Geodromicus (s.str.) macrothorax Kashcheev, 1999

Shavrin, Alexey V., 2022, The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae), Zootaxa 5213 (5), pp. 451-496 : 478-480

Geodromicus (s.str.) macrothorax Kashcheev, 1999

( Figs. 54–59 , 63 )

Geodromicus macrothorax Kashcheev, 1999: 15

Material examined: KAZAKHSTAN: EAST KAZAKHSTAN: 1 ♂: 30 km S Panfilova, Sarybel. Under stones near the river. 27.07.1984. B.V. Iskakov leg. ( ZIN) ; JAMBYL: 2 ♂♂: Chylik R., Sarybastau Gorge. 15.06.1988. V. Kashcheev leg. [paratypes?] ( ZIN) ; 3 ♂♂: Dzhungar, Kokjota Mt., Topolevka. 27.06.2006. V. Kashcheev leg. ( BMNH, cSh) ; 2 ♂♂: same data, Usek R. 10.07.2005 (cSh) ; 4 ♂♂, 2 ♀♀: same data, Argynkaty R. 31.08.2006 (cSh, ZIN) ; 1 ♂, 1 ♀: same data, Keskenterek R. 25.06.1998 (cSh) ; 7 ♂♂, 3 ♀♀: same data, Tentek R., Kokzhar. 25.08.2006 (cSh, MHNG, ZIN) ; ALMATY: 1 ♂, 1 ♀: Chelkydysu , 43°13′02.4″N 80°17′04.7′E′. 2345 m a.s.l. 21.07.2009. V. Kashcheev leg. (cSh) ; 6 ♂♂, 1 ♀: Ile Karatau , Zhynishke R. 22.06.1988. V. Kashcheev leg. (cSh) ; 1 ♂, 1 ♀: Prokhladnaya R. 2000 m a.s.l. 01.09.2008. V. Kashcheev leg. ( ZIN) ; 7 ♂♂, 10 ♀♀: Ketmen Mts .., Shalkydysu R. 17.07.1988. V. Kashcheev leg. ( BMNH, NHMB, cSh, ZIN) ; 1 ♂, 1 ♀: Baynakol R., near Karatogan. 11.09.1988. V. Kashcheev leg. (cSh) ; KYRGYZSTAN: 2 ♂♂: Issyk-Kul region, Kungey Alatau, Cholpon-Ata District , Grigoryevka , 42°.65′N 77°47′E. 2200 m a.s.l. 09.06.2006. R. Ruta leg. (cRR) .

Redescription. Measurements (n=50): HW: 0.86–1.25; HL: 0.57–0.76; OL: 0.27–0.32; TL: 0.12–0.15; AL (averaged): 3.78; PL: 0.88–1.61; PWmax: 0.78–1.86; PWmin: 0.68–1.21; ESL: 1.46–2.29; EW: 1.58–2.36; MTbL (averaged): 1.37; MTrL (averaged): 0.70 (MTrL 1–4: 0.40; MTrL 5: 0.30); AW: 1.21–2.14; AedL: 0.90–1.00; BL: 4.85–7.90.

Habitus as in Figs. 57–59 . Body yellow-brown to black, usually with paler elytra and sometimes paratergites of abdomen; mouthparts, antennae and legs yellow-brown to brown, usually with darker protibia and antennomeres 2–11 or 3–11. Head without or with fine transverse or isodiametric microsculpture, more distinct in middle and/or infraorbital portions; pronotum with very fine transverse microreticulation or without them; abdominal tergites with very dense transverse microreticulation, finer on abdominal tergites VI–VII.

Head 1.5–1.6 times as broad as long, slightly or moderately strongly convex, with very deep and wide subtriangular or subrectangular anterio-median depression; temples convex, less than twice as long as longitudinal length of eyes; interocellar depression deep, subrectangular and moderately wide, separated from infraorbital portions by moderately deep anteocellar foveae, reaching level of apical third of eyes (some specimens without distinct foveae). Punctation sparse and fine, slightly larger, deeper and denser on infraorbital portions. Antenna reaching apical third or middle of elytra when reclined; antennomere 2 short and narrow, 3 distinctly longer than 2, 4 slightly broader and distinctly shorter than 3, 5–8 slightly longer than 4, 9–10 slightly broader than 8, apical antennomere 1.3–1.5 times as long as 10.

Pronotum very convex, from middle or anterior third gradually or sharply narrowed toward short subparallel laterobasal margins in front of subacute hind angles; middle portion without or with very indistinct and narrow longitudinal depression, with shallow and weakly defined or with moderately deep and transverse mediobasal depression. Punctation distinctly denser, larger and deeper than that on head, with interspaces between punctures in middle about as long as one–four nearest punctures, sparser and finer in medioapical and mediobasal portions (some specimens with small oval impunctated area in front of mediobasal depression).

Elytra slightly broader than long, slightly or widely broadened posteriad, 1.4–1.6 times as long as pronotum. Punctation denser, larger, deeper and sometimes coarser than that on pronotum, finer and denser in parascutellar portion and sometimes larger and denser along suture.

Metatibia about twice as long as metarsi.

Abdomen distinctly narrower than elytra.

Male. Pronotum very wide and convex, sometimes significantly enlarged in some specimens ( Fig. 58 ). Profemuri and protarsomeres 1–4 wide. Apical margin of abdominal tergite VIII truncate or rounded. Apical margin of abdominal sternite VIII concave. Aedeagus slender, narrow, with moderately narrow basal portion, gradually narrowed toward rounded or subacute apex; parameres very wide, distinctly broadened toward apical portions, significantly exceeding apex of median lobe, with four long apical and three to four subapical setae; internal sac weakly sclerotized, with narrow and long flagellum ( Figs. 54, 56 ). Lateral aspect of the aedeagus as in Fig. 55 .

Female. Pronotum moderately small, less convex. Profemuri and protarsomeres 1–4 narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded.

Comparative notes. Geodromicus (s.str.) macrothorax differs from the remaining species of the convexicollis group by the shape of very wide and convex pronotum, sometimes with the presence of an indistinct longitudinal depression, and the different shape of the aedeagus with narrow median lobe and very long and wide parameres, significantly exceeding apex of the median lobe, with four long apical and three to four subapical setae.

Distribution. Geodromicus (s.str.) macrothorax is distributed in the Middle Asia, and known from several localities in Kazakhstan and Kyrgyzstan ( Fig. 63 ).

Bionomics. Specimens were collected at elevations from 2000 to 2340 m a.s.l. and were taken from under stones near streams and rivers. It was collected together with specimens of G. (s.str.) convexicollis in some localities of south-eastern Kazakhstan (see above).

Remarks. Geodromicus macrothorax was originally described based on the holotype amd seven paratypes from “Kungey-Alatau, upper flow of Chylik River, Sarybastau Gorge”. According Kashcheev (1999), all types were deposited in ZIN, the collection of the Institute of Zoology in Astana, and the private collection of the author. During the study of collection of V.A. Kashcheev in ZIN, I have not found the type specimens which have been labeled as “ holotype ” or “ paratype ”. Two studied males from Chylik River (see the Material section) apparently refer to the paratypes of the private collection of V. Kashcheev (1953–2012) deposited to ZIN in 2014. It is quite possible that type specimens can be found in this museum later.

Figures

FIGURE 63. Distribution of Geodromicus macrothorax (squares) and G. lebedevianus (circles) in the Middle Asia.

FIGURES 52–56. Aedeagus of Geodromicus lebedevianus (Figs. 52–53: Kazakhstan, Chichkan Valley) and G. macrothorax (Figs. 54–55: Kazakhstan, Chelkydysu; Fig. 56: Kyrgyzstan, Grigoryevka): 52, 54, 56—parameral view, 53, 55—lateral view. Scale bars: 0.1 mm.

FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm.