Oryctophileurus Kolbe, 1878

Perger, Robert & Grossi, Paschoal Coelho, 2013, Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini), ZooKeys 346, pp. 1-16 : 2-4

Oryctophileurus Kolbe, 1878

Type species.

Phileurus nasicornis Burmeister, 1847 (original combination) (Fig. 3).

Species of Oryctophileurus are distinguished from other Phileurini by a combination of the following apomorphic characters: a single cephalic horn with lateral carina, pronotal cavity with ocellate punctures and two teeth or tubercles close to the anterior pronotal margin, teeth vertically positioned at about the same level as the outer eye margins.

Taxonomy and discussion.

The following characters were cited by Endrödi (1985) for Oryctophileurus , Amblyodus , Microphileurus Kolbe, Metaphileurus Kolbe, Trioplus Burmeister, and Goniophileurus Kolbe and are here not considered as generic: outer side of mandible tridentate; antenna 10-jointed; elytra with punctate rows; proleg sexually monomorphic, protarsus not thickened in male. Oryctophileurus and Amblyodus are distinguished from the other mentioned genera by large horns, a distinctly developed and posteriorly carinate pronotal cavity and four external teeth on the protibia ( Endrödi 1985; Ide 1998). However, in Amblyodus the two cephalic horns lack the lateral carina, the punctures in the pronotal cavity are predominately fused to short, transverse ridges, and the teeth close to the anterior pronotal margin are absent. Except for the horn number, these generic differences between Oryctophileurus and Amblyodus have not been recognized by previous workers.

Based on the combination of apomorphic characters of Oryctophileurus that lacks in Amblyodus we suggest retaining the genus Oryctophileurus .

Key to species of Oryctophileurus