Libythea lepita lepita Moore [1858]

Kawahara, Akito Y., 2013, <strong> Systematic revision and review of the extant and fossil snout butterflies (Lepidoptera: Nymphalidae: Libytheinae) </ strong>, Zootaxa 3631 (1), pp. 1-74 : 32-34

publication ID	https://doi.org/10.11646/zootaxa.3631.1.1
publication LSID	lsid:zoobank.org:pub:A355AA9D-3644-4F29-84AA-5D398D2EE6D0
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treatment provided by	Felipe (2021-08-24 23:59:23, last updated by Plazi 2023-11-04 17:58:13)
scientific name	Libythea lepita lepita Moore [1858]
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Treatment

Libythea lepita lepita Moore [1858] View in CoL

( Figs. 7 View FIGURES 2−13 , 19, 31, 44, 59, 125–128)

Libythea lepita Moore [1858] View in CoL : 240.

Libythea celtis Lang 1884: 152 View in CoL [in part, distribution].

Libythea celtis lepita Fruhstorfer 1898: 170 View in CoL .

Libythea (Libythea) celtis var. lepita: Pagenstecher 1901: 4 View in CoL .

Libythea celtis lepita: Bingham 1905: 473 View in CoL , 474.

Libythea celtis chinensis Fruhstorfer 1909b: 209 View in CoL [new synonymy].

Libythea celtis celtoides Fruhstorfer 1909b: 209 View in CoL [new synonymy].

Libythea celtis matsumurae Fruhstorfer 1909b: 209 View in CoL . Shirôzu 1957: 430 [= L. celtis celtoides View in CoL ].

Libythea (Libythea) celtis var. celtoides: Pagenstecher 1911: 5 View in CoL .

Libythea (Libythea) celtis var. chinensis: Pagenstecher 1911: 5 View in CoL .

Libythea (Libythea) celtis var. matsumura: Pagenstecher 1911: 5 .

Libythea lepita celtoides Fruhstorfer 1914: 769 ; pl. 139e.

Libythea lepita matsumurae Fruhstorfer 1914: 769 , pl. 139e.

ibythea lepita View in CoL chinensis Fruhstorfer 1914: 769.

Libythea lepita sophene Fruhstorfer 1914: 769 [new synonymy].

Libythea lepida [sic]: Fruhstorfer 1914: pl. 139e.

Libythea lepita var. celtoides: Matsumura 1919: 24 .

Libythea celtis conjuncta Schawerda 1924: 88 View in CoL . Shields 1984a: 265 [= L. celtis lepita View in CoL ].

Libythea celtis sophene Bang-Haas 1926: 72 .

Libythea lepita View in CoL ab. minomoensis Kasahara 1936: 357, Figs. 2, 4 View FIGURES 2−13 [nomen nudum].

Libythea celtis var. celtoides: Bang-Haas 1937: 99 View in CoL .

Libythea celtis var. chinensis: Bang-Haas 1937: 99 View in CoL .

Libythea celtis var. lepita: Bang-Haas 1937: 99 View in CoL .

Libythea celtis var. matsumurae: Bang-Haas 1937: 99 View in CoL .

Libythea celtis var. sophene: Bang-Haas 1937: 99 .

Libythea lepita matsumae: Kono & Sawamoto 1939: 143 .

Libythea (Libythea) celtis celtoides: Hidaka et al. 1984: 121 View in CoL .

Libythaea [sic] lepita lepita: Gupta & Shukla 1988: 72 .

Libythea celtoides Gorbunov 2001: 148 .

Diagnosis. Libythea lepita lepita is similar to L. l. amamiana and L. l. formosana by sharing similar dorsal wing markings, but differs in that the apical mark in the discal cell is fused with the orange mark between M 3 and CuA 2 by at least 2 mm at the narrowest point. Orange marks on the dorsal surface of the female are generally much larger than marks of the male.

