Ambanus lunatus ( Paik, 1976 )
publication ID |
https://doi.org/ 10.1080/00222930600916050 |
persistent identifier |
https://treatment.plazi.org/id/03FB717D-FFF0-0160-FE28-F9E8FD31FD8B |
treatment provided by |
Carolina |
scientific name |
Ambanus lunatus ( Paik, 1976 ) |
status |
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( Figures 1–3 View Figure 1 View Figure 2 View Figure 3 , 7 View Figure 7 , 8A, B View Figure 8 , 9 View Figure 9 C–F)
Coelotes lunatus Paik 1976, p 84 , Figures 4 View Figure 4 , 19–21 (Dǫ); Paik 1978, p 346, Figure 155 (ǫ).
Ambanus lunatus: Ovtchinnikov 1999, p 64 (transferred from Coelotes ); Namkung 2001, p 393, Figure 28.7a (ǫ); Namkung 2003, p 395, Figure 28.7a (ǫ).
Diagnosis
This species is similar to A. coreana , A. napoloυi Ovtchinnikov, 1999, A. paiki Ovtchinnikov, 1999 , and A. quadratiυulυus ( Paik, 1974) in having female epigynum with atrium very broadly oval, situated posteriorly near epigastric furrow; copulatory pore deep, linear on inner lateral part; copulatory ducts broadly curved with transparent membranes in both posterior margins; male palpal organ with cymbial furrow less than one-third cymbial length; large embolus with curved distal part; and conductor saucer-like, with a rounded distal end situated on centre of papal organ. Ambanus lunatus female can be distinguished by the presence of a semicircular swollen atrial hood; distinctive atrial septum triangularly expanded, originating in posterior plate; chelicerae with three promarginal teeth; and spermathecal stalk overlapped in the middle; the male by an embolus with two distal parts with cup-shaped processes and another divided into two protrusions facing the retrolateral side; cymbial furrow short, about one-fifth the cymbial length.
Description
Measurements (mm). Female/male: habitus length 12.3/12.6; carapace length 4.8/5.7, carapace width 3.3/3.8, carapace height 3.1/4.1; cheliceral length 2.7/2.7, cheliceral width 1.3/1.2, cheliceral fang length 1.4/1.4; sternum length 2.6/2.4, sternum width 2.0/2.0; endite length 1.7/1.7, endite width 1.0/1.0; labium length 0.9/1.0, labium width 0.8/0.9; clypeal height 0.2/0.2; AER 1.0/0.7, PER 1.3/0.9, AME 0.1/0.1, ALE 0.2/0.2, PME 0.2/ 0.2, PLE 0.2/0.2. Eye formula ALE5PLE5PME.AME/ALE5PLE5PME.AME. Palp 5.2/5.4 (1.7/1.9, 0.8/0.6, 1.0/0.4, 1.7/2.5). First leg 12.0/15.2 (3.4/4.1, 1.6/1.9, 2.7/3.4, 2.7/3.6, 1.6/2.2), second leg 10.9/13.7 (3.1/3.8, 1.6/1.8, 2.2/2.8, 2.5/3.3, 1.5/2.0), third leg 10.4/12.9 (2.9/3.6, 1.5/1.7, 1.8/2.2, 2.7/3.5, 1.5/1.9), fourth leg 14.0/17.0 (3.8/4.5, 1.7/ 1.8, 2.9/3.6, 3.9/4.9, 1.7/2.2). Leg formula IV I II III/IV I II III. Abdomen length 6.9/6.0, abdomen width 4.5/3.8, abdomen height 4.7/4.0; ALS 0.5/05, PLS 0.9/0.9 (0.4/0.4+0.5/ 0.5).
