Tiomaniella, Grall & Jäger, 2022

Grall, Elena & Jäger, Peter, 2022, Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae), Zootaxa 5169 (1), pp. 1-25 : 16

Tiomaniella gen. nov.

Figs 66–81 , 94–98

Type species. Tiomaniella ladam spec. nov.

Etymology. The generic name is derived from the name of Tioman Island. The gender is feminine.

Diagnosis. Small Sparassidae with TL <7 ( Figs 94–97 ) belonging to the subfamily Heteropodinae . Males of Tiomaniella gen. nov. can be distinguished from all other Heteropodinae by the subtegular functional conductor ( Figs 66–67 ). Females may be recognised by the following combination of characters: 1. Copulatory organs situated laterally with epigynal pockets located posteriorly, with first winding of internal duct system postero-mediad, 2. Median septum not recognisable, “fusion bubbles” indicating fused seams between lateral lobes and median septum and 3. Internal duct system simple with very broad ducts leading to large spermathecae ( Figs 70–72 ).

Description. See description of type species.

Distribution. Malaysia: Tioman Island ( Fig. 98 : red triangle).

Species included. Only the type species.

Figures

FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea.

FIGURES 66–69. Tiomaniella ladam gen. & spec. nov. holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis.

FIGURES 70–72. Tiomaniella ladam gen. & spec. nov. female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca.

FIGURES 73–81. Tiomaniella ladam gen. & spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left).

FIGURES 82–97. Heteropodinae gen. & spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. & spec. nov.; 86–89 Menarik kecil gen. & spec. nov.; 90–93 Micropoda davisae gen. & spec. nov.; 94–97 Tiomaniella ladam gen. & spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype).