Curtonotum parkeri, Kirk-Spriggs, 2011

Kirk-Spriggs, Ashley H., 2011, A revision of Afrotropical Quasimodo flies (Diptera: Schizophora; Curtonotidae). Part III - the Malagasy species of Curtonotum Macquart, with descriptions of six new species, African Invertebrates 52 (2), pp. 391-391 : 422-425

publication ID	https://doi.org/ 10.5733/afin.052.0212
persistent identifier	https://treatment.plazi.org/id/03FDA115-FFCD-B371-EB99-9920FD28FBA1
treatment provided by	Felipe
scientific name	Curtonotum parkeri
status	sp. nov.

Treatment

Curtonotum parkeri View in CoL View at ENA sp. n.

Figs 5 View Figs 1–8 , 18 View Figs 14–26 , 31 View Figs 27–39 , 65, 68, 71 View Figs 64–72 , 82 View Figs 79–91 , 96 View Figs 92–96

Etymology: The species is named in honour of Frank Parker, in recognition of his contribution to our knowledge of Madagascan Hymenoptera.

Differential diagnosis: This species is closely related to C. coronaeformis sp. n.; the shape, maculae and setation of abdominal sternites 4 and 5 are virtually identical, the apical region of the basiphallus is markedly expanded in both species, with very similar left and right raised keels, and both share the raised and spinose right sclerotised area of the basiphallus. In C. parkeri sp. n., however, the basiphallus is more markedly narrower medially and expanded in the apical third, and the raised and spinose right sclerotised area of the distiphallus is less conspicuously developed, with three short, dark-tipped spinules. The two species occur sympatrically.

Description:

Male (primarily based on ex spirit-preserved HT).

As redescribed for C. balachowskyi , differing in the following respects: Measurements: Overall length 3.65–4.4 mm (n = 4, PT); length of head and thorax combined 2.8 mm; length of thorax and scutellum combined 2.3 mm; wing length 4.1 mm (HT).

Head ( Figs 5 View Figs 1–8 , 18 View Figs 14–26 ). As described for C. gladiiformis sp. n., except: eye height/length ratio: 10:6 (HT); frons ( Fig. 18 View Figs 14–26 ), frons length/width ratio: 5:6 (HT), arista with 7 or 8 long dorsal branches and 3 or 4 ventral branches in addition to terminal fork; gena narrow, eye height/genal height ratio: 12:1 (HT), silver pruinose, slightly darker beyond basal angle; palpus pale brown.

Thorax ( Fig. 5 View Figs 1–8 ). Mesonotum as described for C. boeny ; presutural seta moderately strong, shorter than posterior notopleural seta; supra-alar seta twice as long as posterior dorsocentral seta; postalar setae longer and stronger than acrostichal setae; postpronotum with 6 finer blackbrown setulae; anepisternum silvergrey pruinose, with silveryellow pruinose areas, surface with 20 fine setulae, some larger and arranged in 2 groups of 3 and 4; anepimeron, laterotergite and meron silver-grey to silver-yellow pruinose; katepisternum silver-grey to silver-yellow pruinose, with darker macula in anterior half, dorsal katepisternal seta fine, ca 0.3 length of ventral katepisternal seta, with 16 short, fine setulae at base and along posterior margin.

Scutellum. As described for C. gladiiformis sp. n.

Legs. Fore coxa with 14 brown setulae on anterior surface; mid coxa with 7 brown setulae; fore tibia with ctenidium of 13 spinules.

Wing (as in Fig. 31 View Figs 27–39 ). Veins chestnut-brown, membrane very faintly infuscate brown throughout, very slightly darker in region of dm–cu crossvein; dm–cu crossvein with even arc; haltere dirty white.

Abdomen. Tergite 1 simple, devoid of maculae; tergite 2 with oblique, subovoid brown-black pruinose dorsolateral macula on either side only; tergites 3–5 with large, broad, V-shaped, concolourous median fascia and large, concolourous T-shaped dorsolateral macula, these close or adjoining, but not fully merging with median fascia; lateral margin of tergites 2–5 with subelliptical, concolourous macula in basal half; sternites 4 and 5 as described for C. coronaeformis sp. n.; sternite 6 ( Fig. 82 View Figs 79–91 ) markedly expanded laterally (may appear narrower than Fig. 82 View Figs 79–91 in undissected specimens), narrowed basally with shallow, broad, subtriangular excision apically, with dark brown maculae medially, clothed in long, dense, irregular brown setulae, those at apical margin longer and more prominent.

Terminalia ( Figs 65, 68, 71 View Figs 64–72 ). Hypandrium ( Fig. 65 View Figs 64–72 , hy) long, with broad-based rounded-truncate dorsobasal lobe, posterior bridge dorsally and ventrally produced (subtriangular to slightly angulate in profile); hypandrial arm constricted apically (viewed laterally), with 2 setulae proximal to postgonite, the more lateral strong, ventrally directed, the medial much smaller and weaker (obscured by epandrium on Fig. 65 View Figs 64–72 ), sclerotised area of medial lobes (viewed dorsally), with margins evenly rounded, convex medially, closely abutting, overlapping; postgonite long, thin and straight, with slight undulating anterior margin; epandrium ( Fig. 65 View Figs 64–72 , ep) slightly broader dorsally than ventrally (viewed laterally), evenly rounded on dorsal margin, posterior margin slightly angled, ventral margin with extensive row of long regular to irregular, apically-directed setae; cercus ( Fig. 65 View Figs 64–72 , ce) not prominent, longest setae longer than medium setae on dorsal margin of epandrium; surstylus ( Fig. 65 View Figs 64–72 , ss) long, widest basally, slightly curved in apical ⅔; phallus (as in Figs 68 View Figs 64–72 , ph, bp, dp, 71, bp, dp) C-shaped, strongly sclerotised, brown; phallapodeme ( Fig. 68 View Figs 64–72 , ph) fused to basiphallus, subtriangular (viewed laterally), with basal margin developed into two flat, narrow, subtriangular projections in basal 0.4, bifurcated at point of connection with hypandrium; ejaculatory apodeme ( Fig. 68 View Figs 64–72 , ea) free, duct inserted at junction of phallapodeme and basiphallus; basiphallus ( Fig. 68 View Figs 64–72 , bp) broad in basal third, with slight internal bulge at first bend, very markedly narrowed in apical ⅔ (viewed dorsally); apical section ( Fig. 71 View Figs 64–72 , bp) extremely broad, right lateral margin evenly rounded with semicircular raised margin, left lateral margin with distinct raised keel; distiphallus ( Figs 68, 71 View Figs 64–72 , dp) broad basally, sclerotised area extensive, divided medially, with thin ventrally-directed spine in membranous window, right lateral section with three short, dark-tipped spines (viewed dorsolaterally), apical section narrow, bent and spindle-like.

