Dolichoraphidia, Liu, Xingyue, Lu, Xiumei & Zhang, Weiwei, 2016

Liu, Xingyue, Lu, Xiumei & Zhang, Weiwei, 2016, New genera and species of the minute snakeflies (Raphidioptera: Mesoraphidiidae: Nanoraphidiini) from the mid Cretaceous of Myanmar, Zootaxa 4103 (4), pp. 301-324 : 302-303

Genus Dolichoraphidia gen. nov.

( Figs. 1–4 )

Type species: Dolichoraphidia aspoecki sp. nov.

Diagnosis. Minute size. Head ovoid, with portion posterior to compound eyes slightly shorter than eye length and tapering caudad, and with clypeus slightly shorter than compound eye length; compound eyes large, ovoid; ocelli absent; antenna extremely long, at least twice as long as body length, with 100 or more flagellomeres. Pronotum nearly as long as or slightly longer than head, with a constant height along entire length in lateral view. All tibiae swollen, each tibia with a row of spines on distal half; tibial spurs absent. Wings narrowly elongate. Forewing: Costal space feebly broadened; pterostigma elongate, ~1/5× wing length, starting a distance beyond termination of ScP, with or without pterostigmal crossvein, proximally closed by a crossvein and distally closed by a veinlet of RA; ScP terminating into C at wing midpoint; posterior veinlet of RA continuous with stem of RA; RP with anterior branch proximally angled; two radial cells, one discal cell, and one or two discoidal cells present. Hindwing: Stem of MA proximally fused with MP; an oblique proximal mp-cua present; three radial cells, one discal cell, and one discoidal cell present.

Etymology. From “ dolich -”, long and Raphidia , a common genus-group name of Raphidioptera , in reference to the extremely long antennae of the new genus. Gender: feminine.

Remarks. The new genus is one of the most distinguishable snakeflies based on its extremely elongate antennae, with 100 or more flagellomeres. In all other known genera of Nanoraphidiini the antennae are much shorter, with 20–40 flagellomeres. The wing venation of Dolichoraphidia gen. nov. somewhat resembles that of Lebanoraphidia Bechly & Wolf-Schwenninger, 2011. However, in Dolichoraphidia gen. nov. the wings are much narrower, the Rp is distally bifurcated, and the metatibia is swollen. In Lebanoraphidia the wings are moderately broad, the Rp is simple, and the metatibia is not swollen.

Figures

FIGURE 1. Dolichoraphidia aspoecki gen. et sp. nov. A. Holotype EMTG BA- 001002, habitus in dorsal view; B. Same specimen, habitus in lateral view; C. Same specimen, head and thorax in lateral view; D. Paratype EMTG BA- 002179, habitus in dorsal view; E. Paratype EMTG BA- 002253, habitus in dorsal view. Scale bar = 1.0 mm.

FIGURE 2. Dolichoraphidia aspoecki gen. et sp. nov. A. Holotype EMTG BA- 001002, drawing of fore- and hindwing; B. Paratype EMTG BA- 002179, drawing of fore- and hindwing; C. Holotype EMTG BA- 001002, drawing of head, dorsal view; D. Same specimen, photograph of male genitalia, lateral view; E. Same specimen, drawing of male genitalia, lateral view; F. Paratype EMTG BA- 002179, drawing of female genitalia, lateral view. Scale bar = 1.0 mm (A – C, E – F) and 0.5 mm (D).

FIGURE 3. Dolichoraphidia engeli gen. et sp. nov. A. Holotype EMTG BA- 001300, habtius in dorsal view; B. Same specimen, fore- and hindwing; C. Same specimen, ovipositor in ventral view; D. Same specimen, drawing of ovipositor in ventral view; E. Paratype EMTG BA- 001252, habitus in dorsal view. Arrows indicating pterostigmal crossvein. Scale bar = 1.0 mm.

FIGURE 4. Dolichoraphidia engeli gen. et sp. nov., holotype EMTG BA 001300, drawing of fore- and hindwing. Scale bar = 1.0 mm.