Archaeosmylidia fusca, Makarkin & Yang & Ren, 2014

Makarkin, Vladimir V., Yang, Qiang & Ren, Dong, 2014, A new basal osmylid neuropteran insect from the Middle Jurassic of China linking Osmylidae to the Permian-Triassic Archeosmylidae, Acta Palaeontologica Polonica 59 (1), pp. 209-214 : 210

Archaeosmylidia fusca sp. nov.

Fig. 1 .

Etymology: From Latin fuscus, dark; in reference to dark coloration of the wing.

Holotype: Specimen CNU−NEU−NN2009103.An excellently−preserved, nearly complete forewing.

Type locality: Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.

Type horizon: Jiulongshan Formation, Bathonian, Middle Jurassic ( Gao and Ren 2006).

Description.—Forewing about 15.0 mm long (as preserved; estimated complete length about 15.5 mm), 5.7 mm wide. Costa anteriorly thick, especially basally. Trichosors prominent along hind margin, hardly visible along anterior apical margin, absent proximally. Costal space typical for Osmylidae , moderately dilated. Subcostal veinlets simple (two forked near costal margin); three basal−most markedly stouter than rest; strongly curved in medial part of costal space; closely spaced, especially in pterostigmal region. No crossveins between subcostal veinlets. Pterostigma barely evident. Subcostal space relatively broad, especially in distal portion; with 16 crossveins detected, of these three basal−most stouter, distal ones weak, poorly−discernible. Stem of Rs smooth, not zigzagged. R1 space very narrow, comparable in width with subcostal space. Proximal nygma clearly visible, located distal to origin of Rs1; distal nygma not detected. Ten crossveins, more or less regularly spaced in R1 space (between R1 and Rs); all well proximal to fusion of Sc and R1, no crossveins distal to this fusion detected. Crossveins between branches of Rs scarce, mostly form three irregular gradate series. Rs with 15 branches, not forked before outer gradate series; distal branches nearly straight, only slightly smoothly curved, not sinuous. Origin of M appears to originate from R at wing base. M deeply forked, slightly distal to origin of Rs1. MA and MP parallel before marginal dichotomous branching. Cu dividing into CuA and CuP near wing base; CuA and CuP sub−parallel before branching of CuP. CuA pectinate, with six oblique branches, mostly once forked. CuP relatively short, few−branched with two very oblique branches. 1A long, pectinate, with eight simple oblique branches. 2A pectinate, with seven simple oblique branches. 3A not preserved. Wing membrane fuscous, costal space slightly lighter except near costa.

Stratigraphic and geographic range.— Type locality and horizon only.

Figures

Fig. 1. Forewing of osmylid neuropteran insect Archaeosmylidia fusca gen. et sp. nov. (Osmylidae) from the Jiulongshan Formation of Daohugou, China. Holotype CNU−NEU−NN2009103. Photograph (A) and line drawing of venation (B). Scale bars 5 mm. Abbreviations: 1A, 2A, anal veins; C, costa; CuA, anterior cubitus; CuP, posterior cubitus; MA and MP, anterior and posterior branches of media (M); ng, nygma; R1, first branch of radius (R); Rs, radial sector; Sc, subcosta.