Jurakempynus, Wang & Liu & Ren & Shih, 2011

Wang, Yongjie, Liu, Zhiqi, Ren, Dong & Shih, Chungkun, 2011, New Middle Jurassic Kempynin Osmylid Lacewings from China, Acta Palaeontologica Polonica 56 (4), pp. 865-869 : 866

Genus Jurakempynus nov.

Type species: Jurakempynus sinensis sp. nov., see below.

Etymology: From Jura and kempynus, referring to age of the specimen (the Middle Jurassic). Gender is masculine.

Diagnosis.—Forewing: membrane with many fuscous fragmentary spots; costal cross−veins forked distally; one cross−vein sc−r1 close to the wing base, oblique; some fuscous spots resembling cross−veins distributed between Sc and R1; CuA forked in middle, and each branch with complicated pectinate branches; CuP forming some distal complicated branches; A1 well−developed, forming many pectinate branches. Hindwing: membrane hyaline, with fuscous spots on the outer margin; MP region expanded, forming two rows of cells; MP 2 with a thin additional vein at base; CuA long, CuP as long as the half of CuA, with 3–4 pectinate branches.

Remarks.— Jurakempynus has forked costal cross−veins in forewing, which can differentiate it from most other subfamilies (Protosmylinae, Spilosmylinae, Stenosmylinae , Porisminae, Eidoporisminae and Gumillinae ). The only subfamilies that Jurakempynus is likely assigned to are Osmylinae and Kempyninae . However, the new genus exhibits some kempynine−like features that are distinctly different from Osmylinae : MP space relatively expanded in hindwing, forming 2–3 rows of cells ( MP space generally slightly broadened, but never forming multiple rows of cells in Osmylinae ); MP 2 branching complicatedly in distal half of hindwing ( MP 2 commonly simple, only with some distal simple branches in Osmylinae ); base of MP 2 with an additional vein (absence of the additional vein in Osmylinae ). In Kempyninae three extant genera ( Kempynus , Clydosmylus , and Australysmus ) have the similar MP region in the hindwing, and MP region in Clydosmylus forms many rows of cells, highly simliar to Jurakempynus . MP 2 forms complicated distal forks, which is an autapomorphy of Kempyninae ( New 1989). Based on aforementioned characters that Jurakempynus shares with Kempyninae , we consider Jurakempynus should be assigned to Kempyninae instead of erecting a new high level taxon.