Therates vitalisi ida Mandl, 1954

Matalin, Andrey V. & Wiesner, Jürgen, 2023, Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae), Zootaxa 5256 (5), pp. 401-433 : 413-414

publication ID	https://doi.org/10.11646/zootaxa.5256.5.1
publication LSID	lsid:zoobank.org:pub:E5B9708E-D626-4923-825A-A9FD59AE4024
DOI	https://doi.org/10.5281/zenodo.7758908
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scientific name	Therates vitalisi ida Mandl, 1954
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Therates vitalisi ida Mandl, 1954 View in CoL

Figs 19–21 View FIGURES 15–28 , 47 View FIGURES 29–49 , 71, 72 View FIGURES 65–72 , 103, 104 View FIGURES 99–114 , 167 View FIGURES 152–174 .

Therates fruhstorferi ida Mandl, 1954: 159 View in CoL (Type locality— China, Kuatun , Fukien).

Therates fruhstorferi ida Mandl, 1954 View in CoL — Mandl 1955: 335; Cassola 1985: 510; Hua 2002: 5.

Therates fruhstorferi vitalisi W. Horn, 1913 View in CoL — Wiesner 1988: 40, 1992: 92; Lorenz 1998: 38, 2005: 117; Puchkov & Matalin 2003: 115.

Therates vitalisi W. Horn, 1913 View in CoL — Wang 2009: 110; Putchkov & Matalin 2017: 245; Wiesner 2020: 95.

Type material. HOLOTYPE, ♂ (by original designation)—“Kuatun (2300 m), 27.40n. Br., 117.40ö. L., J. Klapperich, 25.6.1938 (Fukien)” [printed violet label], “Type Therates, Fruhstorferi, Ida m, Ing. K. Mandl ” [printed and handwritten red label], “Holotypus” [printed red label], “ Therates fruhstorferi , ida Mandl” [typed white label], “ MUSEUM KOENIG, BONN ” [printed orange label] ( ZFMK). GoogleMaps PARATYPES: 1♀ —“ Kuatun (2300 m) 27.40n. Br., 117.40ö. L., J. Klapperich, 30.5.1938 (Fukien)“ [printed violet label], “ CoType Therates Fruhstorferi Ida m., Ing. K.Mandl “ [printed and hand-written red label], “ Coll. Ing. K. Mandl ” [printed light blue label]; GoogleMaps 1♀ —“KUATUN, FUKIEN, China, 6.8.46, (TSCHUNG SEN.)” [printed and hand-written white label], “ParaType Therates Fruhstorferi , ssp. Ida m., Ing. K.Mandl “ [printed and hand-written red label], “ Coll. Ing. K. Mandl ” [printed light blue label]; GoogleMaps 1♀ — “KUATUN, FUKIEN, China, 21.7.46, (TSCHUNG SEN.)” [printed and hand-written white label], “ParaType Therates, Fruhstorferi ssp., Ida m., Ing. K. Mandl ” [printed and hand-written red label], “ Coll. Ing. K. Mandl ” [printed light blue label] (all NHMB) GoogleMaps .

Addition material. CHINA: 1♀ —Kuatun, Fukien /Tschung Sen/, 6 Sept. 1946, coll. Klapperich ( JW) .

Reference. CHINA, Zhejiang Prov.: Wuyanlin ( Wang 2009).

Diagnosis. This subspecies is distinguished from the nominotypical subspecies by distinctly transverse labrum in male—LW/LL = 1.14 vs. LW/LL = 1.0–1.08 ( Fig. 71 View FIGURES 65–72 vs. Fig. 65, 69, 70 View FIGURES 65–72 ), less serrated antennomeres 9–10, as well as red-brown or brown fore- and middle femora.

Redescription. TL = 11.8 mm in male, 12.3–13.0 mm (mean = 12.78 mm, n = 4) in females.

