculpture ( Fig. 120B View FIGURE 120 ); with dark grey marks on pronotum laterally, median part of propodeum and basal and anterolateral part of 1 st metasomal tergite ( Fig. 120A,C,E View FIGURE 120 )......................................... A. paenicarinus sp. nov. - Occipital carina strong and complete medio-dorsally, more or less evenly rounded ( Fig. 110D View FIGURE 110 ); body entirely pale yellow ( Fig. 110A View FIGURE 110 )......................................................................... A. mutilus sp. nov.

137 (131) Ovipositor and ovipositor sheaths strongly upcurved ( Fig. 57A View FIGURE 57 )........................... A. curvicauda sp. nov.

- Ovipositor and ovipositor sheaths not strongly upcurved ( Figs. 127A View FIGURE 127 , 178A View FIGURE 178 )................................ 138

138 (137) Metasomal tergites 1–4 black except for a parallel-sided ivory white mark down middle of tergite 2 and occasionally a very small pale area midposteriorly on tergite 1 ( Figs 148E View FIGURE 148 , 178E View FIGURE 178 )............................................. 139

- Metasoma with more extensive pale marks, at least antero-medial part of tergite 4 with large pale mark ( Figs 127F View FIGURE 127 ).. 140

139 (138) Antenna with fewer than 38 [35] flagellomeres; propodeum rather strongly arched in dorsal profile ( Fig. 148D View FIGURE 148 ); hind femur, tibia and tarsus largely brown yellow ( Fig. 148A View FIGURE 148 ); claws with pecten of 7 or 8 long, fine, pale brown teeth on basal lobe, these splaying and distinctly separated, and the penultimate ones slightly longer (see Fig. 2D View FIGURE 2 )............................................................................................ A. rectanguliguttatus sp. nov.

- Antenna with more than 39 [41] flagellomeres; propodeum shallowly arched in dorsal profile ( Fig. 178C View FIGURE 178 ); hind femur, tibia and tarsus largely black ( Fig. 178A View FIGURE 178 ); claws with at most a few small pecten teeth on basal part of basal lobe, not conspicuous........................................................................ A. tobiasi sp. nov.

140 (138) Midlongitudinal propodeal carina not extending to posterior or propodeum, replaced by transverse crenulae; lateral margin of propodeum rufous, mesosoma without distinct dark line along side ( Fig. 127C View FIGURE 127 ); fore wing vein 1-CU1 0.55 x 2-CU1 ( Fig. 127E View FIGURE 127 ); largely orange yellow, without dark marks on back of head ( Fig. 127D View FIGURE 127 ), lateral parts of thorax or lateral part of propodeum more or less uniformly honey yellow without conspicuous dark line ( Fig. 127A,C View FIGURE 127 )... A. paulmarshi sp. nov.

- Midlongitudinal propodeal carina extending to posterior or propodeum though sometimes irregular posteriorly ( Fig. 47F View FIGURE 47 ); lateral margin of propodeum black, mesosoma with distinct dark line along side; fore wing vein 1-CU1 usually less than x 2-CU1; coloration variable......................................................................... 141

141 (140) Hind femur and tibia patterned, at least apical 0.25 of hind femur dark brown to black ( Figs 47A View FIGURE 47 , 186A View FIGURE 186 )........... 142

- Hind femur and tibia uniformly pale coloured, whitish, yellow or orange yellow ( Figs 81A View FIGURE 81 , 82A View FIGURE 82 , 95A View FIGURE 95 )............ 143

142 (141) Hind femur whitish basally with apical 0.7 dark yellow-brown, hind tibia distinctly banded with narrow white ring near base, followed by two broad dark brown-yellow ( Fig. 186A View FIGURE 186 ); mesoscutum brown with dark marks on each lateral lobe and on medial lobe ( Fig. 186F View FIGURE 186 )........................................................ A. variifemurus sp. nov.

- Hind femur more or less uniformly brown beyond pale base ( Figs 47A View FIGURE 47 ); mesoscutum largely orange brown ( Fig. 47E View FIGURE 47 )...................................................................................... A. conina sp. nov.

143 (141) Mesoscutum with extensive dark markings, nearly black along line of notauli ( Fig. 81D View FIGURE 81 ); face with dark medial mark ( Fig. 81B View FIGURE 81 ); legs rather robust with tarsal articles markedly deepening towards apex ( Fig. 81A View FIGURE 81 ); antenna with more than 37 [41] flagellomeres; body more than 5 [5.8] mm long............................................ A. guidaae sp. nov.

- Mesoscutum and face uniformly yellow or brown yellow ( Fig. 147B,D View FIGURE 147 ) OR if with dark marks, the lines of the notauli not dark ( Fig. 82C View FIGURE 82 ); legs rather less robust ( Fig. 82A View FIGURE 82 ); antenna with fewer than 36 [27–35] flagellomeres; body less than 4.5 mm long....................................................................................... 144

144 (143) Body largely brown yellow with dark brown to black mark on lateral lobes of mesoscutum and metapleuron ( Figs 82A,C View FIGURE 82 )....................................................................................... A. hei sp. nov.

- Body largely cream white with dark brown line laterally including mark on lateral part of back of head, side of pronotum, upper part of mesopleuron and lateral part of propodeum ( Figs 95A,C View FIGURE 95 )...................................... 145

145 (144) Occipital carina strongly arched medio-dorsally into a lobe where it is rather weakly defined ( Fig. 95C View FIGURE 95 ); fore wing vein 1- CU1 less than 0.44 x 2-CU1; antenna with fewer than 29 [27] flagellomeres................. A. lobocarinus sp. nov.

- Occipital carina evenly rounded medio-dorsally or forming a point, not formed into a lobe; fore wing vein 1-CU1 more than 0.45 [0.46–0.5] x 2-CU1; antenna with more than 30 [31–34] flagellomeres.............................. 146

146 (145) Metasomal tergite 4 brown on anterior 0.7 ( Fig. 147F View FIGURE 147 ); occiput coriaceous ( Fig. 147D View FIGURE 147 ); head brown posteriorly but without distinct mark dorsolaterally on occipital carina ( Fig. 147D View FIGURE 147 )................................... A. ptraci sp. nov.

- Metasomal tergite 4 entirely white ( Fig. 79F View FIGURE 79 ); occiput aciculate ( Fig. 79D View FIGURE 79 ); head with small dark mark next to occipital carina dorsolaterally ( Fig. 79D View FIGURE 79 )........................................................ A. glutinum sp. nov.

147 (115) Claws strongly pectenate with 12 or more strong, diverging teeth running from base of basal lobe to base of claw proper; body length 5.5–7.2 mm; females with antenna with large pale band medially and metasomal tergites 3–6 black ( Fig. 171A View FIGURE 171 , right)........................................................................ A. subson sp. nov.

- Claws without conspicuous pecten; smaller wasps; coloration of female not as above......................... 148

148 (147) Metasoma boldly patterned with dark marks ( Figs 19C View FIGURE 19 , 57E, G View FIGURE 57 , 103E View FIGURE 103 , 150C View FIGURE 150 )................................. 149

- Metasoma largely white, without dark markings OR at most with weak grey markings on base and sides of 1 st tergite ( Figs 13E View FIGURE 13 , 44C View FIGURE 44 , 169D View FIGURE 169 )................................................................................ 152

149 (148) Hind coxa largely dark brown to black ( Fig. 19A View FIGURE 19 ); midlongitudinal propodeal carina incomplete, absent on posterior 0.25................................................................................. A. areelucki sp. nov.

- Hind coxa entirely brown yellow ( Figs 103E View FIGURE 103 , 150C View FIGURE 150 ); midlongitudinal propodeal carina complete though may be slighly irregular posteriorly.............................................................................. 150

150 (149) Ovipositor and ovipositor sheaths strongly upcurved ( Fig. 57A,F View FIGURE 57 ); metastoma with tergite 3 brown yellow, darker medially ( Fig. 57F View FIGURE 57 ); pterostigma largely white with narrow grey brown border ( Fig. 57A View FIGURE 57 )............... A. curvicauda sp. nov.

- Ovipositor and sheaths straight ( Fig. 103A View FIGURE 103 ); metasoma not as above, tergite 3 darker laterally ( Figs 103G View FIGURE 103 , 150E View FIGURE 150 ); pterostigma with distal 0.5 dark ( Fig. 103A View FIGURE 103 )........................................................... 151

151 (150) Malar space (distance from lowest part of hypostomal carina to bottom of eye, in frontal aspect) more than 0.6 [0.7] x height of eye ( Fig. 150B View FIGURE 150 ); speculum well developed ( Fig. 150C View FIGURE 150 ); mesosoma largely cream white with dark brown black lateral stripe, metanotum and propodeum (Fig,. 150C)......................................... A. reus sp. nov.

- Malar space (distance from lowest part of hypostomal carina to bottom of eye, in frontal aspect) less than 0.6 [0.5] x height of eye ( Fig. 103B View FIGURE 103 ); speculum absent, specular area entirely aciculate ( Fig. 103E View FIGURE 103 ); mesosoma largely orange yellow with dark brown to black marks on side of pronotum, metanotum and propodeum ( Fig. 103E View FIGURE 103 )..... A. mediomaculatus sp. nov.

152 (148) Fore wing vein 3-SR shorter than vein r ( Fig. 57A View FIGURE 57 )....................................... A. curtivena sp. nov.

- Fore wing vein 3-SR at least 1.2 x longer than vein r ( Figs 13A View FIGURE 13 , 44D View FIGURE 44 )....................................... 153

153 (152) Frons with depressions behind antennal sockets distinctly differentiated from rest of frons and with at least a weak “V” or “Y”-shaped carina in front of anterior ocellus ( Figs 44B View FIGURE 44 , 162D View FIGURE 162 ); ocelli usually relatively large, the maximum diameter of posterior ocellus often greater than shortest distance between posterior ocellus and eye ( Figs 13D View FIGURE 13 , 44B View FIGURE 44 ); pterostigma with apical half dark grey ( Fig. 44D View FIGURE 44 ); propodeum and 1 st metasomal tergite completely yellow or brown yellow ( Fig. 44A View FIGURE 44 ) 154

- Frons more or less flat ( Fig. 170D View FIGURE 170 ) or if with distinct depressions behind antennal sockets then these merging gradually with rest of frons and without “V” or “Y”-shaped carina in front of anterior ocellus; ocelli sometimes with maximum diameter of posterior ocellus less than 0.8 than shortest distance between posterior ocellus and eye (Figs); pterostigma usually completely pale ( Figs 73A View FIGURE 73 , 169A View FIGURE 169 ); propodeum and base of 1 st metasomal tergite usually marked with grey brown ( Fig. 73E View FIGURE 73 )..... 156

154 (153) Codon at 1543–45 coding for A; codon at 1582–84 coding for L; codon at 1594–96 coding for S; codon at 1597–99 coding for S; codon at 1720–23 coding for L; codon at 1807–09 coding for I; codon at 1816–18 coding for V................................................................................................ A. alboluteus sp. nov.

- Codon at 1543–45 coding for S; codon at 1582–84 coding for M; codon at 1594–96 coding for T; codon at 1597–99 coding for G; codon at 1720–23 coding for M; codon at 1807–09 coding for M; codon at 1816–18 coding for I............ 155

155 (154) Codon at 1888–1890 coding for A....................................................... A. codon sp. nov.

- Codon at 1888–1890 coding for T.................................................. A. similicodon sp. nov.

156 (153) Codon at 1732–1734 coding for A; codon at 1795–1797 coding for M; codon at 1816–1818 coding for L; codon at 1819–1821 coding for T; codon at 1884–1886 coding for P; codon at 1900–1902 coding for T.. A. fuscomedius sp. nov.

- Codon at 1732–1734 coding for G or S; codon at 1795–1797 coding for T or I; codon at 1816–1818 coding for I; codon at 1819–1821 coding for I; codon at 1884–1886 coding for S; codon at 1900–1902 coding for M................... 157

157 (156) Codon at 1732–1834 coding for S................................................ A. subfuscomedius sp. nov.

- Codon at 1732–1834 coding for G................................................................... 158

158 (157) Codon at 1795–1797 coding for I................................................... A. paenicarinus sp. nov.

- Codon at 1795–1797 coding for T................................................................... 159

159 (158) Base at 1751 A; base at 1814 G; base at 1929 T........................................ A. mediofuscus sp. nov.

- Base at 1751 G; base at 1814 A; base at 1929 C..................................... A. submediofuscus sp. nov.

160 (114) Fore wing vein r> 1.25 x length of vein 3-SR ( Fig. 30G View FIGURE 30 , 187E View FIGURE 187 ) AND fore wing vein 1-1A medially with distinctly thickened and curved zone ( Fig. 30G View FIGURE 30 , 187E View FIGURE 187 ); occipital carina medially obliterated ( Figs 30D View FIGURE 30 ); ocelli large, distance across ocellar triangle 7 or 8 x shortest distance between posterior ocellus and eye ( Figs 30D View FIGURE 30 , 55D View FIGURE 55 , 177D View FIGURE 177 ); hind wing vein M+CU more than 1.9 x length of 1-M ( Fig. 30G View FIGURE 30 ); basal lobes of 1 st metasomal tergite sharply angularly projecting ( Fig. 115E View FIGURE 115 ).................................................................................................. 161

- Fore wing vein r <1.15 x length of vein 3-SR ( Figs 48E View FIGURE 48 , 107A View FIGURE 107 ); fore wing vein 1-1A without markedly thickened and curved zone medially ( Fig. 106D View FIGURE 106 ); occipital carina, ocelli and hind wing vein M+CU and 1 st tergite variable....... 166

161 (160) 4 th metasomal tergite exposed and distinctly longitudinally striate on basal half medially ( Fig. 55F View FIGURE 55 ); speculum absent, entie aea aciculate.................................................................................... 162

- 4 th metasomal tergite smooth or aciculate (if exposed) without longitudinal striation; speculum variable............ 164

162 (161) Occiput aciculate with quite deep punctures associated with setae; midlongitudinal carina of propodeum complete; colour largely brown yellow ( Fig. 30A View FIGURE 30 ) ..................................................................... 163

- Occiput aciculate-coriaceous ( Fig. 55D View FIGURE 55 ); midlongitudinal carina of propodeum absent posteriorly ( Fig. 55E View FIGURE 55 ); colour largely cream white, centre of propodeum pale brown ( Fig. 55E View FIGURE 55 ).............................. A. cramum sp. nov.

163 (162) Hind wing vein M+CU less than 2.0 [1.9] x 1-M ............................................ A. nivori sp. nov.

- Hind wing vein M+CU more than 2.0 [2.2] x 1-M....................................... A. brevimedius sp. nov.

164 (161) Fore wing vein 1-CU1 0.85 x as long as vein 2-CU1....................................... A. tmaliaae sp. nov.

- Fore wing vein 1-CU1 less than 0.8 x as long as vein 2-CU1 ( Fig. 177A View FIGURE 177 ).................................... 165

165 (164) Fore wing vein 1-CU1 less than half length of vein 2-CU1 ( Fig. 90A View FIGURE 90 ); codon at positions 1795–97 coding for M; codon at positions 367–9 coding for M...................................................... A. lavaeolous sp. nov.

- Fore wing vein 1-CU1 more than half length of vein 2-CU1 ( Fig. 187E View FIGURE 187 ); codon at positions 1795–97 coding for T; codon at positions 367–9 coding for I........................................................ A. vetinarii sp. nov.

166 (160) Tergite 3 entirely smooth and shiny ( Fig. 134G View FIGURE 134 ); legs robust............................... A. polititergus sp. nov.

- Tergite 3 at least coriaceous, usually rugulose or longutudinally striate at least on basal third.................... 167

167 (166) Ocelli minute and ocellar triangle very small, transverse diameter of posterior ocellus less than 0.3 x distance between stemmaticum and occipital carina, and distance across ocellar triangle less than 0.7 (0.68) distance between stemmaticum and occipital carina ( Fig. 107C View FIGURE 107 ); fore wing vein 1-CU1 0.8–1.1 x 2-CU1; basal 3 metasaomal tergites dark brown to black.......................................................................... A. microophthalmos sp. nov.

- Ocelli not minute and ocellar triangle larger, transverse diameter of posterior ocellus more than 0.35 times distance between stemmaticum and occipital carina and/or distance across ocellar triangle more than distance between stemmaticum and occipital carina; fore wing vein 1-CU1 shorter than 2-CU1 OR IF rarely up to 1.1 x 2-CU1 THEN ocelli normal sized OR occiput with distinct wide median furrow; metasoma differently coloured..................... 168

168 (167) Distance across stemmaticum approximately equal to shortest distance between eyes; fore wing grey medially with clear apex ( Fig. 159A View FIGURE 159 ); antennae largely orange yellow becoming dark towards tip ( Fig. 159A View FIGURE 159 )......... A. scottshawi sp nov.

- Distance across stemmaticum less than 0.9 x shortest distance between eyes; forewing not as above, without clear apex ( Figs 37D View FIGURE 37 , 106B View FIGURE 106 ); antenna usually largely black, sometimes with white apex ( Fig. 37A View FIGURE 37 )........................ 169

169 (168) Occipital carina complete medio-dorsally ( Figs 37D View FIGURE 37 , 48B View FIGURE 48 , 77B View FIGURE 77 ), very rarely in species where carina produced anteriorly to form a point, extreme centre may be weak or incomplete ( Fig. 64C View FIGURE 64 )........................................ 170

- Occipital carina distinctly incomplete medio-dorsally, curving anteriorly near level of posterior ocelli and becoming obliterated, line seldom demarked by clear transition in sculpture ( Figs 99B View FIGURE 99 )................................ 199

170 (169) Hind wing vein 2-SC+R distinctly transverse ( Figs 37F View FIGURE 37 , 48A View FIGURE 48 ) AND hind wing subbasal cell entirely glabrous ( Figs 37G View FIGURE 37 , 48F View FIGURE 48 ) AND antenna with white apex ( Fig. 37A View FIGURE 37 ) [not known in glabribasalis sp. nov.]; fore wing 1st subdiscal cell with distinct longitudinal glabrous zone in distal part of cell.................................................. 171

- Hind wing vein 2-SC+R longitudinal or interstitial OR if transverse THEN hind wing subbasal cell almost entirely setose ( Fig. 106C View FIGURE 106 ); fore wing 1st subdiscal cell usually without distinct longitudinal glabrous zone in distal part of cell; antenna and mesonotum variable.......................................................................... 173

171 (170) Mesosoma largely honey-yellow dorsally and ivory to cream ventrally with black lateral line running from side of pronotum, upper part of mesopleuron and side of propodeum ( Fig. 48A,D View FIGURE 48 ).................. A. connudatum sp. nov.

- Mesosoma at least with lateral lobes and anterior of medial lobe of mesoscutum and most of metanotum and propodeum black.......................................................................................... 172

172 (171) Scutellum and posterior posterior of middle lobe of mesonotum cream ( Fig. 77D View FIGURE 77 ); metasomal tergites 1–3 black with triangular cream marks posteriorly on tergites 1 and 2 ( Fig. 77F View FIGURE 77 )........................ A. glabribasalis sp. nov.

- Scutellum and mesonotum entirely black ( Fig. 37C,F View FIGURE 37 ); metasomal tergites 1–3 black with with only small cream mark posteriorly on tergite 2 ( Fig. 37E View FIGURE 37 )........................................................ A. calvus sp. nov.

173 (170) Tarsal claws with pecten composed of line of approximately 10–15, parallel, adpressed teeth giving appearance of a silvery lamella ( Fig. 2C View FIGURE 2 )................................................................................. 174

- Tarsal claws without conspicuous pecten OR if with pecten THEN it is composed of 3 –7 widely separated teeth (see Fig. 2B View FIGURE 2 ) OR composed of 7–10 longer, setiform teeth that diverge, the longer being subapical ( Fig. 2D View FIGURE 2 ).............. 180

174 (173) Occipital carina produced mid-anteriorly to form a point and giving rise to a short carina ( Fig. 129B,F View FIGURE 129 ); codon at position 1831–1833 coding for I.......................................................................... 175

- Occipital carina evenly rounded anteriorly ( Fig. 132D View FIGURE 132 ) or if pointed, not giving rise to a short midlongitudinal carina ( Figs 128C View FIGURE 128 ); codon at position 1831–1833 coding for V..................................................... 176

175 (174) Tergite 3 distinctly finely longitudinally striate on anterior 0.4 ( Fig. 129C View FIGURE 129 ); tergites 1 and 2 anteriorly dark grey-brown ( Fig. 129C View FIGURE 129 ); bases 1704, 2025, 2070 G, G, C respectively................................. A. pectunguis sp. nov.

- Tergite 3 hardly striate on anterior 0.4 ( Fig. 129G View FIGURE 129 ); tergites 1 and 2 anteriorly largely pale ( Fig. 129G View FIGURE 129 ); bases 1704, 2025, 2070 A, A, T respectively....................................................... A. pectunguisella sp. nov.

176 (174) Mesosoma largely cream-white or brown yellow with or without black line running along upper part of pronotum and mesopleuron and along side of propodeum ( Figs 106E View FIGURE 106 , 132C View FIGURE 132 )............................................ 177

- Mesosoma either largely black or largely orange-red ( Figs 38C View FIGURE 38 )........................................... 179

177 (176) Mesosoma entirely cream-white to yellow; metasomal tergite 5 without any dark mark, palest of the three species in this complex ( Fig. 132 View FIGURE 132 )................................................................. A. pinnulae sp. nov.

- Mesosoma largely cream-white with black line running along upper part of pronotum and mesopleuron and along side of propodeum ( Figs 22A View FIGURE 22 , 106E View FIGURE 106 )...................................................................... 178

178 (177) Metasomal tergite 3 with a pair of lateral black marks ( Fig. 22E View FIGURE 22 ); propodeum without midlongitudinal carina, with weak smooth medial line on anterior 0.2 only ( Fig. 22F View FIGURE 22 ); base 1656 G; base 1854 T; base 1894 A........... A. atuin sp. nov.

- Metasomal tergite 3 with a single large transverse black mark ( Fig. 106F View FIGURE 106 ); propodeum with distinct midlongitudinal carina on basal 0.6 ( Fig. 106C View FIGURE 106 ); base 1656 A; base 1854 C; base 1894 G............................. A. mericeti sp. nov.

179 (176) Flagellum black basally becoming white from approximately the midpoint ( Fig. 38A View FIGURE 38 ); mesoscutum with lateral lobes and anterior of middle lobe largely black, posteromedially cream ( Fig. 38D View FIGURE 38 ); codon at position 277–9 coding for M; posterior of metasomal tergite 3 and all of tergite 4 orange............................................. A. canus sp. nov.

- Flagellum entirely black or basally yellow-brown becoming dark or black towards apex ( Fig. 128A View FIGURE 128 ); mesoscutum largely orange-red with propodeum somewhat darker; codon at position 277–9 coding for I; metasomal tergites 3 and 4 entirely black ( Fig. 128F View FIGURE 128 )................................................................. A. pectopulicis sp. nov.

180 (173) Fore tibia long and curved, more than 1.5 x length of fore tarsus ( Figs 122A View FIGURE 122 , 123A View FIGURE 123 ); pronotum laterally with coarse radially arranged rugae medially and posteriorly ( Fig. 123E View FIGURE 123 ); 3 rd metasomal tergite less transverse ( Figs 122F View FIGURE 122 , 123G View FIGURE 123 ) 181

- Fore tibia not especially elongate and more or less straight, less than 1.5 x length of fore tarsus ( Fig. 137A View FIGURE 137 ); pronotum more or less coriaceous posterolaterally without strong radial or parallel pattern of rugae ( Fig. 137F View FIGURE 137 ); 3 rd metasomal tergite more transverse ( Fig. 137E View FIGURE 137 )........................................................................ 182

181 (180) Base 1571 A; base 1625 C; base 1637 G; base 1694 C; base 1733 G........................... A. palmae sp. nov.

- Base 1571 G; base 1625 T; base 1637 A; base 1694 T; base 1733 C........................ A. palmatipes sp. nov.

182 (180) Antenna with more than 49 flagellomeres penultimate flagellomeres (those approximately 10 from apex) approximately 1.5–2 x longer than wide ( Fig. 4B View FIGURE 4 )................................................................. 183

- Antenna with fewer than 48 flagellomeres OR if with approximately 50 flagellomeres THEN penultimate flagellomeres (those approximately 10 from apex approximately 2.5–3 x longer than wide ( Fig. 4A View FIGURE 4 )........................ 186

183 (182) Face largely honey-yellow with small brown or darker mark just below each antennal socket ( Fig. 31B View FIGURE 31 ); codon at positions 1597–99 coding for N; codon at positions 1783–85 coding for I; codon at positions 1882–84 coding for N........ 184

- Face largely brown with clypeus (and sometimes small central area above clypeus), malar region, and narrow line next to eye ivory-white to yellowish ( Fig. 137B View FIGURE 137 ); codon at positions 1597–99 coding for S; codon at positions 1783–85 coding for M; codon at positions 1882–84 coding for A............................................... A. prillae sp. nov.

184 (183) Lateral lobes of mesoscutum dark, middle lobe of mesoscutum with a brown to black mark at midlength ( Fig. 31F View FIGURE 31 ); propodeum more coarsely sculptured; base 1869 C.................................... A. brunniguttatus sp. nov.

- Lateral lobes of mesoscutum variable, but middle lobe without brown transverse mark at midlength ( Fig. 45C View FIGURE 45 ); propodeum more finely sculptured; base 1869 T................................................................. 185

185 (184) Mesoscutum largely honey coloured except narrowly brown-black posterolaterally ( Fig. 45C View FIGURE 45 ); metasomal tergite 2 with a large cream coloured mark medio-posteriorly, often extending to postero-lateral part of tergite and sometimes to midanterior of tergite ( Fig. 45F View FIGURE 45 ); bases 1557, 1830, 1875 and 2079 T, G, C and A respectively....... A. complexus sp. nov.

- Mesoscutum more extnsively dark marked, least with lateral lobes largely brown to black; metasomal tergite 2 with only a small rufous mark medio-posteriorly ( Fig. 84F View FIGURE 84 ); bases 1557, 1830, 1875 and 2079 C, A, A and G respectively............................................................................................ A. herrena sp. nov.

186 (182) Metasoma entirely black ( Fig. 69F View FIGURE 69 ); mesoscutum black except for midposterior part of middle lobe ( Fig. 69C View FIGURE 69 )............................................................................................. A. faciei sp. nov.

- Coloration not as above.......................................................................... 187

187 (186) Third segment of maxillary palp swollen, only 4 x longer than wide ( Fig. 183B View FIGURE 183 ); side of mesopleuron largely evenly foveate-rugose ( Fig. 183E View FIGURE 183 )....................................................... A. turgidipalpus sp. nov.

- Third segment of maxillary palp slender ( Figs 179B View FIGURE 179 ); side of mesopleuron usually aciculate with foveate or rugose sculpture restricted to dorsal parts and precoxal sulcus.................................................. 188

188 (187) Basolateral flange of 1st metasomal tergite not strongly and angularly produced and not concave posteriorly, forming a smooth lobe virtually reaching spiracle ( Figs 154E,G View FIGURE 154 ) AND antenna with more than 40 flagellomeres............. 189

- Baso-lateral flange of 1st metasomal tergite angularly produced and distinctly concave posteriorly ( Fig. 36F View FIGURE 36 ) OR antenna with fewer than 39 flagellomeres................................................................... 192

189 (188) Hind wing vein 2-SC+R strongly transverse ( Fig. 179E View FIGURE 179 ).................................. A. transvena sp. nov.

- Hind wing vein 2-SC+R not transverse.............................................................. 190

190 (189) Metasoma almost entirely dark brown to black except for paler brown to cream coloured area at the base of tergite 2 ( Fig. 154G View FIGURE 154 ); propodeum with very coarse rugose sculpture ( Fig. 154E View FIGURE 154 ) and with distinct, rather ridge-like posterolateral protuberances ( Fig. 154D View FIGURE 154 ); metasomal tergite 3 very shiny with longitudinal striation almost parallel ( Fig. 154G View FIGURE 154 ).......................................................................................... A. rivulus sp. nov.

- Metasoma with far more extensive pale markings ( Figs 9E View FIGURE 9 , 99G View FIGURE 99 ); propodeum less coarsely sculptured ( Fig. 99E View FIGURE 99 ) and without distinct posterolateral ridge-like protuberances ( Fig. 99C View FIGURE 99 ); longitudinal striation of tergite 3 usually distinctly curved and diverging from midline ( Figs 9E View FIGURE 9 , 99G View FIGURE 99 )..................................................... 191

191 (190) Antenna with fewer than 31 (29) flagellomeres; 3 rd metasomal tergite almost entirely black ( Fig. 99G View FIGURE 99 )................................................................................................. A. magratae sp. nov.

- Antenna with more than 32 (34) flagellomeres; 3 rd metasomal tergite almost entirely white, narrowly black laterally ( Fig. 9E View FIGURE 9 ).............................................................................. A. achingae sp. nov.

192 (188) 1 st metasomal tergite with inverted wine-glass shaped black mark.............................. A. calicis sp. nov.

- 1 st metasomal tergite not patterned as above............................................................ 193

193 (192) Propodeum entirely black ( Fig. 174D View FIGURE 174 ).................................................. A. teatimei sp. nov.

- Propodeum pale yellow medially, black laterally ( Fig. 161F View FIGURE 161 ).............................................. 194

194 (193) Frons virtually flat, without distinct depressions behind antennal sockets and without sublateral carina paralleling eye ( Fig. 161C,D View FIGURE 161 )..................................................................... A. sesquipunctatus sp. nov.

