Trichodromeus trilobatus Shavrin, 2021

Shavrin, Alexey V., 2021, Four new species of the genus Trichodromeus Luze, 1903 from Nepal and Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), Zootaxa 4991 (2), pp. 353-362 : 360-361

Trichodromeus trilobatus Shavrin , sp.n.

( Figs. 4 , 12–13 )

Type material examined: Holotype, ♂ [dissected]: ‘ PAKISTAN: Azad | Jammu & Kashmir, NW | Junkar, 4000m, 1.–10.VIII.2003 ’ <printed>, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed>, ‘ HOLOTYPE | Trichodromeus | trilobatus sp.n. | Shavrin A. V. des. 2021’ <red, printed> ( CSC).

Paratype: 1 ♂, 5 ♀: same data as the holotype (1 ♂, 1 ♀: CS; 4 ♀♀: CSC). All paratypes with red printed label: ‘ PARATYPE | Trichodromeus | trilobatus sp.n. | Shavrin A. V. des. 2021’.

Description. Measurements (min-max; n=7): HW: 1.04–1.22; HL: 0.85–1.01; OL: 0.23–0.31; LT: 0.23–0.27; AL (holotype): 3.76; PL: 1.07–1.21; PWmax: 1.25–1.41; PWmin: 0.93–1.09; ESL: 1.74–2.02; EW: 1.82–1.93; MTbL (holotype): 1.50; MTrL (holotype): 0.69 (MTrL 1–4: 0.37; MTrL 5: 0.32); AW: 1.76–1.94; AedL: 1.30; BL: 6.20–8.50 (holotype: 6.35).

Body, mouthparts, antennae and legs black; apical part of apical segment of maxillary palpi, apical part of tibia and tarsi yellow-brown. Head with moderately coarse, transverse microsculpture, finer in middle; microreticulation of pronotum as that on head, finer and denser in medioapical and basal portions, some specimens without meshes in mediobasal third. Habitus as in Fig. 4 .

Head 1.2 times as broad as long, with strongly convex portions between ocelli and eyes; anterio-median impression moderately deep and wide; temples about as long as longitudinal length of eyes; interocellar depression deep, moderately narrow, subrectangular, separated from infraorbital ridges by deep, strongly convergent latero-anteriad anteocellar foveae, almost reaching level of apical third of eyes. Ocelli small, located slightly below level of posterior margins of eyes. Punctation fine and sparse, somewhat larger and denser on inraorbital ridges. Antennae reaching middle or apical third of elytra when reclined; basal antennomere moderately wide, more than twice as long as broad, antennomere 2 significantly shorter and narrower than basal antennomere, 3 distinctly longer than 2, 4–9 slightly longer than 3, 10 slightly shorter than 9.

Pronotum slightly broader than long, from widest anterior portion gradually narrowed posteriad, anterior margin rounded, not protruded anteriad; middle portion with moderately deep and narrow longitudinal depression, sometimes connected with small transverse mediobasal depression in mediobasal third, two paratypes with indistinct, moderately long irregular impressions on each side of longidudinal depression in middle. Punctation denser and coarser than that on head, sparser in mediobasal third.

Elytra about as broad as long, slightly widened posteriad; hind margins straight. Punctation slightly larger and sparser than that on pronotum, coarser on prescutellar portion.

Abdomen about as broad as elytra, with two very large transverse tomentose spots in middle of abdominal tergite IV and two small spots in middle of abdominal tergite V. Punctation very fine and dense.

Male. Protarsomeres 1–4 very wide. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII somewhat straight. Median lobe wide and subparallel in middle, slightly widened in preapical third and narrowed toward rounded apex, with long and moderately wide dorsal plates, slightly curved latero-apically on each side of apical portion of median lobe; parameres distinctly exceeding apex of median lobe, with long apical setae ( Fig. 12 ). Lateral aspect of aedeagus as in Fig. 13 .

Female. Protarsomeres 1–4 narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded.

Comparative notes. Based on the general shape of the body, T. trilobatus sp.n. is similar to T. jaegeri sp.n. and T. kaliyuga sp.n., from which it can be distinguished by the larger and somewhat darker body and different shape of the apical part of the median lobe.

Distribution. The species is known only from the type locality in Azad Kashmir, Pakistan.

Etymology. The specific epithet is derived from the Latin tri (three) and lobatus, -a, -um (lobed), referring to the shape of the apical portion of the median lobe of the aedeagus.

Figures

FIGURES 1–4. Habitus of Trichodromeus: 1—T. acutipenis (holotype), 2—T. jaegeri (paratype), 3—T. kaliyuga (holotype), 4—T. trilobatus (paratype). Scale bar: 1.0 mm.

FIGURES 12–13. Aedeagus of Trichodromeus trilobatus (holotype): 12—parameral view, 13—lateral view. Scale bar: 1.0 mm.