Mannerheimia cachemirica Coiffait, 1982

Shavrin, Alexey V., 2024, On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), Zootaxa 5474 (5), pp. 584-592 : 585-587

Mannerheimia cachemirica Coiffait, 1982

( Figs 1–2 , 6–8 )

Mannerheimia cachemirica Coiffait, 1982: 146 ; Shavrin 2021: 307

Mannerheimia vicina Coiffait, 1982: 146 syn. n.

Mannerheimia vicina : Shavrin 2021: 307

Type material examined. Holotype of Mannerheimia cachemirica Coiffait, 1982 ♂ ( Fig. 1 ; dissected): ‘Kashmir, Aru | lg.H.Franz, Okt. 1977 ’ <printed>, ‘TYPE’ <red, printed>, ‘ Mannerheimia | cachemirica | H. COIFFAIT det. 19[printed]76’ <handwritten in black> ( NMW). Paratype ♂ (dissected; aedeagus missing): first labels as that in the holotype, with red printed label: ‘PARATYPE’ ( NMW). All types with additional printed label: ‘ Mannerheimia | cachemirica Coiffait, 1982 | Shavrin A. V. det. 2024’.

Holotype of Mannerheimia vicina Coiffait, 1982 ♂ ( Fig. 2 ; dissected): ‘Kashmir, Chandanwari | b.Pahalgam, 2800-3100 m | lg.H.Franz, Okt.1977 ’ <printed>, ‘TYPE’ <red, printed>, ‘ Mannerheimia | vicina | H. COIFFAIT det. 19[printed]76’ <handwritten in black>, ‘ Mannerheimia | cachemirica Coiffait, 1982 | Shavrin A. V. det. 2024’ <printed> ( NMW).

Redescription. Measurements (n=3): HW: 0.47–0.55; HL: 0.32–0.42; OL: 0.12; AL (holotype): 0.92; PL: 0.37; PW: 0.62–0.72; ESL: 0.75–0.90; EW: 0.84–0.96; MTbL (holotype): 0.54; MTrL (holotype): 0.30 (MTrL 1–4: 0.18; MTrL 5: 0.12); AW: 0.85–1.03; AedL: 0.35–0.39; BL: 2.55 (holotype of M. cachemirica )–2.65 (holotype of M. vicina ).

Habitus as in Figs 1–2 . Body reddish-brown, with distinctly darker head and abdomen (pronotum and elytra of holotype of M. cachemirica distinctly paler; paratergites of abdomen paler); antennomeres 6–11 brown; mouthparts, antennomeres 1–5, legs and paratergites of abdomen yellow. Body shiny, without microsculpture except for middle portion of head (holotype of M. cachemirica ) with very indistinct longitudinal meshes and abdomen with fine and dense transverse microreticulation (other specimens without meshes on head), distinctly coarser in abdominal tergite III–IV. Head with irregular and sparse, moderately large and deep punctation, denser in middle; middle portion of neck with very sparse and fine punctation; punctation of pronotum denser, larger and deeper than that on head, finer and sparser in middle (holotype of M. cachemirica ), with impunctate middle and mediobasal portion; scutellum without (holotype of M. cachemirica ) or with several very fine punctures in middle; punctation of elytra about as that on pronotum or denser, larger and deeper (holotype of M. cachemirica ), finer and sparser in middle along suture, punctures not forming disitnct longitudinal rows except for paratype with fused five rows of serial punctures; abdominal tergites with very fine and sparse punctation.

Head 1.3–1.4 times as broad as long, indistinctly elevated in middle, with very short and shallow anteocellar foveae about as long as diameter of ocellus; anterior portion between antennal insertion and anterior margins of eyes slightly concave. Eyes very large, convex. Ocelli large, well visible, situated at about level of posterior margins of eyes; distance between ocelli slightly greater than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi distinctly more than twice as long as preceding segment, from widest basal third gradually narrowed toward subacute apex. Antennae reaching basal portion of elytra when reclined; basal antennomere ovoid, slightly less than three times as long as broad, antennomere 2 distinctly shorter than basal antennomere, 3 narrower and slightly longer than 2, 4 small, about as long as broad, 5–6 distinctly longer and broader than 4, 7–8 slightly broader than 6, 9–10 indistinctly longer and broader than 8, apical antennomere about as long as two preceding antennomeres, from middle gradually narrowed toward subacute apex.

Pronotum 1.6–1.9 times as broad as long, 1.3 times as broad as head, widest in about middle, markedly more narrowed posteriad than anteriad; apical margin rounded, about as long as somewhat straight posterior margin; anterior angles widely rounded, indistincly protruded anteriad; posterior angles obtuse, with slightly sinuate laterobasal margins; each laterobasal portions with distinct wide impressions.

Elytra slightly broader than long, slightly widened apicad, twice longer than pronotum; apical margins straight or rounded. Surface of elytra in middle with distinct, irregular and transverse elevations between punctures (holotype of M. cachemirica ). Hind wings fully developed.

Metatarsus 1.8 times as long as metatibia.

Abdomen about as broad as elytra or slightly broader, with two small transverse tomentose wing-folding spots in middle of abdominal tergite V; apical margin of abdominal tergite VII with distinct narrow palisade fringe.

Male. Apical margins of abdominal tergite VIII and sternite VIII widely and rather deeply concave. Aedeagus with wide basal portion, gradually narrowed apically, with narrow median lobe and small rounded apex; parameres moderately wide, distinctly broadened in apical portions, slightly not reaching apex of median lobe; internal sac narrow and moderately long ( Fig. 6, 8 ). Lateral aspect of aedeagus as in Fig. 7 .

Female unstudied.

Comparative notes. Based on the general shape of the body and the aedeagus, M. cachemirica is similar to Nepalese M. emodensa , from which can be distinguished by the slightly narrower elytra, finer punctation of the pronotum and the elytra, slightly longer preapical antennomeres, and shorter parameres with markedly narrower apical portions.

Distribution. Mannerheimia cachemirica is known from two localities in Kashmir, India.

Bionomics. The holotype of M. vicina was collected at altitudes from 2800 to 3100 m a.s.l. Detailed bionomical data are unknown.

Remarks. Mannerheimia cachemirica was originally described based on three specimens from “ Inde, Cachemire, Aru…”. Mannerheimia vicina was originally described based on a male and a female from “ Inde, Cachemire, Chandanwari, près Pahalgam…”. I studied the holotype and the paratype of M. cachemirica and the holotype of M. vicina , and during the study I have not found sufficient morphological differences between them. The holotype of M. cachemirica has slightly more transverse pronotum and distinctly elevated surface between punctures in the middle of the elytra, but the aedeagus of it is similar to the type of M. vicina . The body of the paratype of M. cachemirica is almost identical to that of the holotype of M. vicina . Thus, I synonymize M. vicina with M. cachemirica , the species which has page priority in the same article ( Coiffait 1982).

Figures

FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm.

FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm.