Cardiacephala schildi ( Cresson, 1930 ) Ferro & Marshall, 2018

Ferro, Gustavo Borges & Marshall, Stephen A., 2018, A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae), Zootaxa 4429 (3), pp. 401-458 : 446-447

Cardiacephala schildi ( Cresson, 1930) new combination

( Figs. 80–81 )

Plocoscelus schildi Cresson, 1930: 334 .

Description: Body length 7 mm, wing 4 mm. Palpus mostly dark brown, outer face slightly white. Clypeus brown with pale microsetulae. First flagellomere black covered with golden setulae. Fronto-orbital plates brown with a black spot posteriorly. Postpronotal lobe and scutellum brown. Row of katepisternal setae brownish. Mid and hind femora brown, slightly darkened on medial portion. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. Oviscape dark brown, apex black.

Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Frontal vitta with a narrowed depression centrally, reaching ocellar triangle. Inner vertical seta insertion flat, without tubercle. Paracephalon smooth, without swelling (not extending beyond the vertex). Vertex rounded. Line of black supracervical setae strong.

Thorax: Mesonotum with white pruinose and irregularly darkened. Scutellum upturned, triangular. Femora with sparse black setulae. Mid and hind tibia flattened, but sulcus on outer face weak. Wing with stigmatal and discal bands, apex clear with a dark and narrow preapical pigmentation. Crossvein r-m with small dark spot. R4+5 and M1+2 meeting at the wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling.

Abdomen: Tergites with brownish setulae. Female abdomen: Oviscape with pale microsetulae. Single spermathecal duct apically broad, spermatheca cylindrical and elongated. Paired spermathecal duct uniformly thick with an apical swelling before splitting into stems. Paired spermathecal duct stems thin and smooth in basal third, apical two thirds broad and convoluted. Paired spermathecae striate and spherical ( Fig. 81 ).

Type material: Holotype (1 ♂, USNM): COSTA RICA. Higuito, San Mateo. Coll. Pablo Schild (white label). Type No. 27264 (orange label). USNMENT 0 0 677015.

Material examined: COSTA RICA. Puntarenas, Osa Peninsula 2.5 km S Rincon, - 50 m, 8º 42’ 1 N 83º30’50 W, trail nr. station, human dung, 10–11.VIII.2001, S. A. Marshall (2 ♀, DEBU 00 168243, DEBU 00168270); San Jose, San Carlos, Riosparaiso Res. Pecari Stn., 16 km NNE Quepos, 400 m, 9º33’53” N 84º7’32” W, 15.II.2003, S. A. Marshall (1 ♀, DEBU 00205738); San Jose, San Carlos, Riosparaiso Res. Pecari Stn., 16 km NNE Quepos, 400 m, 9º33’53” N 84º7’32” W, 15.II.2003, S. A. Marshall (1 ♀, DEBU 00205735, sequence DEBUA 288–17).

Distribution: Costa Rica.

Comments: This species is probably closely related to C. harenosa , although the female genitalia are unlike any other Cardiacephala . Male genitalia dissections were not done because the holotype is the only known male.

Figures

FIGURES 80–81. Cardiacephala schildi. 80, female, lateral view; 81, female genitalia, ventral view.