Paroxyethira dzumac, Wells & Johanson, 2012

Wells, A. & Johanson, K. A., 2012, Review of the New Caledonian species of Paroxyethira Mosely, 1924 (Trichoptera: Hydroptilidae), Zootaxa 3478, pp. 330-344 : 337-338

Paroxyethira dzumac , new species

Figs 19, 20, 39

The male is similar to that of P. hamata , new species, having strongly asymmetric gonopods, but is distinguished by having the phallic sheath with an apical rather than subapical process, and pair of widely separated setae; and left gonopod more slender and lacking a brush of black setae. The form of the phallic sheath resembles that of P. atypica , but the apical process is short, not curving broadly. Both P. dzumac and P. hamata have an elongate dorsal process on the female terminalia, but in P. dzumac the process is narrow and straplike, not expanded subapically.

Male antennae each with 19 flagellomeres; female antennae each with 19 flagellomeres.

Forewing length. Male 2.1–2.7 mm (n=6); female 2.2–2.5 mm (n=10).

Male genitalia ( Figs 19, 20): Abdominal sternite VII with small sharp median spur. Median process on abdominal sternite VIII elongate spatulate; length around 3.5x width at middle. Abdominal segment IX dorsally with distal margin deeply concave; lateral angles produced into lateral lobes that are slender in ventral view; elongate, as long as ventral process of VIII. Gonopods quite dissimilar, right stout, tapered distally to narrow apex; brush of stout black setae on mesal margin, longest near mid length; closely joined at base to very slender left gonopod without brush of setae; bearing 3–4 setae distally. Phallic apparatus slender, straight, elongate; constricted at about 1/6 length at which constriction arises a short, threadlike titillator. Phallic sheath elongate, sheathlike; with straplike apical process and single large apical seta, second similar seta at about half length.

Female terminalia ( Fig. 39). Abdominal sternites VII–VIII fused; broad-based; abruptly constricted to form narrow tube for distal 2/3rds; apico-ventrally truncate; apico-dorsally bearing curved, straplike sclerotised process. Dorsally tergite VIII discrete; whole structure densely setose ventrally, setal bases prominent. Segment X undivided.

Holotype male: Province Sud, Mt. Dzumac, source stream of Ouinne River , downstream crosspoint to mountain track, 22°01.997'S, 166°28.486'E, 795 m, over about 30 m waterfall, 18.xi–4.xii.2003, Malaise trap, loc#031, leg. K.A. Johanson ( MNHP).

Paratypes: Same data as for holotype— 7 male, 21 female (1 male, 1 female on slides) ( SMNH); Province Sud, Mt. Dzumac, source stream of Ouinne River , near crosspoint to mountain track, 22°02.439'S, 166°28.646'E, 805 m, 18.xi.–4.xii.2003, Malaise trap, loc#029, leg. K.A. Johanson — 2 males ( SMNH) .

Other material examined: Province Sud, Mt. Dzumac, source stream of Ouinne River , near crosspoint to mountain track, 22°02.073'S, 166°28.460'E, 810 m, 18.xi–4.xii.2003, Malaise trap, loc#030, leg. K.A. Johanson — 1 female ( SMNH) ; Province Sud, Mt. Dzumac, source stream of Ouinne River , downstream crosspoint to mountain track, 22°01.997'S, 166°28.486'E, above waterfall, 795 m, 3.xii.2003, light trap, loc#061, leg. K.A. Johanson — 1 female ( SMNH) .

Etymology: Dzumac, derived from the type locality.

Remarks: In a sample from a stream on Mt. Dzumac, most of the males are uniformly of this form. Among them, however, are 2 males that resemble these others, but are distinct, having the spine on the phallic sheath stouter and subapical, and the left gonopod more slender and lacking a brush of stout dark setae. They are described below tentatively as a distinct species, P. hamata , new species.