Type Specimens Examined. Libythea celtis celtoides Fruhstorfer 1909 , HOLOTYPE: ♀, Japan ( BMNH). Libythea celtis chinensis Fruhstorfer 1909 , HOLOTYPE: ♂, China: Sze-tsehuan, H. G. Smith ( BMNH) . Libythea celtis formosana Fruhstorfer 1909 , HOLOTYPE: ♂, Formosa [ Taiwan], H. Fruhstorfer ( BMNH) . Libythea celtis matsumurae Fruhstorfer 1909 , HOLOTYPE: ♂, Japan: Yesso, Sapporo, H. Fruhstorfer ( BMNH) . Libythea lepita Moore [1858] , HOLOTYPE: ♂, North India, Col. Buckley ( BMNH) .

Other Specimens Examined. (261♂, 79♀): BHUTAN: Punakha: 1♂, 21-III-1927, F. M. Bailey ( AMNH) . CHINA: Beijing: 1♂, 22-VII-1997, H. Ogura ( TFC) . Fujian: Kuatun , 3500 ft [1067 m], 1♂, VI-1896 ( BMNH) . Hubei [Hebei]: Shennongjia County, Gushui : 1♂, 7-VI-1991 ( TFC) ; 1♂, 26-VI-1991 ( TFC) ; 3♂, VII-1997 ( TFC) . Mupin : 1♂, 12-14000 ft [3658–4267 m], D. C. Graham ( USNM) . Sichuan: Bashan, Daba Shan Mountains , 1600–1900 m: 2♂, 13-VII-1994 ( TFC) . Dujianyan City : 1♂, 1-20-VII-1994 ( TFC) . Jianxi: Xiushui : 1♂, V-1982 ( TFC) . Tibet: C. Pang : 1♂, 19-VI-1913, F. M. Bailey ( AMNH) . Dre : 4♂, 14-VII-1913, F. M. Bailey ( AMNH) ; 7500 ft [2286 m], 1♂, 4-VII-1913 ( BMNH) . Yigrong Valley : 8♂, 7♀, 4-5-VII-1913, F. M. Bailey ( AMNH) ; 1♂, 14-IV-1932, F. M. Bailey ( AMNH) . Rima [Lima]: 1♂, 6-VII-1911, F. M. Bailey ( AMNH) . Po Me, Layoting : 1♂, 12-III-1947 ( AMNH) . Rong Valley : 1♀, 6-VII-1913, F. M. Bailey ( AMNH) . Zayul, Chikung : 1♂, 7-III-1911 ( AMNH) . Shanxi: Xian : 1♂, 14-III-1994 ( TFC) . Yunnan: 5♂ ( USNM) ; Loutsechiang : 3♂ ( AMNH) ; 2♂ ( USNM) . INDIA: 3♂ ( AMNH) ; 1♂, 1♀ ( USNM) . ‘ Himalayas’ : 1♀, Jacob Doll ( AMNH) . ‘ Tassienlou’ : 1♂, B. Neumögen ( USNM) . Arunachal Pradesh: Abor, Chindro V., 5500 ft [1676 m], 1♂, 4-VI-1913 ( BMNH) ; Mishmi, Imulin : 1♀, 5-IV-1913 ( AMNH) . Sikkim: 5♂, 2♀, 25-VII-1927, F. M. Bailey ( AMNH) . Himachal Pradesh: Dalhousie , 1♂, 16- V-1891 ( BMNH) ; 1♂, 13-X-1891 ( BMNH) . Jutogh , 1♂, 2♀ 19-IV-1878 ( BMNH) . Kangra, Hocking : 1♂, B. Neumögen ( USNM) . Kasauli , 2♂, 1♀, 23-25-VIII-1903 ( BMNH) . Kullu : 1♀, 10-VIII-1992 ( TFC) ; 4♂ ( BMNH) . Shimla , 2♂, 1♀ ( BMNH) ; 2000 ft [610 m], 1♀, [4]-[X]-1931, F. M. Bailey ( BMNH) . Near Shimla , 8000 ft [2438 m], 1♂, VII-1922, C. H. Stockley ( BMNH) . Shimla Hills, Mahasu : 1♀, 19-V-1931, F. M. Bailey ( AMNH) . Kumaon : 3♂, 1♀, Doherty ( CMNH) . Kumaon, Sattar : 1♂, 22-IV-1992 ( TFC) . Madhya Pradesh: 1♀, 10-V-1931, F. M. Bailey ( AMNH) . Uttaranchal: Mussoorie , 5400 ft [1646 m]: 1♀, 19-V-1962, J. P. Donahue ( MSUC) ; 5500 ft [1676 m], 1♀, 6-VI-1912, H. D. Peile ( BMNH) ; 1♂, 5-V-1913, H. D. Peile ( BMNH) ; 1♀, 5-V-1916, H. D. Peile ( BMNH) ; 1♂, 28-V-1916, H. D. Peile ( BMNH) ; 1♀, 20-VIII-1916, H. D. Peile ( BMNH) ; 1♂, 27-VIII-1916, H. D. Peile ( BMNH) . Mussoorie, Dehra Dun district : 2♂, 17-V-1961, E. M. Shull ( AMNH) . West Bengal: Teesta Valley : 2♂, III-IV, 1927, F. M. Bailey ( AMNH) . JAPAN: Aichi: 1♂, Asuke, 11-VIII-1989, A. Y. Kawahara and O. Kawahara ( AYK) . Fukuoka: Mt. Inunaki : 2♂, 10-VI-1950, T. Shirozu ( MGCL) ; Shikamajima : 1♂, 28-VI-1947, R. J. J. ( MGCL) . Gifu: 2♀, 4-8-VI-1919 ( USNM) ; 2♂, 1♀, 30-V-1924 ( USNM) ; 1♂, 11-VI-1924 ( USNM) . Gunma: Mount Asama : 1♂, 11-IX-1923, S. Satake ( CMNH) . Hyogo: Kobe : 1♂, 14-VI-1900, E. Scarlett ( BMNH) ; 2♂, 18- IV-1934 ( CAS) ; 1♂, 18-IV-1934, S. Yamada ( USNM) ; 1♀, 22-VI-1934, S. Yamada ( USNM) . Mount Futatabi : 1♂, 11-VI-1940, H. Masuda ( CAS) . Rokko : 3♂, 1♀, 10-IV-1925 ( AMNH) . Kumamoto: Haki , 2♂, 1-VII-1893, A. E. Wileman ( BMNH) ; Makinohara , 1♂, 18-VI-1893, A. E. Wileman ( BMNH) ; Mamibara , 1♂, 23-VII-1893, A. E. Wileman ( BMNH) . Kyoto: 1♂, 17-X-1918, I. Sugitani ( AMNH) ; 1♂, 10-VII-1928 ( USNM) ; 1♂, 6-VI-1939, T. Shirozu ( MGCL) ; 1♂, 1-VI-1950, Y. Okada ( CAS) ; 3♂, 6-7-VII-1950 ( AMNH) ; 2♂, VI-1954 ( USNM) ; 1♂, VI- 1959 ( CAS) ; 2♂, 6-VIII-1980, G. A. Gorelich ( CMNH) . Arashiyama , 1♂, H. Fruhstorfer ( BMNH) . Kibune : 7♂, 10-VI-1951, B. Weber ( MGCL) ; 2♀, 19-IV-1953, B. Weber ( MGCL) . Kurama : 2♂, 2♀, 14-X-1928, I. Sugitani ( CAS) ; 1♂, 21-VI-1929, I. Sugitani ( CAS) . Iwate: Iwaizumi-cho : 1♂, 4-VIII-1998, M . Chiba ( TFC) . Kanagawa: Yokohama : 1♂, 1♀, Pryer ( BMNH) ; 1♂, III-1901, J. Hashimoto ( AMNH) ; 1♂, 3-V-1902 ( CUIC) . Nagano: Kamikochi : 3♂, 2-VIII-1917 ( USNM) . Karuizawa : 1200 m: 1♂, 2-VIII-1975, I. Hashimoto ( CAS) ; 1206 m: 1♀, 26-VIII-1974, P. J. and M. C. Shanahan ( CAS) . Oiwake : 1♂, 2♀, 1885, H. Pryer ( BMNH) ; 1885, H. Pryer ( BMNH) ; 1♂, 8-VIII-1956 ( LACM) . Yatsugatake : 1♂, 12-VII-1953 ( CVC) . Nagasaki: Tsushima, Toyotama : 2♂, 12-VII-1998 ( TFC) . Okayama: 1♂, N. Hirata ( USNM) . Kuboino : 1♂, 17-VII-1995, S. Yamanaka ( TFC) . Osaka: Minomo : 2♂, 1934 ( CAS) . Myoken : 1♂, 1♀, 20-VI-1937, S. Inoue ( CAS) . Nagasaki: Tsushima , 1♂, Sept-Oct., H. Fruhstorfer ( BMNH) . Nara: Amagase , 1♂, 11-VII-1894 ( BMNH) ; Yoshino , 984 ft [300 m], 1♂, 1♀, 9-VI-1901, A. E. Wileman ( BMNH) . Miyazaki: Oteno , 2♂, 6-VI-1893, A. E. Wileman ( BMNH) . Saitama: Agano : 2♂, 26-VI- 1932 ( USNM) . Iruma : 1♂, 29-VI-1981, M. Jinkubo ( AMNH) . Shiga: Yokkaichicho : 1♂, 18-VI-1953 ( CVC) . Shimane: Okinoshima, Dogo Island, Ushiki : 1♂, 4-VII-1982, T. Fujioka ( TFC) ; 1♀, Okinoshima, Tsuma-mura : 9- 10-X-1985, T. Fujioka ( TFC) . Shizuoka: 1♂ ( CAS) ; 1♂, 18-VII-1984 ( KKC) . Fujinomiya : 2♂, 12-VI-1991, K. Kobayashi ( KKC) ; 1♂, 12-VI-1996, K. Kobayashi ( KKC) ; 2♂, 4-14-XI-1998, A. Y. Kawahara ( AYK) ; Agoyama , 150 m: 1♀, 30-V-1999, A. Y. Kawahara ( AYK) ; 1♂, Numakubo, 90 m, 26-V-1999, A. Y. Kawahara ( AYK) . Mount Fuji : 1♂, 16-VII-1969, K. Kobayashi ( KKC) ; 1♀, 14-VIII-1975, K. Kobayashi ( KKC) ; 1♀, 21-X-1977, K. Kobayashi ( KKC) ; 1♀, 1-VI-1983, K. Kobayashi ( KKC) ; 1♂, 22-VI-1983, K. Kobayashi ( KKC) ; 1♂, 4-VIII- 1983, K. Kobayashi ( KKC) ; 1♂, 9-VIII-1995, A. Y. Kawahara ( AYK) . Mount Setsugata : 1♂, 18-VII-1984, K. Kobayashi ( KKC) . Sakamoto : 1♂, 6-VI-1999, A. Y. Kawahara ( AYK) . Shibakawa, Inako : 1♂, 3♀, 24-X-1983, K. Kobayashi ( KKC) . Tochigi: Nikko , 2♀, 1880 ( BMNH) . Tokushima: Bizan : 1♂, 2-VIII-1987, A. Y. Kawahara and O. Kawahara ( AYK) . Tokyo: 1♀, 15-VI-1931, I. Gressitt ( CAS) ; 2♂, 1962-1963, G. Evans ( USNM) . Mount Takao : 2♂, 20-VI-1963 ( CAS) . Tottori: 1♂, Wakazakura-cho, Inaba forest trail: 10-VII-1995, S. Yamanaka ( TFC) . Yamanashi: Hinoharu : 1♀, 20-VI-1988, A. Y. Kawahara ( AYK) ; 1♀, 8-VII-1990, A. Y. Kawahara ( AYK) ; 2♂, 5♀, 21-VI-1991, A. Y. Kawahara ( AYK) . Kamiashigawa, Ohishi River , 880 m: 1♂, 16-VI-1999, K. Kobayashi ( AYK) . Kamikuishiki, Hongo, Ashigawa Valley , 550 m: 1♂, 16-VI-1999, A. Y. Kawahara ( AYK) ; 1♂, Lake Motosu , 14- VIII-1975, K. Kobayashi ( KKC) . Lake Shoji : 1♀, 29-VII-1981, M. Kobayashi ( KKC) . Mitama : 1♂, 1♀, 17-VI- 1983, K. Kobayashi ( KKC) ; 4♂, 25-VI-1997, K. Kobayashi ( KKC) . Mitsu-toge : 1♂, 1-VIII-1984, K. Kobayashi ( KKC) ; Naka-Ashigawa : 1♂, 17-VI-1983, K. Kobayashi ( KKC) . Nanbucho : 1♀, 14-IV-1999, K. Kobayashi ( KKC) ; Sanogawa : 5♂, 3-IV-1996, A. Y. Kawahara ( AYK) ; 1♀, 13-IV-1999, A. Y. Kawahara ( AYK) . Nirasaki : 1♂, 18-VI-1986, K. Kobayashi ( KKC) . Shoji , 1♀ ( BMNH) . Tenshigatake : 2♂, 1♀, 13-VIII-1995, A. Y. Kawahara ( AYK) . Uenohara : 2♂, 1♀, 18-VI-1979, K. Ishizuka ( AMNH) . Yamanakako : 1♂, 19-VIII-1993, K. Kobayashi ( KKC) . Uncertain Prefecture : 1♀ ( BMNH) ; 2♀, Pryer ( BMNH) ; 1♂, IX-1877 ( BMNH) ; 1♀, 4-VI-1901 ( BMNH) ; 1♂, 8-VIII-1901 ( BMNH) ; 1♂, H. M. Holbrook ( CAS) ; 1♂, 19-VII-1911, E. R. Tinkham ( CAS) ; 5♂, 1♀ ( CMNH) ; 3♂, 1♀ ( USNM) ; 1♂, 2-VI ( CUIC) ; 1♀, V-1922 ( CUIC) ; 1♀, 9-VI-1922 ( CUIC) ; 1♂, 12-VI-1960, K. Kobayashi ( KKC) . ‘ Minoo’ : 1♂, VI-1934, K. Etoh ( USNM) . ‘ Hikawa’ : 1♂, 7-VIII-1937 ( USNM) . [ Locality illegible]: 1♂, 28-VI-1978, K. Kobayashi ( KKC) ; 1♂, 17-VI-1976, K. Kobayashi ( KKC) . KOREA: 1♂, 6-VI- 1990, S. Igarashi ( TFC) . National Forest 7 mi [11.27 km] east of Uijongbu : 1♂, 1-IV-1954, T. A. Derse ( AMNH) . NEPAL: Katmandu : 5♂, F. M. Bailey ( AMNH) ; 22♂, 2♀, 27-IV – 11-V-1935, F. M. Bailey ( AMNH) ; 1♂, 11-V- 1936, F. M. Bailey ( AMNH) ; 5♂, 31-V-1937, F. M. Bailey ( AMNH) ; 1♂, 30-V-1938, F. M. Bailey ( AMNH) . Katmandu Valley, Godavari : 1♀, 20-VI-1936, F. M. Bailey ( AMNH) ; 10♂, 2♀, 13-21-V-1938, F. M. Bailey ( AMNH) ; 1♂, 30-V-1977, Smith ( CMNH) ; 1♂, 11-II-1978, Smith ( CMNH) ; Godavari School : 1♂, 1960, E. L. Watrin ( CAS) . Manari : 1♂ ( TFC) . Nagarkot : 1♂, 19-IV-1935, F. M. Bailey ( AMNH) .

Distribution. Central Asia to East Asia: Bangladesh, Bhutan, China, India, Japan, Korea and Nepal.

References

Bang-Haas, O. (1926) Novitates Macrolepidopterologicae, 1. Staudinger Bang-Haas, Dresden-Blasewitz. xx, 238 pp.

Bang-Haas, O. (1937) Catalogus lepidopterorum regionis palaearcticae, 1. Staudinger preis liste palaearktischer lepidopteren, 61. Blasewitz, Dresden. 141 pp.

Bingham, C. T. (1905) The fauna of British India, including Ceylon and Burma, Butterflies, 1. Taylor & Francis, London. 511 pp., 10 pls.

Fruhstorfer, H. (1898) Eine neue Libythea aus Asien. Berliner Entomologische Zeitschrift, 43, 169 - 170. http: // dx. doi. org / 10.1002 / mmnd. 18970420106

Fruhstorfer, H. (1909 b) Drei neue Libythea - Rassen. Entomologische Zeitschrift, 22 (49), 209.

Fruhstorfer, H. (1914) Libythea. In Seitz, A. [Ed.], Die Gross schmetterlinge der Erde, 9. Alfred Kernen, Stuttgart. pp. 767 - 771.

Gorbunov, P. Y. (2001) The butterflies of Russia: classification, genitalia, keys for identification (Lepidoptera: Hesperioidea and Papilionoidea). Russian Academy of Sciences Institute of Plant & Animal Ecology, Thesis. Ekaterinburg. 320 pp.

Gupta, I. J. & Shukla, J. P. N. (1988) Studies on the butterflies of Arunachal Pradesh and adjoining areas, India (Lepidoptera: Acraeidae, Satyridae, Nymphalidae, Riodinidae and Lycaenidae). Records of the Zoological Survey of India Occasional Paper, 109, 1 - 115 + 23 pls.

Hidaka, T., Fujii, H., Unno, K. & Imamori, M. (1984) F i rudo zukan - kar a shashin ni yoru: Ch o. Tokai University Press, Tokyo. 221 pp.

Kasahara, T. (1936) [On an aberration and variation of the marks of Libythea lepita.] Zephyrus, 6, 356 - 359.

Kono, H. & Sawamoto, T. (1939) On the foods of the larvae of the butterflies in Hokkaido. Kontyu, 12 (4), 140 - 143.

Lang, H. C. (1884) Butterflies of Europe, 1. Reeve, L., London. vi, 396 pp.

Matsumura, S. (1919) Thousand insects of Japan Additamenta, 3 [4]. pp. 475 - 742, 1 - 34; pls. 26 - 53. Tokyo.

Moore, F. (1858) A catalog of the Lepidopterous Insects in the Hon. East-India Company [by Horsfield, T. H. & Moore, F.]. v, 14 pp. (introduction), 278 pp. (text), 11 pp. (index), 18 pls. W. M. H. Allen & Co., London.

Pagenstecher, A. (1901) Libytheidae. Das Tierreich, 14, 1 - 18.

Pagenstecher, A. (1911) Libytheidae. In Aurivillius, C. & Wagner, H. [Eds.], Lepidopterorum catalogus, 3. Junk, W., Berlin. 12 pp.

Schawerda, K. (1924) Faunula dolomitana. Mitteilungen der Munchner Entomologischen Gesellschaft 8, 73 - 142 + 1 plate.

Shields, O. (1984 a) A revised, annotated checklist of world Libytheidae. Journal of Research on the Lepidoptera, 22, 264 - 266.

Figures

FIGURES 2−13. Lateral view of the male eighth abdominal tergum of Libythea and Libytheana with scales and hairs removed. 2. Libythea celtis; 3. Libythea collenettei; 4. Libythea geoffroyi; 5. Libythea labdaca; 6. Libythea laius; 7. Libythea lepita; 8. Libythea myrrha; 9. Libythea narina; 10. Libytheana carinenta; 11. Libytheana fulvescens; 12. Libytheana motya; 13. Libytheana terena.

Abbreviations

AMNH	American Museum of Natural History
USNM	Smithsonian Institution, National Museum of Natural History
CMNH	The Cleveland Museum of Natural History
CAS	California Academy of Sciences
CUIC	Cornell University Insect Collection
LACM	Natural History Museum of Los Angeles County

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Lepidoptera
Family	Nymphalidae
Genus	Libythea

1926

Libythea celtis conjuncta

Libythea celtis

Lang, H. C. 1884: 152

1884

Libythea lepita

Moore, F. 1858: 240

1858

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