Harpactea dufouri ( Thorell, 1873 )

Hernández-Corral, Jesús, 2017, The spider family Dysderidae in the Balearic Islands, Zootaxa 4329 (4), pp. 375-391 : 386

Harpactea dufouri ( Thorell, 1873)

Figs 6 , 15–16

Harpactea dufouri Thorell, 1873: 561 (descr. ♂); Denis 1961: 238, fig. 1 (descr. ♂); Brignoli 1978: 114, figs 2–3 (descr. ♂); Pons 1993a: 338; Vadel et al. 2006: 79; Pons & Palmer 1996: 38; Bosselaers & Van Keer 2016: 4.

Harpactea corticalis ; Orghidan et al. 1975: 19; Gines 1982: 60 (misidentifications).

Previous records in the Baleares. Cabrera: Caló dels Palangrer ( Pons 1993a); Can Feliu ( Pons 1993a; Canal de s’Aigo ( Pons 1993a); Monument al Francesos ( Pons 1993a); N’Ensiola ( Pons 1993a); Penyal Blanc ( Pons 1993a); Port de Cabrera ( Pons 1993a) .

Formentera: Without precise locality ( Thorell, 1873).

Majorca: Cala Pi ( Orghidan et al 1975., sub H. corticalis ); Ca’n Sion forest ( Orghidan et al. 1975, sub H. corticalis ); Cueva del Puente ( Orghidan et al 1975., sub H. corticalis );Galilea ( Brignoli 1978; Orghidan et al. 1975, sub H. corticalis ); Illetes ( Orghidan et al. 1975, sub H. corticalis ); Magaluf ( Orghidan et al. 1975, sub H. corticalis ); Manacor, Cova des Pont ( Gines 1982, sub H. corticalis ); Manacor, Cova des Xots (Vadel et al. 2006); Palma de Mallorca ( Orghidan et al. 1975, sub H. corticalis ); s’Albufeira Parco National ( Pons 1993b); Valldemosa ( Brignoli 1978).

Menorca: Without precise locality ( Pons 1991); San Cristobal ( Denis 1961).

New material examined. Ibiza: Ses Feixes (38°54’53.45N0°27’0.41’’E), 5 ♂♂ 7 ♀♀, pitfalls in maquis, 28.XII–8.I.2010 (CJH).