Variation: No significant variation is noted.

Holotype: ♂ “ MADAGASCAR: Tuléar Province / Zombitse National Park / near ANGAP office, 840 m / 22°53.19’S, 44°41.53’E / 28.ii–6.iii.2002 / California Acad of Sciences / R. Harin’Hala, Malaise trap / deciduous spiny forest / MA0213A18 // HOLOTYPE ♂ / Curtonotum / parkeri sp. n. / A.H. Kirk-Spriggs 2010 [red card]” ( CAS). In good condition, some setation of head missing; card-pointed; right wing detached and glued to card; dissected, abdomen and terminalia in micro-vial pinned beneath specimen. GoogleMaps

Paratypes (all labelled: “ PARATYPE ♂ / Curtonotum / parkeri sp. n. / A.H. Kirk-Spriggs 2010 [blue card]”): MADAGASCAR: 3♂ same data as holotype (CAS); same except: 3♂ “ 29.xii.2001 – 5.i.2002, MA0213A10”; 1♂ “ 26.i–5.ii.2002, MA0213A14”; 4♂ “ 14–21.ii.2002, MA0213A16”; 3♂ “ 27. iii–3.iv.2002, MA0213A22”; 1♂ “ 1–9.v.2002, MA0213A27”; 1♂ “ MADAGASCAR: Tuléar / Prov., Zombitse National / Park, near national road / 22°50.43’S, 44°43.87’E / 20–27.iii. 2002, 825 m / R. Harin’Hala, Malaise / trap, deciduous spiny forest / MA0213B21”; 2♂ “ MADAGASCAR: Province / Fianarantsoa, radio tower / 22 km SW of Ilakaka, near / Fianarantsoa /Tuléar border / 22°46.75’S, 45°1.50’E / 27.ii–6.iii.2002, 1100 m / M.E. Irwin & R. Harin’Hala / Malaise in Uapacca forest / MA-02-12-10”; 1♂ “ MADAGASCAR: Tuléar / Province, Andohahela Nat’l / Park, Tsimelahy, Parcelle II / 24°56.21’S, 46°37.60’E / 6–16.i. 2003, 180 m / M.E. Irwin, F. Parker & / R. Harin’Hala, Malaise trap / in transitional forest / MA022012”; 1♂ “ MADAGASCAR: Province / Fianarantsoa, near Isalo / National Park, in dry wash / east of Interpretive Center / 22°37.60’S, 45°21.49’E / 10–17.ii. 2002, 885 m / M.E. Irwin & R. Harin’Hala / Malaise trap in open area / MA0211B15”; same except: 1♂ “ 22–30.vi.2002, MA0211B33”; 1♂ “ 4–13.v.2002, MA0211B27”; 1♂ “ 6–14 xii.2002, MA0211B49”; 1♂ “ 2–12.ii.2003, MA02 11B55”; 1♂ “ MADAGASCAR: Tuléar / Province, Beza Mahafaly / Reserve, Parcelle I near / research station / 23°41.19’S, 44°35.46’E / 7–18.vii. 2002, 165 m / R. Harin’Hala, Malaise / trap in dry deciduous forest / MA-02-14A-29” (all CAS).

Other material examined (all labelled “ Curtonotum parkeri sp. n. det. A.H. Kirk-Spriggs 2010”): MADAGASCAR: Fianarantsoa Prov.: 1♂ near Isalo National Park, in dry wash east of Interpretive Center , 22°37.60'S: 45°21.49'E, 9–17 xi. 2001, 885 m, M.E. Irwin & R. Harin’Hala, Malaise trap in open area (MA0211B02) [in spirit], BMSA(DNA)#0017 GoogleMaps ; 1♂ same except: 14–22.vi.2003 (MA0211B67) [in spirit], BMSA(DNA)#0061; 1♂ radio tower, 22 km SW of Ilakaka near Fianarantsoa / Tuléar border, 22°46.75'S: 45°1.50'E, 27.ii–6.iii.2002, 1100 m, M.E. Irwin & R. Harin’Hala, Malaise in Uapacca forest (MA021210) [in spirit], BMSA(DNA)#0041 GoogleMaps ; 1♂ same except: 16–27.ii.2002 (MA021209) [in spirit], BMSA(DNA)#0060 (all BMSA) .

Distribution ( Fig. 96 View Figs 92–96 ): Occurring in Tapia Forest and Wooded Grassland-Bushland vegetation types in the Central Highlands, Evergreen Rainforest and Dry Deciduous Forest biomes. In the South and South East biogeographical zones and Subarid bioclimatic zone ( Figs 105–107 View Figs 105–107 ; Tables 1–3; Appendix II).