Female. Medium-sized spider slightly shorter than male, found under stones and fallen leaves on the ground of the forests. Carapace elongate, 1.2 times longer than width, moderately narrowed in eye area, and distinctly longitudinal fovea on middle ( Figure 1A View Figure 1 ). AER straight and PER slightly procurved in frontal view; AME smaller than other eyes, separated by as much as their diameter ( Figure 1B View Figure 1 ); and eye ratio 24. Clypeal height slightly longer than AME diameter and without chilum. Chelicerae with numerous long setae; lateral condyle yellowish brown; three promarginal teeth, middle one largest; and two retromarginal teeth subequal in size ( Figure 1C View Figure 1 ). Endites reddish brown, widest at midpart; and labium rectangular, longer than wide ( Figure 1D View Figure 1 ). Sternum shield-shaped, widest at second coxae, and slightly projected between fourth coxae ( Figure 1E View Figure 1 ). Palp ( Figure 1 View Figure 1 G– I) composed of one claw with seven side teeth; tibia with nine trichobothria in three rows (5d-1d-3r), tarsus five in one (5r) and femur with three spines, tibia with five (four, 2-2 on dorsal; one, 0-1 on prolateral), tarsus 14 (five, 2-2-1 on prolateral; five, 0-5 on retrolateral; four, 0-4 on ventral). Legs ( Figures 1F View Figure 1 , 7 View Figure 7 ) yellowish brown without ring patterns; length of first leg (patella+tibia) always shorter than carapace length; trochanters not notched; tibia with 11–17 trichobothria in four rows (4p-5d-4d-4r on first leg, 4p-5d-4d-4r on second, 3p-5d-4d-4r on third, 1p-6d-3d-1r on fourth), metatarsi six to seven in one row (seven on first leg and second, six on third and fourth), tarsi six to seven in one row (seven on first leg, third and fourth, six in second); tarsal organ situated close to distal end of tarsus, slightly anterior of distal trichobothrium; tarsi with three claws, upper claws with 9–12 side teeth (12 on first leg, 11 on second, nine on third and fourth), lower with one to two (one on first leg, two on second, third and fourth). Leg spination ( Table I): leg I femur with four spines, tibia seven (one, 0-0-1 on prolateral; six, 2-2-2a on ventral), metatarsus eight (two, 1-1 on prolateral; six, 2-2-2 on ventral), tarsus without spine; leg II femur with four spines, tibia six (one, 0-0-1 on prolateral; five, 1-2-2a on ventral), metatarsus nine (three, 0-1-2 on prolateral; six, 2-2-2 on ventral), tarsus without spine; leg III femur five spines, tibia 10 (four, 1-0-1 on prolateral; retrolateral, six, 2-2-2a on ventral), metatarsus 16 (10, 1-2-2 on prolateral; retrolateral, six, 2-2-2 on ventral), tarsus five (one, 0-1 on retrolateral; four, 0-1- 1 on prolateral and ventral); leg IV femur four spines, tibia 12 (two, 1-0-1 on dorsal; four, 1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 16 (10, 1-2-2 on prolateral and retrolateral; six, 2-2-2 on ventral), tarsus five (four, 0-1-1 on prolateral and retrolateral; one, 0-1 on ventral). Abdomen ovoid with scattered brownish yellow spots and chevrons on dorsal side ( Figure 1A View Figure 1 ). Cribellum absent. Spinnerets brown; ALS cylindrical with horizontal apical part; and PLS almost twice as long as other spinnerets and two segments, second segment slightly longer than proximal one.
Female epigynum and genitalia ( Figures 1J View Figure 1 , 2A, B View Figure 2 , 9C, D View Figure 9 ): epigynal teeth absent; atrial septum distinctive, triangularly expanded, originating in posterior plate; atrial slit or true copulatory pore deep, linear on inner part; atrial hood semicircular and swollen horizontally; copulatory ducts broadly curved with transparent membranes; spermathecal heads small cylindrical processes, situated at anterior of spermathecae; spermathecae short, overlapped, with indistinct stalks and bases; fertilization ducts very small, arising from the posterior spermathecae.
Male. Medium-sized spiders slightly longer than female. Carapace elongate, 1.6 times as long as width, moderately narrowed in eye area, and distinctly longitudinal fovea on middle ( Figure 3A View Figure 3 ). AER and PER almost straight in frontal view; AME smaller than other eyes, separated by slightly less than their diameter, and eye ratio 26 ( Figure 3B View Figure 3 ). Clypeal height slightly longer than AME diameter and without chilum. Endites reddish brown, widest at mid-part; and labium rectangular, slightly longer than wide ( Figure 3C View Figure 3 ). Sternum shieldshaped, widest at second coxae, and not produced between fourth coxae ( Figure 3D View Figure 3 ). Chelicerae with numerous long setae, lateral condyle yellowish brown, with three promarginal teeth, middle one largest, and two retromarginal teeth subequal in size ( Figure 3E View Figure 3 ). Legs ( Figures 3F View Figure 3 , 7 View Figure 7 ) yellowish brown; length of first leg (patella+tibia) always shorter than carapace length; trochanters not notched; tibia with 16–24 trichobothria in four rows (5p-7d-4d-6r on first leg, 6p-6d-6d-6r on second, 4p-6d-5d-7r on third, 2p-6d- 4d-4r on fourth), metatarsi six to eight in one row (eight on first, third and fourth leg; six on second), tarsi 8–10 in one row (10 on first leg, nine on second and fourth, eight on third); tarsal organ situated close to distal end of tarsus, slightly anterior of distal trichobothrium; tarsi with three claws, upper claws with 9–13 side teeth (13 on first leg, 12 on second, 10 on third, nine on fourth), lower with zero to one (zero on first leg, one on second, third and fourth). Leg spination ( Table I): leg I femur with four spines, tibia seven (one, 0-0-1 on prolateral; six, 2-2-2a on ventral), metatarsus eight (two, 0-1-1 on prolateral; six, 2-2-2 on ventral), tarsus without spine; leg II femur with five spines, tibia six (one, 0-1-1 on prolateral; five, 1-1-2a on ventral), metatarsus nine (three, 0-1-2 on prolateral; six, 2-2-2 on ventral), tarsus without spine; leg III femur with six spines, tibia 10 (four, 0-1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 17 (one, 1-0-0 on dorsal; 10, 1-2-2 on prolateral and retrolateral; six, 2-2-2 on ventral), tarsus four (three, 1-1-1 on prolateral; two, 0-1-1 on retrolateral; one, 0-1 on ventral); leg IV femur with four spines, tibia 12 (two, 1-0-1 on dorsal; four, 1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 17 (one, 1-0-0 on dorsal; 10, 1-2-2 on prolateral and retrolateral; six, 2-2-2 on ventral), tarsus six (three, 1-1-1 on prolateral; two, 0-1-1 on retrolateral; one, 0-1 on ventral). Abdomen ovoid, with scattered brownish yellow spots and chevrons on dorsal side ( Figure 3A View Figure 3 ). Cribellum absent. Spinnerets brown, ALS cylindrical, with horizontal apical part and PLS almost twice as long as other spinnerets and with two segments, second segment slightly longer than proximal one.
Male palp ( Figures 3 View Figure 3 G–J, 8A, B): patellar apophysis absent; RTA modified with ITA; cymbial furrow short, about one-fifth cymbial length; tegular sclerite weakly sclerotized and longitudinally situated on tegulum; conductor broadly saucer-like with many minute denticles, rounded distal end situated on centre of papal organ; conductor dorsal apophysis situated on upper embolus; embolus broad and short, wound counterclockwise, two distal parts with cup-shaped processes and another divided into two protrusions facing the retrolateral side.
Specimens examined
Two females, 21 July 1984, two females, 24 July 1984, two females, 22 September 1984, three females, 3 October 1984, five females, five males, 3 November 1984, two females, 11 November 1984, one female, 14 April 1985, two females, 21 April 1985, three females, 3 May 1985, one female, four males, 18 May 1985, Mt. Yebong, Gyonggi-do, K. S. Lee; one male, 6 September 1986, same locality, J. H. Kim; one female, 19 January 2006, Simbog cave, Chungcheongbuk-do, B. W. Kim.
Distribution
Korea (Mt. Yebong, Mt. Songni).
Ambanus coreana ( Paik, 1992) nov. comb.
( Figures 4–6 View Figure 4 View Figure 5 View Figure 6 , 7 View Figure 7 , 8 View Figure 8 C–F, 9A, B)
Coelotes sp. (A) Namkung and Yoon 1975, p 39, Figure 3 View Figure 3 (ǫ).
Alloclubionoides coreana Paik 1992, p 9 , Figures 1–8 View Figure 1 View Figure 2 View Figure 3 View Figure 4 View Figure 5 View Figure 6 View Figure 7 View Figure 8 (D „, in Clubionidae View in CoL ); Namkung 2003, p 448, Figure 34.31a, b („). Misidentified male.
Alloclubionoides coreanus: Platinick 1997, p 702 (in Clubionidae View in CoL ).
Coelotes paikaeunensis Kim and Jung 1993, p 2 , Figures 1 View Figure 1 , 6 View Figure 6 , 7 View Figure 7 (Dǫ); Namkung 2001, p 390, Figure 28.4a, b (ǫ). Misidentified female.
Ambanus paikaeunensis: Wang 2002, p 28 (transferred from Coelotes ); Namkung 2003, p 394, Figure 28.6a, b (ǫ). Misidentified female.
Ambanus lunatus: Namkung 2001, p 393 , Figure 28.7b („); Namkung 2003, p 395, Figure 28.7b („); Kim and Cho 2002, p 185, Figures 361–366 (ǫ „). Misidentified.
Diagnosis
This species is similar to A. lunatus , A. napoloυi Ovtchinnikov, 1999, A. paiki Ovtchinnikov, 1999 , and A. quadratiυulυus. The female of A. coreana can be distinguished by the presence of an oblique (/ \) shaped atrial hood; atrial septum small and broadly obscure; chelicerae with two promarginal teeth; spermathecal stalk overlapped in lower; and the male by an embolus with distal part with cup-shaped process attached slender embolus tip facing down; cymbial furrow short at about one-quarter the cymbial length.
Description
Measurements (mm). Female/male: Habitus length 12.7/10.3; carapace length 6.7/5.5, carapace width 4.0/3.4, carapace height 3.7/3.6; cheliceral length 3.0/2.5, cheliceral width 1.8/1.3, cheliceral fang length 1.7/1.3; sternum length 3.0/2.5, sternum width 2.4/1.9; endite length 2.1/1.6, endite width 1.1/0.9; labium length 0.9/0.9, labium width 0.8/0.8; clypeal height 0.2/0.2; AER 0.8/0.9, PER 1.1/1.2, AME 0.1/0.1, ALE 0.2/0.2, PME 0.2/0.2, PLE 0.2/0.2. Eye formula ALE5PLE5PME.AME/ALE5PLE5PME.AME. Palp 6.0/5.4 (2.0/1.9, 0.9/0.6, 1.2/0.4, 1.9/2.5). First leg 14.2/14.0 (4.0/3.8, 2.0/1.8, 3.1/3.1, 3.2/3.3, 1.9/ 2.0), second leg 12.2/12.6 (3.3/3.5, 1.8/1.6, 2.4/2.5, 3.0/3.1, 1.7/1.9), third leg 11.8/12.0 (3.2/3.3, 1.7/1.6, 2.1/2.1, 3.2/3.3, 1.6/1.7), fourth leg 16.1/16.0 (4.1/4.2, 2.0/1.7, 3.4/3.2, 4.5/4.6, 2.1/2.3). Leg formula IV I II III/IV I II III. Abdomen length 8.1/5.8, abdomen width 5.3/3.6, abdomen height 5.0/3.4; ALS 0.5/0.5, PLS 1.1/1.0 (0.5/0.5+0.6/0.5).
Female. Medium-sized spiders distinctly longer than male, found under stones and fallen leaves on the ground of the forests. Carapace elongate, 1.7 times as long as width, moderately narrowed in eye area, and distinctly longitudinal fovea on middle ( Figure 4A View Figure 4 ). AER and PER slightly procurved in frontal view; AME smaller than other eyes, separated by as much as their diameter; and eye ratio 28 ( Figure 4B View Figure 4 ). Clypeal height twice as long as AME diameter and without chilum. Chelicerae with numerous long setae; lateral condyle yellowish brown; two promarginal teeth, outer larger than inner; and two retromarginal teeth subequal in size ( Figure 4C View Figure 4 ). Endites reddish brown, widest at mid-part; and labium rectangular, longer than wide ( Figure 4D View Figure 4 ). Sternum shield-shaped, widest at second coxae, and slightly projected between fourth coxae ( Figure 4E View Figure 4 ). Palp ( Figure 4 View Figure 4 G–I) composed of one claw with seven side teeth; tibia with nine trichobothria in three rows (5d-1d-3r), tarsus five in one (5r) and femur with four spines, tibia six (three, 1-2 on dorsal and prolateral), tarsus 17 (one, 1-0 on dorsal; six, 3-2-1 on prolateral; five, 2-2-1 on retrolateral; five, 1-0-4 on ventral). Legs ( Figures 4F View Figure 4 , 7 View Figure 7 ) yellowish brown without ring patterns; length of first leg (patella+tibia) always shorter than carapace length; trochanters not notched; tibia with 22– 25 trichobothria in four (five on second leg) rows (6p-7d-5d-7r on first leg, 6p-6d-6d-2r-4r on second, 5p-7d-5d-5r on third, 6p-6d-6d-5r on fourth), metatarsi seven to eight in one row (seven on first, third and fourth leg, eight on second), tarsi eight to nine in one row (nine on first leg, eight on second, third and fourth); tarsal organ situated close to distal end of tarsus, slightly anterior of distal trichobothrium; tarsi with three claws, upper claws with 9–12 side teeth (12 on first leg, 11 on second, nine on third and fourth), lower with two to three (two on first, second and third leg, three on fourth). Leg spination ( Table I): leg I femur with four spines, tibia seven (one, 0-0-1 on prolateral; six, 2-2-2a on ventral), metatarsus eight (two, 1-0-1 on prolateral; six, 2-2-2 on ventral), tarsus without spine; leg II femur with four spines, tibia seven (two, 0-1-1 on prolateral, five, 2-1-2a on ventral), metatarsus 11 (three, 1-1-1 on prolateral; two, 0-1-1 on retrolateral; six, 2-1-3 on ventral), tarsus one (1-0-0) on ventral; leg III femur eight spines, tibia 10 (four, 1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 16 (10, 1-2-2 on prolateral and retrolateral; six, 2-2-2 on ventral), tarsus five (two, 0-2 on prolateral; one, 0-1 on retrolateral; two, 0-1-1 on ventral); leg IV femur four spines, tibia 10 (four, 1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 15 (five, 1-2-2 on prolateral; four, 1-1-2 on retrolateral; six, 2-2-2 on ventral), tarsus six (0-1-1 on prolateral, retrolateral and ventral). Abdomen ovoid with scattered brownish yellow spots and chevrons on dorsal side ( Figure 4A View Figure 4 ). Cribellum absent. Spinnerets brown, ALS cylindrical, with horizontal apical part and PLS almost twice as long as other spinnerets and with two segments, second segment slightly longer than proximal one.
Female epigynum and genitalia ( Figures 4J View Figure 4 , 5A, B View Figure 5 , 9A, B View Figure 9 ): epigynum teeth absent; atrial septum small, broadly obscure, originating in posterior plate; atrial slit or true copulatory pore deep, linear on inner retrolateral part; atrial hood oblique (/ \) shaped; copulatory ducts broadly curved with transparent membranes; spermathecal heads small cylindrical processes, with some slender tentacles on distal part, situated at anterior of spermathecae; spermathecae short, overlapped, with indistinct stalks and bases; fertilization ducts very small, arising from the posterior margin of spermathecae.
Male. Medium-sized spider distinctly shorter than female. Carapace elongate, 1.6 times as long as width, moderately narrowed in eye area, and distinctly longitudinal fovea on middle ( Figure 6A View Figure 6 ). AER straight and PER slightly procurved in frontal view; AME smaller than other eyes, separated by slightly less than their diameter and eye ratio 35 ( Figure 6B View Figure 6 ). Clypeal height slightly longer than AME diameter and without chilum. Endites reddish brown, widest at mid-part; and labium rectangular, slightly longer than wide ( Figure 6C View Figure 6 ). Chelicerae with lateral condyle yellowish brown with three promarginal teeth, middle one largest, and two retromarginal teeth subequal in size ( Figure 6D View Figure 6 ). Sternum shield-shaped, widest at second coxae, and not projected between fourth coxae ( Figure 6E View Figure 6 ). Legs ( Figures 6F View Figure 6 , 7 View Figure 7 ) yellowish brown; length of first leg (patella+tibia) always shorter than carapace length; trochanters not notched; tibia with 22–25 trichobothria in four rows (7p- 6d-6d-5r on first leg, 7p-6d-5d-6r on second, 6p-6d-4d-6r on third, 6p-7d-3d-6r on fourth), metatarsi 6–13 in one row (13 on first leg, eight on second, six on third, seven on fourth), tarsi eight to nine in one row (nine on first leg, eight on second and third, nine on fourth); tarsal organ situated close to distal end of tarsus, slightly anterior of distal trichobothrium; tarsi with three claws, upper claws with 10–13 side teeth (13 on first leg, 12 on second, 11 on third, 10 on fourth), lower with zero to three (zero on first leg, three on second and fourth, one on third). Leg spination (Table II): leg I femur with four spines, tibia seven (one, 0-0-1 on prolateral; six, 2-2-2a on ventral), metatarsus seven (two, 0-1-1 on prolateral; six, 2-1-2 on ventral), tarsus without spine; leg II femur with four spines, tibia 10 (two, 1-1 on prolateral; five, 1-1-2a on ventral), metatarsus nine (three, 1-2 on prolateral; six, 2-2-2 on ventral), tarsus without spine; leg III femur six spines, tibia 10 (four, 1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 16 (10, 1-2-2 on prolateral and retrolateral; six, 2-2-2 on ventral), tarsus four (two, 0-1-1 on prolateral; two, 0-1-0 on retrolateral and ventral); leg IV femur four spines, tibia 10 (four, 0-1-1 on prolateral and retrolateral; six, 2-2-2a on ventral), metatarsus 17 (10, 1-2-2 on prolateral and retrolateral; seven, 3-2-2 on ventral), tarsus five (four, 0-1-1 on prolateral and retrolateral; one, 0-1 on ventral). Abdomen ovoid with scattered brownish yellow spots and chevrons on dorsal side ( Figure 6A View Figure 6 ). Cribellum absent. Spinnerets brown; ALS cylindrical with horizontal apical part; and PLS almost twice as long as other spinnerets and with two segments, second segment as long as proximal one.
Male palp ( Figures 6 View Figure 6 G–J, 8C, D): patellar apophysis absent; RTA modified with ITA; cymbial furrow short, about one-quarter of cymbial length; tegular sclerite weakly sclerotized and longitudinally situated on tegulum; conductor broadly saucer-like with many minute denticles, rounded distal end positioned on centre of papal organ; conductor dorsal apophysis situated on distal embolus; embolus broad and short, wound counterclockwise, distal part with cup-shaped process attached slender embolus tip facing down.
Specimens examined
One female, 9 October 1973, Mt. Gamak, near the demilitarized zone, Gyeonggi-do, J. Namkung; one female, 4 June 1993, three females, two males, 9 December 1993, Mt. Paikwun , Gyeonggi-do, C. H. Jung; one female, five males, 23 October 2001, two females, five juveniles, 11 August 2004, Korean national arboretum of Gwangrung ( KNAG), Gyeonggi-do, T. S. Kwon .
Distribution
Korea (KNAG, Mt. Paikwun, Mt. Gamak).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
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Phylum |
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Class |
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Order |
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Family |
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Genus |
Ambanus lunatus ( Paik, 1976 )
Kim, Byung-Woo & Lee, Woncheol 2006 |
Ambanus coreana ( Paik, 1992 )
Kim & Lee 2006 |
Ambanus paikaeunensis:
Wang 2002: 28 |
Ambanus lunatus
: Namkung 2001: 393 |
Ambanus lunatus
: Ovtchinnikov 1999: 64 |
Alloclubionoides coreanus:
Platinick 1997: 702 |
Coelotes paikaeunensis
Kim and Jung 1993: 2 |
Alloclubionoides coreana
Paik 1992: 9 |
Coelotes lunatus
Paik 1976: 84 |
Coelotes
Blackwall 1841 |
Coelotes
Blackwall 1841 |