Ischnosoma quadriguttatum quadriguttatum ( Champion, 1923 )

Kocian, Matúš & Schülke, Michael, 2018, New species and records of Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) from Nepal, Zootaxa 4442 (4), pp. 501-522 : 516-518

Ischnosoma quadriguttatum quadriguttatum ( Champion, 1923)

Figs 28 , 45 .

NEPAL: Baglung: Dhaulagiri, Baglung Lekh , 10–15 km NW Baglung, 2350–2550 m , 28°18'36''N, 83°31'56''E, 10.–12.V.2004, leg. J. Schmidt, 10 exs. (cKle, cSch); Bhojpur: Valley NW Phedi, 1900 m, 25.V.1997, leg. W. Schawaller, 1 ex. (SMNS); Dailekh: Talpokhari S Dailekh, 1800 m, 29.V.1988, leg. W. Schawaller, 1 ex. (SMNS); Dhawalagi: Annapurna South Himal, SE of Narcheng, left riverside of Bele Khola , 2300 m , 28°30'40''N, 83°41'33''E, 25./ 26.V.2001, leg. G. Hirthe, 1 ex. (cHir); Dolakha: Rolwaling Himal, Lapchi Kang, range below Ting Sang La, 2300–2500 m, 1.IX.1999, leg. J. Schmidt, 1 ex. (NME); Dolpa–Mustang–Myagdi: right banks of Lethe Khola near Lethe, 2400 m, 5.–7.V.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); S slope, Dhaulagiri mountains, N Banduk, 2400–2600 m, 28°28'35''N, 83°35'05''E, 8.V.2009, leg. J. Schmidt, 8 exs. ( NME) ; Dhaulagiri mountains, upper Marang Khola valley., 28°29'50''N, 83°27'37''E, 2500–2700 m, 16.V.2009, leg. J. Schmidt, 1 ex. ( NME), 1 ex. (cSch); Dhaulagari Himal, S-slope, N Banduk village, 1900–2300 m , 28°27'22''N, 83°35'13''E to 28°28'07''N, 83°35'10''E, 6.V.2009, leg. J. Schmidt, 4 exs. (NME), 1 ex. (cSch); SW Dhaulagiri Himal, Dhara Khola Vali, 28°30'36''N, 83°18'16''E, 1900 m, 21.–22.V.2012, leg. J. Schmidt, 6 exs. (NME); S slope, Dhaulagari mountains, above Pathlekharka , 2500–2700 m, 28°32'19''N, 83°29'25''E, 12.V.2009, leg. J. Schmidt, 2 exs. ( NME) ; SW Dhaulagiri, Maraini , 2400–2800 m, 11.V.2012, 28°31'07''N, 83°15'49''E, leg. J. Schmidt, 2 exs. ( NME) ; Gorkha: Darondi Khola, above Barpak, 3300– 3000 m, 11.VIII.1983, rhododendron forest, leg. J. Martens & W. Schawaller [243], 1 ex. (SMNS); Darondi Khola, below Barpak to Doreni, 1100– 900 m, 12.VIII.1983, forest remnants, leg. J. Martens & W. Schawaller [246], 1 ex. (SMNS); Buri Gandaki, Nyak to lower Chuling Khola valley, 2450–2870 m, 2.VIII.1983, Pinus excelsa , mountain pasture, leg. J. Martens & W. Schawaller [227], 1 ex. ( SMNS) ; same data, but 2270–2450 m, 1 ex. ( SMNS) ; Ilam: road Ilam to Birtamode, Karphok, 1700 m, bushes, agricultural area, 2.IV.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); Mai Pokhari, 2100–2200 m, 9.– 10.IV.1988, Castanopsis forest remnant, leg. J. Martens & W. Schawaller [319], 1 ex. (SMNS); Kaski: above Dhumpus, Berlese, 2100 m, 8.–10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Dhumpus, Laubwald, 2100 m, 8.–10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Pothana, 2000 m, 27.– 29.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); Pokhara, souther bank of Phewa Lake, creek, sifted, 800–900 m, 8.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Annapurna region , Khare to Pothana, rhododendron forest, 1950–2000 m, 10. V. 2001, leg. G. Hirthe, 1 ex. (cHir); Annapurna mountains , Tadapani to Chuile, 2450– 2550 m, 4.V.1999, leg. C. Krüger & G. Hirthe, 1 ex. (cHir); Lalitpur: Mount Phulchoki , 1800–2000 m, 25.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. ( SMNS) ; above Godawari, Kunako Khola, sifted, 1750 m, 6.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Lamjung: Nawronkot near Larjung, Thakhola area , 2900–3000 m, leg. H. Franz, 1 ex. ( NHMW) ; Manang : Annapurna mountains, Bagarchhap–Temang, 2400–2700 m, 17.IV.1999, leg. C. Krüger & G. Hirthe, 1 ex. (cHir); Panchthar: Paniporua, 2300 m, 16.–20.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [328], 1 ex. ( SMNS) ; Parbat: Annapurna mountains, Bhickok to Deurali , 2100 m, 7.V.1999, leg. C. Krüger & G. Hirthe, 3 exs. (cHir, cSch); Annapurna region, West Mardi Himal, above Deurali, 2100–2200 m , 28°18'57''N, 83°49'59''E, 10. V. 2001, leg. G. Hirthe, 4 exs. (cHir, cSch); Rasuwa: Bagmati province, Gul Ghanjyang, 2600 m, 6.IV.1981, leg. I. Löbl & A. Smetana, 1 ex. ( MHNG) ; Sankhuwasabha: above Pahakhola, 2600–2800 m, 31.V.–3.VI.1988, Quercus semecarpifolia, Rhododendron , leg. J. Martens & W. Schawaller [404], 3 ex. (SMNS); Arun Valley, between Mure and Hurure , 2050–2150 m, 9.–17.VI.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [412], 2 ex. ( SMNS); N Furure, floodplain near creek , 27°30'44''N, 87°16'14''E, 2100 m, 7.XII.1998, sifted, leg. M. Hartmann, 1 ex. (NME); Arun Valley, W above Tashigaon , 2500–2700 m, 26.– 27.V.2014, 27°36'33''N, 87°13'28''E, leg. J. Schmidt, 1 ex. ( NME) ; Arun Valley, near Tashigaon , 2200–2400 m, 12.+ 28.–29.V.2014, 27°37'N, 87°14'E, leg. J. Schmidt, 2 ex. ( NME) ; Sindhupalchok: Barahbise region , Ting – Sang – La, leg. H. Franz, 1 ex. ( NHMW) ; Solukhumbu: below Pangum, 2500 m, 14.–15.V.1997, leg. W. Schawaller, 3 exs. (SMNS); Umg. Sutje near Lughla, Khumbu, leg. H. Franz, 2 exs. (NHMW); Surkhet: Bheri Khola bridge, 500 m, 24.–25.V.1998, leg. W. Schawaller, 1 ex. ( SMNS) ; N Surkhet, 1600–2000 m, 28.V.1998, leg. W. Schawaller, 1 ex. (SMNS); Syangja: Manaslu mountains, Dudh Pokhari Lekh, upper Dardi Khola valley, 2600– 2300 m, 15.–17.IV.2003, leg. J. Schmidt, 2 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, upper Phulinagiri Madi, 2500 m, 19.–21.IV.2003, leg. J. Schmidt, 10 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, below Helam Pokhari, 2000 m, 22.IV.2003, leg. J. Schmidt, 1 ex. (NME); Manaslu mountains, S of Bara Pokhari, 2300 m, 8.IV.2003, leg. J. Schmidt, 24 exs. ( NME, cSch); Manaslu mountains, Bara Pokhari lekh, Chandi Khola valley , 11.–12.IV.2003, 2000–2300 m, leg. J. Schmidt, 1 ex, ( NME) ; Manaslu mountains , S of Bara Pokhari, 2000 m, 7.IV.2003, leg. J. Schmidt, 1 ex. ( NME) ; Bara Pokhari Lekh , 2500 m, 2.IV.1999, leg. G. Hirthe & C. Krüger, 2 exs. (cHir, cSch); Taplejung: confluence of Kabeli and Tada Khola, 1000 m, 23.–25.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [344], 5 exs. ( SMNS) ; ascent to pasture Lassetham from Omje Khola, 2400–3150 m, 6.V.1988, mixed Quercus - Tsuga - Rhodonendron forest, leg. J. Martens & W. Schawaller [357], 1 ex. ( SMNS).

Remarks. I. quadriguttatum quadriguttatum is widespread in Southeast Asia. In Nepal, it is the most common species and present everywhere ( Fig. 45 ). A second subspecies, I. quadriguttatum japonicum Kocian, 2003 was described from Japan.

Figures

FIGURE 45. Distribution of Ischnosoma quadriguttatum quadriguttatum (Champion) in Nepal (material examined and literature records).

FIGURES 25–31. Extruded internal sac of aedeagus from lateral view (25–27), apical portion of male sternite VII with position of dark long setae (28–30), apical portion of male sternite VIII (31). Ischnosoma jumla sp. n. (25), I. nepalicum (26), I. nepalense (27), I. quadriguttatum quadriguttatum (28), I. travei (29, 31), I. gratiosum (30). Scale bars: a (Figs 28–31), b (Figs 25–27).