Euscorpius kabateki, Kovařík & Šťáhlavský, 2020

Kovařík, František & Šťáhlavský, František, 2020, Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia, Euscorpius 315, pp. 1-37 : 16-22

Euscorpius kabateki sp. n.

( Figures 80–114 , 184 , Table 2 )

http: //zoobank. org / urn: lsid: zoobank. org: act: A813698C- 63F9-40B2-B2EA-45DE501FF7D4

Euscorpius sicanus (in part): Parmakelis et al. 2013: fig. 1 (Locality 60), fig. 2–3 (FESP5).

Euscorpius sicanus complex (in part): Fet et al., 2018: 129, fig. 3.

TYPE LOCALITY AND TYPE REPOSITORY. Greece, Mt. Parnassos , Ski Center Parnassos, 38°32.4‘N 22°34.8‘E, 1800 m a. s. l. ; FKCP.

TYPE MATERIAL. Greece, Mt. Parnassós , Ski Center Parnassou, 38°32.4'N 22°34.8'E, 1800 m a. s. l., 2♂ 4♀1♀ juv. (holotype and paratypes), 5– 6. VI.2007, leg. P. Kabátek, 6. VI.2007, 2♀1♀ juv. (paratypes), leg. V. Ryjáček , FKCP.

ETYMOLOGY. The species epithet is a patronym honoring Petr Kabátek, the collector of holotype of the new species.

DIAGNOSIS (♂ ♀). Total length 28–37 mm. Color reddish yellow. Pedipalp patellar external trichobothria numbers: 5–6 eb, 4 eb a, 2 esb, 4 em, 4 est, 7 et, ventral aspect of patella with 9–11 trichobothria. Pectinal teeth number 9–10 in males, 7–9 in females. Chelicerae yellow, without reticulation. Males with pedipalp fingers type C, females with fingers undulate. Dorsal metasomal carinae on III–IV segments granulated. Dorsolateral metasomal carinae on II–IV segments absent. Ventrolateral metasomal carinae on II–IV segments present and smooth. Metasoma V ventrally with median carina indicated. Chela length/width ratio, 2.3 in males and 2.4–2.5 in females. Metasoma IV length/width ratio, 2.1–2.2 in males and 1.7–1.8 in females. Telson length/depth ratio, 2.2–2.3 in males and 3.2–3.3 in females.

DESCRIPTION (♀ ♂). Total length about 28–37 mm in both sexes. The habitus is shown in Figs. 80–83 . For position and distribution of trichobothria of pedipalps, see Figs. 107–113 . Sexual dimorphism, see below in the description.

Coloration ( Figs. 80–83 ). Base color uniformly reddish yellow to orange. Chelicerae yellow, without reticulation.

Carapace and mesosoma ( Figs. 80–88 ). Carapace finely granulated with several smooth areas; carinae are absent. The anterior margin of the carapace is almost straight. Carapace with two lateral eyes. Tergites finely granulated, without developed carina. Tergite VII lacking lateral and median carinal pairs. Sternites III–VII smooth and lustrous, lacking lateral and median carinae. Stigmata small, narrow ellipical. Pectinal teeth number 9–10 (1 x 9, 3 x 10) in males and 7–9 (2 x 7, 3 x 8, 7 x 9) in females, fulcra are present. Pectines have three marginal lamellae and 4–5 middle lamellae.

Metasoma and telson ( Figs. 91–98 ). Metasoma very sparsely hirsute. Metasoma, mainly in dorsal aspect and whole metasoma V very finely granulated. Dorsal metasomal carinae on II–V segments distinctively irregularly granulated; dorsolateral carinae absent; ventrolateral carinae present, smooth on segments II–IV and granulated on segment V. Metasoma V ventrally granulated with median carina developed, on metasoma I–IV ventral median carina is absent. Anal arch with small pigmented granules. Telson smooth, elongate in females and swollen in males, with annular ring indicated in females and developed in male. Aculeus short more curved in male.

Pedipalps ( Figs. 99–114 ). Pedipalps very sparsely hirsute. Patella with 26–27 (5–6 eb [8 x 5, 12 x 6], 4 eb a, 2 esb, 4 em, 4 est, 7 et) external and 9–10 (2 x 9, 15 x 10, 3 x 11) ventral trichobothria. Chela with 4 ventral trichobothria, of which V 4 is located on external surface near Eb 1. Femur granulated; patella rather smooth. Femur has granulated developed carinae; ventroexternal carina is incomplete. Patella with 5 complete carinae including irregular wide externomedian. DPS well developed. Manus dorsally with fine solitary granules, almost absent in females; median carina missing, other carinae are smooth. Male with pedipalp fingers type C, female with fingers undulate.

Legs ( Figs. 89–90 ). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules.

Measurements. See Table 2 .

AFFINITIES. This species belongs to the subgenus Euscorpius Thorell, 1876 , s. str.; it is morphologically similar to E. sicanus (C. L. Koch, 1837) from Italy (Sicily), which differs by having darker coloration. The presence of more than 4 (usually 5) trichobothria in eb series is a signature trait of a monophyletic ‘ Euscorpius sicanus complex’ (see Fet et al., 2003; Parmakelis et al., 2013).

COMMENTS. Fet et al. (2018: 129) mentioned that “‘ Euscorpius sicanus complex’ representatives are found across Central Greece, including Mt. Parnassos”.After E. sicanus was recently restricted to its type locality of Sicily ( Tropea, 2017), the Greek populations of this diverse complex remained unnamed. DNA data are available for this species in Parmakelis et al. (2013) under FESP5 (Locality 60; a specimen was collected by J. O. Rein in Lilaia, Mt. Parnassos).

DISTRIBUTION. Greece, Mt. Parnassos ( Fig. 184 ).

Figures

Figure 184. Map of distribution of five species described here.

Figures 80–83: E. kabateki sp. n. Figures 80–81. Male holotype in dorsal (80) and ventral (81) views. Figures 82–83. Female paratopotype in dorsal (82) and ventral (83) views. Scale bar: 10 mm.

Figures 84–90: E. kabateki sp. n. Figures 84, 86–87, 89–90. Male holotype, carapace and tergites I–IV (84), coxosternal area and sternites III–V (86), left legs III–IV, retrolateral aspect (89–90 respectively), and chelicerae and eyes with part of carapace (87). Figures 85, 88. Female paratopotype, carapace and tergites I–III (85), coxosternal area and sternites III–IV (88).

Figures 91–98: E. kabateki sp. n. Figures 91–94. Female paratopotype, telson lateral (91), and metasoma and telson lateral (92), dorsal (93), and ventral (94) views. Figures 95. Male paratype, telson lateral. Figures 96–98. Male holotype, metasoma and telson lateral (96), dorsal (97), and ventral (98) views. Scale bars: 5 mm (92–94, 96–98).

Figures 99–106. E. kabateki sp. n., male holotype, pedipalp segments. Chela dorsal (99), external (100) and ventral (101) views. Patella dorsal (102), external (103) and ventral (104) views. Trochanter and femur dorsal (105) and ventral (106) views.

Figures 107–114. E. kabateki sp. n., female paratopotype, pedipalp segments. Chela dorsal (107), external (108) and ventral (109) views. Patella dorsal (110), external (111) and ventral (112) views. Trochanter and femur dorsal (113) and ventral (114) views. Trichobothrial pattern is indicated by white circles.