Milania agrion ( Séguy, 1948 ) Šifner, 2010

Šifner, František, 2010, Two new genera and species of the family Scathophagidae (Diptera) from the Palaearctic and Oriental Regions with additional faunistic records, Acta Entomologica Musei Nationalis Pragae 50 (2), pp. 609-618 : 611-612

Milania agrion ( Séguy, 1948) comb. nov.

( Figs. 1–5 )

Norellisoma agrion Séguy, 1948: 169 .

Norellisoma agrion: FUKUHARA & KARAHASHI (1966) : 217; HIRONAGA & SUWA (2005): 207; ŠIFNER (2008a): 127 .

Type material examined. HOLOTYPE: J, ‘ Japan, Chuzenji , 11.7.1917, Edme Gallois’ [yellow label] / ‘ Type’ [red label] / ‘ Museum Paris’ / ‘ Norellisoma agrion , J, Type, E. Séguy vid. 48’ ( MNHN) . PARATYPES: J, ‘ Japan, Chuzenji , 12.9.1917 ’ / ‘ Museum Paris, coll. J. Hervé – Bazin 1923’ / ‘ Cotype Norellisoma agrion J, Cotyp., E. Séguy vid.48 ’; ♀, ‘ Japan, Chuzenji , 29.7.1917, Edme Gallois’ / ‘ Cotype Museum Paris, coll. J. Hervé – Bazin 1923’ / ‘ Norellisoma agrion ♀, Cotyp., E. Séguy vid. 48 ’; ♀, ‘ Japan, Chuzenji , 12.9.1917, Edme Gallois’ / ‘ Museum Paris, coll. J. Hervé – Bazin 1923’ (all MNHN) .

Additional material examined. JAPAN: HONSHU: 1 J, ‘ Japan, Nagano, Ynago-yu, 1500–1600m, Mt. Yatsugatake , 28.vi.1989, M.Suwa leg.’ / ‘ Norellisoma agrion J, Det.M. Suwa 1994 ’; 1J 1♀, ‘ Japan, Iwate, Mt.Hayachine , 1000– 1400m, 12.–13.vii.[19]80, Maasaki Suwa’ / ‘ Norellisoma agrion Séguy,1948 , J/ ♀, Det. T. Hironaga 1999’ ( FSPC) .

Redescription. Body length 14–16 mm.

HEAD. Two orbital bristles, 2–3 frontal bristles, frontal vitta, parafacials, frons, genae and antennae completely yellow; arista plumose; vibrissal callus plain with one vibrissa; palpi yellow, narrow with one apical bristle; inner vertical bristles long, outer vertical bristles short.

THORAX. Setation consisting of 4–5 dorsocentral bristles (prescutellar pair very long), one short humeral bristle, two notopleural bristles, one prealar bristle, one supraalar and two postalar bristles, two pairs of scutellar bristles of equal size, one katepisternal bristle; all bristles conspicuously fine and acrostichal bristles very sparse. Wings with brown veins, legs completely yellow with long yellow hairs.

ABDOMEN. Male. Sternites 3 and 4 rectangular, with long yellow hairs; lobes of sternite 5 straight, narrowed apically, completely covered with long yellow hairs and with short thorns medially ( Fig. 1 ); cerci fused medially, in posterior view arched, with black bristles, surstyli apically arched forward ( Figs. 2 and 3 ). Epiphallus straight and dilated apically, pregonite basally very narrow, widely dilated apically with two short subapical bristles, postgonite very long and dilated apically ( Fig. 4 ). Female. Sternites 5 and 6 long, with one pair of black bristles caudally; sternite 7 slightly dilated caudally, laterally sclerotized and membranous in middle; sternites 8 long, oval, narrowed apically, with short and fine apical setae ( Fig. 5 ).

Distribution. Japan ( SÉGUY 1948, FUKUHARA & KARAHASHI 1966, HIRONAGA & SUWA 2005, ŠIFNER 2008a).

Figures

Figs. 1–10. 1–5 – Milania agrion (Séguy, 1948) comb. nov.; 6–10 – Milania longiabdomina (Sun, 1992) comb. nov. 1–4, 6–9 – J: 1, 6 – abdominal sternites 3 to 5; 2–3, 7–8 – cerci and surstyli (caudal and lateral views); 4, 9 – penis apparatus with pregonites and epiphallus; 5, 10 – ♀: abdominal sternites 5 to 8, with two spermathecae. Scale bars: 0.5 mm (Figs. 1–3, 6–8, 10); 0.1 mm (Figs. 4, 9).