Head shining blue with violet reflection; orbital plates indistinctly striated in posterior third; frons smooth, slightly elevated, wide semi-circular (see above), practically vertical, slightly expanded forward (see at side), frontal sulci deep and poorly convergent in anterior 2/3, clearly divergent in posterior third; occiput slightly concave in anterior margin. Mandibles in male brown underside, yellow-brown topside with light-brown teeth and apical molar; in females black except yellow-brown basal third of topside. Labial palpi yellow-brownish, except brown apical palpomeres; palpomeres 1 and 2 of maxillary palpi light brown, palpomeres 3 and 4 dark brown. Antennae do not extend posteriorly to the shoulders, scape brown on anterior side and black on posterior side; antennomeres 2–5 black with indistinctly blue tinge and brown apices, antennomeres 9 and 10 in male with serrated anterior lower margins; in female less delated with slightly protruding margins ( Fig. 47 View FIGURES 29–49 ). Labrum distinctly transverse in male( Fig. 71 View FIGURES 65–72 ), LW/LL = 1.14, indistinctly transverse in females ( Fig. 72 View FIGURES 65–72 ), LW/LL = 1.0–1.07 (mean = 1.05, n = 4), blackbrownish with medium-sized or small apical yellow spot separated from central apical teeth.

Pronotum shining blue with light violet tinge; indistinctly transverse, PW/PL = 1.0–1.07 (mean = 1.03, n = 5), apical lobe practically equal wide throughout; thorax shining blue with violet reflection.

Fore and middle femora red-brown or brown with blue-tinged base and knees; hind femora black with blueviolet lustre except pale-yellow base; fore and middle tibiae red-brown; hind tibiae black or brown-black, all tibia with blue tinge; tarsi black with light blue tinge; HTbL/HTaL = 1.21–1.27 (mean = 1.24, n = 5).

Elytra black-blue with purple reflection, practically sub-parallel, EL/EW = 1.98–2.1 (mean = 2.02, n = 5); punctuation in basal half except humeral area deep and regular, then shallow and sparse, apical quarter practically smooth; scutellum black with blue tinge; apical margin indistinctly cut. Elytral pattern presented by small basal portion of humeral lunule, small basal dot poorly visible in some specimens and elongated slightly oblique central dot ( Figs 19, 21 View FIGURES 15–28 , 103, 104 View FIGURES 99–114 ).

Aedeagus with short sharply curved blunt apex and distinct lateral carinae ( Fig. 167 View FIGURES 152–174 ); AL = 2.9, EL/AL = 2.52.

Distribution. CHINA: Fujian, Zhejiang Provinces ( Fig. 184 View FIGURES 184 ).

References

Cassola, F. (1985) Memoire sur l'Eurychile di Franco Andrea Bonelli e attuali conoscere sul genre Therates Latreille (Coleoptera, Cicindelidae). Bollettino del Museo Regionale di Scienze Naturali, Torino, 3 (2), 499 - 514.

Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597

Hua, L. - Z. (2002) List of Chinese Insects. Volume II. Zhongshau (Sun Yat - sen) University Press - Huayu Nature Book Trade Co. Ltd, Guangzhou China, 612 pp.

Lorenz, W. (1998) Systematic list of extant ground beetles of the word (Insects Coleoptera Geadephaga : Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). First Edition. Published by the author, Tutzing, Germany, 503 pp.

Lorenz, W. (2005) Systematic list of extant ground beetles of the Word (Insecta Coleoptera Geadephaga : Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 530 pp.

Mandl, K. (1954) Zur Kenntnis der Cicindeliden (Col.) Sud-Chinas. Bonner Zoologische Beitrage, 5, 157 - 161.

Mandl, K. (1955) Zur Kenntnis der Cicindeliden (Col.) Sud-Chinas (III Teil). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen, 6, 334 - 340.

Puchkov, A. V. & Matalin, A. V. (2003) Subfamily Cicindelinae Latreille, 1802. In: Lobl, I & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Appolo Book, Stenstrup, pp. 99 - 118.

Wang, Y. - P. (2009) Cicindelidae. In: Wang, Y. - P. (Ed.), Insecta of Wuyanlin, Zhejiang and analysis of its forest health. Science Press, Beijing, p. 110. [in Chinese, with English summary]

Wiesner, J. (1988) Die Gattung Therates Latr. und ihre Arten. 15. Beitrag zur Kenntnis der Cicindelidae (Coleoptera). Mitteilungen der Munchner Entomologischen Gesellschaft, 78, 5 - 107.

Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt, Checklist of the Tiger Beetles of the World. Vol. 27. Beitrag zur Kenntnis der Cicindelidae. Verlag Erna Bauer, Keltern, 364 pp.

Wiesner, J. (2020) Checklist of the tiger beetles of the world. 2 nd Edition. Edition winterwork, Borsdorf, 540 pp.

Figures

FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png.

FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels.

FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales.

FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype.

FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes

FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm.