Hepatus Latreille, 1802

Lima, Daniel, Silva, Rafael Costa da, Aguilera, Orangel, Pinheiro, Allysson Pontes & Santana, William, 2023, Brazilian Miocene crabs I. Taxonomic review of Cyclocancer tuberculatus Beurlen, 1958 and Hepatella amazonica Beurlen, 1958 (Pancrustacea, Decapoda, Brachyura), Papéis Avulsos de Zoologia 63, pp. 1-9 : 3

Genus Hepatus Latreille , in Bosc, 1801

Hepatus Latreille , in Bosc, 1801: 55, 56 [type species: Calappa angustata Fabricius, 1798 , by monotypy, junior synonym of Hepatus pudibundus (Herbst, 1785) ].

Hepatus Latreille, 1802: 22 [type species: Calappa angustata Fabricius, 1798 , by monotypy, junior synonym of Hepatus pudibundus (Herbst, 1785) ].

Hepatulus Fowler, 1912: 590 [type species: Hepatus fasciatus Latreille, 1803 ; unnecessary replacement name for Hepatus Latreille , in Bosc, 1801].

Hepatoides Balss, 1957: 1612 [unnecessary replacement name for Hepatus Latreille , in Bosc, 1801].

Cyclocancer Beurlen, 1958: 15 [type species Cyclocancer tuberculatus Beurlen, 1958 , by monotypy].

Included species: Hepatus beurleni †† nomen nov.; H.biformis †† Collins & Todd in Todd & Collins, 2005; H. bottomsi †† Blow,2003; H.chiliensis † H. Milne-Edwards,1837 ; H. epheliticus ( Linnaeus, 1763) ; H. gronovii Holthuis, 1959 ; H. guraboensis †† Collins in Collins et al., 2009; H. kossmanni Neumann, 1878 ; H. lineatus † Rathbun, 1898 ; H. lineatinus †† Collins & Todd in Todd & Collins, 2005; H. nodosus †† Collins & Morris, 1976; H. pauli †† Collins, Garvie & Mellish, 2014; H. praecox †† Collins, Donovan & Dixon, 1996; H. pudibundus (Herbst, 1785) ; H. scaber Holthuis, 1959 ; H. spinimarginatus †† Feldmann, Schweitzer & Encinas, 2005.

Remarks: The authorship of Hepatus has been generally attributed to Latreille (1802: 22) (e.g., Desmarest & Bosc, 1830; Holthuis, 1959; Glaessner, 1969; Ng et al., 2008; De Grave et al., 2009; Schweitzer et al., 2010). Hepatus was recently attributed to Bosc(1801:55,56)(seeWoRMS,2023). Bosc (1801: 29, 48) makes it clear that he used some results of Latreille′s manuscript – which was in preparation. More importantly, Bosc (1801: 29, 48) attributed authorship to Latreille (see Opinion 1133: 97 by C. Dupuis, Case 1879, BZN 1979: 97 for Talitrus Bosc, 1802 ). According to the ICZN (1999: Art. 50.1.1) "if it is clear from the contents that some person other than an author of the work is alone responsible both for the name or act and for satisfying the criteria of availability other than actual publication, then that other person is the author of the name or act″.Thus,the authorship for Hepatus should therefore be Latreille, in Bosc (1801: 55, 56) (type species Calappa angustata Fabricius, 1798 , by monotypy). Desmarest & Bosc (1830: 42, 209), in the second edition of Bosc (1801, 1802) attributed the authorship of Hepatus to Latreille. Here we follow Dupuis (Opinion 1133: 97) (for more details see ICZN Case 1879; Bousfield & Holthuis, 1969; Dupuis, 1975; Holthuis, 1975).