Majorca: Albufera Parque Natural (39°47'53''N3°6'27''E), 10m, 2 ♂♂, litter at border of salt marsh, 3.IV.2003 (CRB); S’Albufera PN (N 39°33' 51''N3°0' 30''E), 10m, 2 ♂♂ 2 ♀♀, litter in Pinus forest, 21.III.2015 (CRB); Andratx, Col de Sa Gramola (39°39.290'N 2°24.507'E), 80m, 1 ♂, forest clearing, 26.III.2015 (CRB); Aranjassa SW, road to Llucmajor (39°31′15″N 2°49′28E), 150m, 5 ♂♂ 3 ♀♀, pitfalls in Pistacea maquis, 20–26.III.2015 (CRB); Banyalbufar (39°40'20''N2°29'52''E), 150m, 6 ♂♂ 6 ♀♀, litter in Platanus forest, 1.IV.2003 (CRB); Cala San Vicente (39°55’10’’N13°3’16’’E), 5 ♂♂ 3 ♀♀, 3–9.III.1971, D. J. Clark leg. (BMNH); Coll de Soller (39°47.231'N2°47'825'E), 520m, 1 ♂ 5 ♀♀, litter in Quercus forest, 24.III.2015 (CRB); Estellencs (39°39'12.9''N 2°29'1.0''E), 150 m, 1♂, ground layer near stream, 20.IV.2013 (CJL); Estellencs East. (39°39.342'N 2°29.244'E), 180m, 1 ♂ 3 ♀♀, degraded Pinus forest, 26.III.2015 (CRB); Felanitx E., Can Gelat (39°26'50''N3°14'41''E), 80m, 1 ♀, stones in open Pinus forest, 4.IV.2003 (CRB); Felanitx, Puig Sant Salvador (39°27'20''N3°11'7"E), 490m, 1 ♂ 1 ♀, stones in Quercus maquis, 4.IV.2003 (CRB); Palma de Mallorca, Castillo de San Carlos (39°32'47.4 ‘‘N2°37'16.8''E), 16 m, 1♀, pine grove, 29.IV.2014 (CJL); Montuiri E., Puig de Sant Miquel (39°33' 51 ‘‘N3°0' 30'E), 250m, 3 ♀♀, sieving litter in open planted forest, 23.IX.2014 (CRB); Pollença, Puig de Santa Maria (39°51'36'N 3°5'55''E), 180m, 6 ♂♂ 5 ♀♀, seeving litter in Quercus forest, 21.III.2015 (CRB); Pollença, Cala Boquer (39°56'N3°8'E), 1 ♂ 1 ♀, 10–12.III.1971, D. J. Clark leg. (BMNH); Port de Pollença (39°54'46 ‘‘N3°5'24''E), 45m, 5 ♀♀, litter in small Quercus forest, 22.III.2015 (CRB); Puig de Inca (39°43'45''N2°57'23''E), 280m, 2 ♀♀, stones in open Quercus forest, 3.IV.2003 (CRB); Puig de Randa (39°31'36''N2°55'30''E), 520m, 1 ♀, stones in maquis, 5.IV.2003 (CRB); idem, 3 ♀♀, 23.IX.2014, 1 ♀, 30.IX.2014 (CRB), 2 ♂♂, 27.X.2014 (CJL), 1 ♂, 20.III.2015 (CRB); road Sant Elm-La Trapa (39°35'11''N2°21'25''E), 350m, stones in Pinus forest, 1 ♂, 1.IV.2003 (CRB); Sa Rápita W., Sa Punta Negra (39°21'22''N2°54'43''), 10m, 1 ♂ 1 ♀, litter along small lagoon, 5.IV.2003 (CRB); Sant Elm (39°34' 40.3''N' 2°21' 34.6''E), 30m, 6 ♀♀, along riverbed, 30.X.2014 (CJL); Valldemossa S., Torrent d’Avall (39°42'38''N2°37'18''E), 420m, 4 ♂♂ 4 ♀♀, stones in Quercus forest, 2.IV.2003 (CRB).

Comments. The female of this species was only very recently described ( Bosselaers & Van Keer 2016). Drawings of both vulva and the male bulb are presented here (figs 15–16). The lateral oblique protrusions seen on the vulva are not present in all the females inspected.

Distribution and habitat. This is the only Harpactea species present in the Balearic islands. It is found in deciduous and pine forests and other shaded or semi-shaded habitats with presence of leaf litter. Citations from Majorca of Harpactea corticalis (Simon, 1882) by Orghidan et al. (1975) and ( Gines 1982) are considered misidentifications (see below). Recently, the species has also been cited in Catalonia (Bosselaers & Van Keer 2016).

Figures

FIGURES 1 – 7. Fig. 1. Dysdera arnedoi Lissner new species. Female, dorsalateral view. Figs 2 – 3. Dysdera crocata C. L. Koch (from St Elm). 2. Male, dorsal view; 3. Female, dorsal view; Figs 4 – 5. Dysdera lata ReUss, 1834; 4. Male, dorsal view; 5. Female, dorsal view. Fig. 6. Harpactea dufouri (Thorell, 1839); Male, dorsal view. Fig. 7. Parachtes riberai Bosmans new species; Male, dorsal view.

FIGURES 8 – 16. Figs 8 – 14. Dysdera arnedoi Lissner new species; 8. Prosoma, dorsal view; 9. Left chelicera, ventral view; 10. FemUr IV, dorsal view; 11. MetatarsUs and tarsUs I, lateral view; 12. Tarsal claw, pectination only drawn for one claw; 13. VUlva, dorsal view; 14. VUlva, ventral view. Figs 15 – 16. Harpactea dufouri (Thorell, 1873). 15. Male palp, retrolateral view (from Estellencs); 16. VUlva, dorsal view (from Palma, Castilla de San Carlos). Scale bars: 8: 2 mm, 9: 0.5 mm, 10, 11: 1 mm, 12: 0.1 mm, 13 – 16: 0.2 mm.