- Frons distinctly depressed behind antennal sockets ( Figs 27D View FIGURE 27 , 64C View FIGURE 64 , 92D View FIGURE 92 ) and with sublateral carina paralleling eye and sometimes then curving to form posterolateral border of frontal depressions.................................. 195

195 (194) Mesopleuron including precoxal sulcus more or less evenly aciculate, precoxal sulcus virually absent, indicated by only a weak depression with very faint striae or a very short, narrow line of just 3 or 4 small foveae, area below wings not or hardly longitudinally striate ( Figs 20C View FIGURE 20 , 27C View FIGURE 27 ).......................................................... 196

- Anterior of mesopleuron and precoxal sulcus conspicuousley more coarsely sculptured rugulose to striate, area of mesopleuron below wings longitudinally striate ( Figs 70D View FIGURE 70 , 92C View FIGURE 92 )........................................... 197

196 (195) Episternal scrobe distinct, area above it slightly swollen and specular area with distinctly reduced sculpure ( Fig. 27C View FIGURE 27 ); codon at 1894–96 ATT coding for I; codon at 1972–74 TTA coding for L........................ A. binkyi sp. nov.

- Episternal scrobe virtually absent, area above it not swollen and specular area without differentiated sculpture ( Fig. 20C View FIGURE 20 ); codon at 1894–96 GTT coding for V; codon at 1972–74 TTT coding for F...................... A. ascrobi sp. nov.

197 (195) Hind wing vein m-cu interstitial........................................................ A. liessa sp. nov.

- Hind wing vein m-cu antefurcal ( Fig. 64D View FIGURE 64 )............................................................ 198

198 (197) Hind coxa with trace of fine oblique striation on basodorsal and basomedial surface; base 1901 G. A. divergerus sp. nov.

- Hind coxa without a trace of oblique striation on basodorsal and basomedial surface ( Fig. 70D View FIGURE 70 ); base 1901 A........................................................................................ A. flannelfooti sp. nov.

199 (169) Hind wing without vein m-cu ( Fig. 99F View FIGURE 99 )............................................................... 00

- Hind wing with vein m-cu (see Fig. 154F View FIGURE 154 )............................................................. 205

200 (199) Antenna with fewer than 31 (29) flagellomeres.......................................... A. magratae sp. nov.

- Antenna with more than 31 (33–40; precise number unknown in rincewindi sp. nov.) flagellomeres............... 201 201 (200) Propodeum entirely black ( Figs 63C View FIGURE 63 , 152D View FIGURE 152 , 167E View FIGURE 167 ); codon at 1807–09 coding for V or M; codon at 1825–27 coding for S or V............................................................................................. 202

- Propodeum largely pale, at least broadly anteriorly and almost reaching posterior ( Figs 135D View FIGURE 135 , 160A View FIGURE 160 ); codon at 1807–09 coding for I; codon at 1825–27 coding for T.......................................................... 204

202 (201) Face with large brown mark, occipital carina dorso-laterally brown to dark brown; codon at 1684–86 ciding for I; codon at 1714–16 coding for V; codon at 1789–91 coding for A; codon at 1825–27 coding for S............ A. stohelit sp. nov.

- Face and back of head uniformly pale yellow; codon at 1684–86 ciding for I; codon at 1714–16 coding for V; codon at 1789–91 coding for S; codon at 1825–27 coding for V................................................... 203

203 (202) Codon at 1798–1800 coding for L; codon at 1807–09 coding for M; codon at 310–312 coding for V.................................................................................................. A. dimorphus sp. nov.

- Codon at 1798–1800 coding for F; codon at 1807–09coding for V; codon at 310–312 coding for I. A. rincewindi sp. nov.

204 (201) Antenna with more than 37 [39] flagellomeres; fore wing vein 2-CU1 1.3 x 1-CU1 ( Fig. 160E View FIGURE 160 ); codon at 1657–59 coding for I; codon at 2044–46 coding for I................................................... A. selachiii sp. nov.

- Antenna with fewer than 37 [35] flagellomeres; fore wing vein 2-CU1 1.2 x 1-CU1 ( Fig. 135E View FIGURE 135 ); codon at 1657–59 coding for V; codon at 2044–46 coding for V................................................... A. ponderi sp. nov.

205 (199) 3 rd metasomal tergite of female with broad transverse black band that is produced into a point medioposteriorly ( Figs 39F View FIGURE 39 , 85F View FIGURE 85 , 94F View FIGURE 94 )..................................................................................... 206

- 3 rd metasomal tergite of female variably coloured but not as above......................................... 208

206 (205) Codon at 1816–1818 coding for V.................................................. A. lipwigi sp. nov. (part)

- Codon at 1816–1818 coding for I................................................................... 207

207 (206) 1 st metasomal tergite with a pale median line ( Fig. 39F View FIGURE 39 ); codon at 1825–27 coding for T; codon at 2041–43 GCA coding for A............................................................................... A. caprinus sp. nov.

- 1 st metasomal tergite solid dark brown-black on basal 0.8 ( Fig. 85F View FIGURE 85 ); codon at 1825–27 coding for M; codon at 2041–43 ACA coding for T..................................................................... A. hircus sp. nov.

208 (205) Basal lobes of 1 st metasomal tergite more or less evenly rounded, posteriorly not or hardly emarginate, or if weakly emarginate, more or less straight anteriorly, the steepest angles (i.e. most close to transverse) converging posteriorly at 20–45 degrees ( Figs 94E View FIGURE 94 , 146F View FIGURE 146 , 166F View FIGURE 166 )................................................................ 209

- Basal lobes of 1 st metasomal tergite more strongly and sharply protruding, posteriorly strongly emarginate, the steepest angles (i.e. most close to transverse) converging posteriorly at more than 90 degrees ( Figs 89E View FIGURE 89 , 109E View FIGURE 109 )............. 214

209 (208) Body entirely or almost entirely cream to yellow, at most with a few brown patches on propodeum and 1 st and 2 nd metasomal tergites ( Figs 151E View FIGURE 151 , 172E View FIGURE 172 )................................................................ 210

- Body boldly marked with dark brown or black, tergites 2 and 3 at least with large black lateral stripes ( Figs 93E View FIGURE 93 , 94F View FIGURE 94 ) 211

210 (209) Antenna with fewer than 39 [37–38] flagellomeres; fore wing vein 3-SR less than 1.5 [1.25] x vein r ( Fig. 172A View FIGURE 172 ); hind wing vein 2-SC+R longitudinal...................................................... A. sutthisani sp. nov.

- Antenna with more than 39 [40] flagellomeres; fore wing vein 3-SR more than 1.5 [1.8] x vein r ( Fig. 151A View FIGURE 151 ); hind wing vein 2-SC+R virtually interstitial...................................................... A. ridcullyi sp. nov.

211 (209) Fore wing 2 nd submarginal cell very elongate, vein 3-SR more than 2 (2.3) x length of 2-SR ( Fig. 166E View FIGURE 166 ); middle lobe of mesoscutum with a pair of longitudinal brown lines ( Fig. 166F View FIGURE 166 ).............................. A. stibbonsi sp. nov.

- Fore wing 2 nd submarginal cell not elongate, vein 3-SR less than than 2 (1.5–1.7) x length of 2-SR ( Fig. 146E View FIGURE 146 ); middle lobe of mesoscutum either uniformly pale yellow ( Fig. 146D View FIGURE 146 ) or with a single brown patch ( Fig. 94D View FIGURE 94 )................. 212

212 (211) Pterostigma largely dark grey brown, pale cream brown basally and apically ( Fig. 94A View FIGURE 94 ); fore wing vein 3-SR 1.7 x 2-SR; fore wing veins 2-SR and 2-M forming an angle of approximately 75 degrees................................. 213

- Pterostigma with basal 0.4 pale, distal part dark grey brown ( Fig. 146E View FIGURE 146 ); fore wing vein 3-SR 1.75 x 2-SR ( Fig. 146E View FIGURE 146 ); fore wing veins 2-SR and 2-M forming an angle of approximately 85 degrees ( Fig. 146E View FIGURE 146 )......... A. pteppicymoni sp. nov.

213 (212) Codon at 1792–4 ATA coding for M..................................................... A. lipwigi sp. nov.

- Codon at 1792–4 ACA coding for T............................................... A. lipwigduplicitus sp. nov.

214 (208) 2 nd metasomal tergite approximately 1.35 x wider posteriorly than long ( Fig. 89E View FIGURE 89 ); codon at 1594–96 ATA coding for M; codon at 1606–08 GTT coding for V; codon at 1714–16 GTT coding for V................... A. klopfsteinae sp. nov.

- 2 nd metasomal tergite less than 1.2 (1.15 or less) x wider posteriorly than long ( Figs 51C View FIGURE 51 , 158F View FIGURE 158 ); codon at 1594–96 ACA coding for T; codon at 1606–08 ATT or ATC coding for I; codon at 1714–16 ATT coding for I.................. 215

215 (214) Ocelli rather large, distance across posterior ocelli approximately 4 x shortest distance between posterior ocellus and eye................................................................................................ 216

- Ocelli rather smaller, distance across posterior ocelli approximately 2.5–3.0 x shortest distance between posterior ocellus and eye ( Figs 62B View FIGURE 62 , 101D View FIGURE 101 )......................................................................... 218

216 (215) 3 rd tergite coarsely striate ( Fig. 51B View FIGURE 51 ); codon at position 1930–32 ACA coding for T; codon at 1972–74 TTA coding for L.............................................................................. ... A. contemptus sp. nov.

- 3 rd tergite finely striate ( Figs 158F View FIGURE 158 ); codon at position 1930–32 GCA coding for A; codon at 1972–74 TTC or TTT coding for F......................................................................................... 217

217 (216) Propodeum densely setose with fine rugulose sculpture ( Fig. 158D View FIGURE 158 ); codon at 1720–22 GTA coding for V; codon at position 1807–1809 ATT coding for I................................................. A. sacharissa sp. nov.

- Propodeum less densely setose, with coarser sculpture ( Fig. 109D View FIGURE 109 ); codon at 1720–22 ATT coding for I; codon at position 1807–1809 ACT coding for T............................................................ A. morti sp. nov.

218 (215) Metapleuron cream ( Fig. 62A View FIGURE 62 ); codon at 1972–74 TTA coding for L........................... A. deyoyoi sp. nov.

- Metapleuron brown ( Fig. 101C View FIGURE 101 ); codon at 1972–74 TTC coding for F........................... A. malichi sp. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Braconidae

Genus

Aleiodes

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