Caelostomus latus Kavanaugh and Rainio, 2016

Kavanaugh, David H. & Rainio, Johanna, 2016, Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar, Proceedings of the California Academy of Sciences 63 (7), pp. 201-268 : 207-209

Caelostomus latus Kavanaugh and Rainio , sp. nov.

Figure 4

TYPE MATERIAL.— Holotype ( Figs. 4A–B ), a female, in CAS, labeled: “ MADAGASCAR

Ranomafana NP. Talatakely, Trail C, 350 m. 10.5.2005 ”/ “ HOLOTYPE Caelostomus latus

Kavanaugh & Rainio sp. n. 2015” [red label]. TYPE LOCALITY.— Madagascar, Fianarantsoa Province, Ranomafana National Park .

DERIVATION OF SPECIES NAME.— The species epithet, latus , is the Latin adjective meaning broad or wide, a reference to relatively broad pronotum of the holotype specimen and, presumably, other members of this species.

RECOGNITION.— Size moderate for genus, SBL = 6.3 mm. Based on Jeannel’s (1948) key, this species is a member of subgenus Caelostomus s. str. It’s members ( Fig. 4A ) share with those of other members of the subgenus the following features: head with frons and frontal furrows smooth, impunctate; elytral base distinctly margined from humerus medially to base of stria 3 or 2; elytral striae 6 and 7 deeply impressed throughout, not effaced apically; lateral elytral intervals narrow, moderately convex; apical part of umbilicate series of setiferous pores not inserted in a deep groove; and female apical abdominal ventrite with (two or) three pairs of setae subapically. Within the subgenus, the following features are shared only with members of two species of Jeannel’s convexiusculus group, namely Caelostomus convexiusculus Tschitschérine (1899) and Caelostomus alluaudi Jeannel (1948) : SBL greater than 6.0 mm, elytra short and broad, ratio EL/EW = 1.19, elytral disc convex; and elytral intervals distinctly punctate throughout, including in apical part. The holotype female of C. latus is slightly larger than members of these other two species, has a distinct and broad transverse depression between the eyes seen best in lateral view ( Fig. 4C ) (depression absent from members of the other two species). The pronotum in the holotype of C. latus is much wider than in C. convexiusculus members, slightly wider than in C. alluaudi members, and less depressed anterolaterally than in members of either of the other species; and the pronotal basal foveae are straighter and more sharply defined laterally than in members of the other two species. The elytra are less abruptly rounded laterally and apically than in C. alluaudi members but similar to shape in to those in C. convexiusculus members. Finally, anterior tarsomeres 1 to 3 of the female holotype of C. latus are distinctly toothed medioapically ( Fig. 4D ), a feature not seen in females of the other two species.

GEOGRAPHICAL DISTRIBUTION.— At present, known only from the type locality.

HABITAT DISTRIBUTION.— The unique holotype female was collected by hand in secondary montane forest in the Talatakely area .

Figures

FIGURE 4. Digital images of holotype female of Caelostomus latus Kavanaugh & Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Head, left lateral aspect (arrow points to tranverse depression between eyes); D. Right anterior tarsus, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.