Heterogomphus ulysses Burmeister 1847

Alvarez, Héctor Jaime Gasca, da, Claudio Ruy Vasconcelos, Fonseca & Ratcliffe, Brett C., 2008, Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon, Insecta Mundi 2008 (61), pp. 1-62 : 25

Heterogomphus ulysses Burmeister 1847

( Fig. 39-45 )

Heterogomphus ulysses Burmeister 1847: 232

DESCRIPTION. Length: 47.2-52.5 mm (males); 46.7-49.1 mm (females). Width: 22.7-27.3 mm (males); 21.4-24,5 mm (females). Color: Dark reddish brown to black.

Males. Head: Frons with long, recurved horn, sides of horn subparallel, apex rounded and truncate ( Fig 40 ). Eye canthus with rounded apex, surface rugose, anterior margin weakly crenulate. Clypeus reflexed, truncate, apex broad with notch at center ( Fig. 41 ), surface smooth to punctate. Antenna with 10 segments, club subequal in length to segments 2-7. Mandibles with a single, large tooth with rounded apex. Pronotum: Majors with broad, high prominence, posterior surface convexly rounded; apex cariniform, with emargination at center. Sides with broad, tuberculiform process ( Fig. 40 ). Base with marginal bead. Surface of fovea with small, sparse, setigerous punctures; setae dense, moderately long, reddish brown. Lateral margins rugose. Minors with frontal horn and pronotal prominence variably reduced. Elytra: Impressed sutural stria present. Surface occasionally with weak furrows in the central area. Deep punctures present near humerus. Sides with irregular rows of ocellate and deep punctures behind humerus. Apex with small setae, reddish brown setae. Pygidium: Surface rugopunctate, base with transverse band of long, dense, reddish brown setae. Apical half smooth, densely setigerous in small specimens. Legs: Protibia quadridentate. Apex of posterior tibia crenulate, with a row of thick spinules ( Fig. 42 ). First tarsomere of posterior tarsus long, quadrangular. Venter: Prosternal process long, apex rounded, densely setose; setae long, reddish brown. Parameres: Elongated, oval, strongly contracted forward apices ( Fig. 43-44 ).

Females. As males except in the following respects: Head: Frons with small, conical tubercle. Clypeal surface strongly punctate. Mandibles smaller in size. Pronotum: Surface of anterior half strongly rugose, posterior half smooth, with sparse punctures. Pygidium : Apex with small, reddish brown setae. In lateral view, basal half convex, anterior half weakly concave.

DIAGNOSIS. Heterogomphus ulysses may be distinguished by the following combination of characters: apex of metatibia with a row of spinules, sides of pronotum with a small tuberculiform process, pronotal fovea deep and densely setigerous, frontal horn long with bilobed apex.

Figures

Figure 39-44. Heterogomphus ulysses. 39) Habitus. 40) Male head and pronotum (lateral view). 41) Male head (dorsal view). 42) Metatibia (scale line: 5 mm). 43) Parameres frontal view. 44) Parameres lateral view (scale line: 1 mm).

Figure 45. Distribution of Heterogomphus ulysses in Brazilian Amazonia.