Agnathus decoratus ( Germar, 1818 )

Jelínek, Josef & Kubáň, Vítězslav, 2009, A review of the genus Agnathus (Coleoptera: Pyrochroidae: Agnathinae), with description of Agnathus secundus sp. nov. from China, Acta Entomologica Musei Nationalis Pragae 49 (1), pp. 253-281 : 262-266

Agnathus decoratus ( Germar, 1818)

( Figs. 5–9 )

Notoxus decoratus Germar, 1818: 229 . Type locality: Central Germany, Sachsen-Anhalt, Halle env., near river.

Agnathus ornatus Megerle von Mühlfeld’ in GERMAR (1818): 232 (unavailable name). – GEMMINGER & HAROLD (1869): 2062 (catalogue; in synonymy; unavailable name); SEIDLITZ (1898): 363 (comment; unavailable name); ABDULLAH (1974): 24 (comment; unavailable name).

Agnathus decoratus: GERMAR (1825) :(4), pl. 4. (characters); DEJEAN (1834): 216 (catalogue; France); DEJEAN (1836): 237 (catalogue; France); DUPONCHEL (1842): 199 (dictionary); KIESENWETTER (1849): 53 (catalogue); LAFERTÉ- SÉNECTÈRE (1849): 295 (characters; biology; distribution; localities in France and Germany); GAUBIL (1849): 237 (list of species; distribution); REDTENBACHER (1849): 609 (characters; locality in Austria); MULSANT & REY (1856a): 114 –118, pl. I, figs. 1–4 (descriptions of larva and pupa); MULSANT & REY (1856b): 114 –118, pl. I, figs. 1–4 (descriptions of larva and pupa); DOHRN (1858): 69 (catalogue); REDTENBACHER (1858): 635 (characters; locality in Austria); BACH (1859): 282 (characters); SCHAUM (1859): 75 (catalogue); GUTFLEISCH & BOSE (1859): 430 (characters); LACORDAIRE (1859): 523 (larva; pupa; characters), 532 (characters; distribution); SCHAUM (1862): 77 (catalogue); FAIRMAIRE (1863): 458 (characters), pl. 100, figs. 500, 500a,b; MARSEUL (1863): 191 (catalogue); MARSEUL (1866): 86 (catalogue); BACH (1866): xlii (list of species; Germany); MULSANT & REY (1866a): 269, figs. 1–4 (characters of adult), 271 (biology; distribution; locality in France), 272, figs. 5–7 (characters of larva), 274, fig. 8 (characters of pupa), 282 (list of species); MULSANT & REY (1866b): 6 (list of species), 181, figs. 1–4 (characters of adult), 183 (biology; distribution; locality in France), 184, figs. 5–7 (characters of larva), 186, fig. 8 (characters of pupa); STEIN (1868):86 (catalogue); RÓZSAY (1868): [page?] (list of species; locality in Slovakia); GEMMINGER & HAROLD (1869): 2062 (catalogue; in Pythidae ); REITTER (1870): 123 (list of species; biology; locality in Moravia); BRANCSIK (1871): 73 (list of species; biology; locality in Slovenia); STIERLIN & GAUTARD (1871): 224 (list of species; locality in Switzerland); ABEILLE DE PERRIN (1874): [page?] (study); REDTENBACHER (1874): 138 (characters; locality in Austria); BERTOLINI (1874): 155 (list of species; distribution in Italy); SEIDLITZ (1875): 368 (list of species; distribution); GOZIS (1875): 65 (catalogue; France); STEIN & WEISE (1877): 123 (catalogue; distribution); DELHERM DE LARCENNE (1877): [page?] (list of species; locality in France); BAUDI DI SELVE (1877): 407 (localities in Italy; biological note); SCHNEIDER & LEDER (1878): 248 (list of species; locality in Georgia); SCHLOSSER KLEKOVSKI (1878): 568 (key), 571 (characters; localities in Croatia); BUYSSON (1879): 151 (colour plate with 1 fig.); BUYSSON (1880): 72 (biology); HEYDEN (1883): 138 (catalogue; distribution); REY (1884):428 (comments in favour of the placement in Anthicidae ); MARSEUL (1887): 344 (catalogue); BIELZ (1887): 81 (list of species; locality in Romania); SCHILSKY (1888): 92 (list of species; distribution in Germany, Prussia, Silesia, Moravia and Austria); QUEDENFELDT (1888): (8) (locality in Germany); BAUDI DI SELVE (1889): 145 (list of species; distribution in Piemonte); GERHARDT (1890):295 (list of species; locality in Moravia); SEIDLITZ (1890): 528 (list of species; distribution); STIERLIN (1890): 160 (list of species; localities in Switzerland); SEIDLITZ (1891): 567 (list of species; distribution); HEYDEN (1891): 256 (catalogue; distribution); KUTHY (1897): 133 (list of species; localities in Hungary, Slovakia and Romania); SEIDLITZ (1898): 360 (larval characters), 362 (characters; comments; biology; distribution; localities); BUYSSON (1898): 129 (biology); DESBROCHERS DES LOGES (1900): 4 (characters; localities in France), 39 (list; distribution in France); WARNIER (1901): 121 (catalogue); BARTHE (1902): 57 (list of species; France); REITTER (1906): 462 (catalogue; distribution); FORMÁNEK (1907): 52 (biological note); EGGERS (1908): 5 (biological note); REITTER (1908): 21 (biological data; locality in Moravia); SCHIL- SKY (1908): 126 (list of species; distribution in Germany, Prussia, Silesia, Moravia and Austria); BUYSSON (1910): 106 (biology; distribution; localities in France); GERHARDT (1910): 279 (list of species; locality in Moravia); REITTER (1911): 413 (characters in key; biological data; localities in Germany); FALCOZ (1912): 19, 51 (list, in Lagriidae ; note; locality in France); KUHNT (1912): 730 (key; distribution); PETRI (1912): column 231 (list of species; locality in Romania); ŁOMNICKI (1913): 117 (list of species; Silesia); JAKOBSON (1915): 1017 (list of species; distribution; locality in Georgia; bibliography), pl. 79, fig. 20; PURKYNĚ (1926): 94 (biological data; locality in western Ukraine); ABOT (1928): 234 (list of species; biological data; locality in France); WINKLER (1928): column 900 (catalogue; distribution); STAMMER (1929): 24 (biological data); FLEISCHER (1930): 299 (list of species; biological data); PORTEVIN (1931): 109 (key; characters), fig. 214; GALIBERT (1932): 307 (list of species; biological data; locality in France); FLEISCHER (1933): 105 (list of species; biological data); PORTA (1934): 78 (key; distribution in Italy); ROUBAL (1934): 177 (biological data and reflections on bionomy; localities in former Czechoslovakia); ROUBAL (1936): 298 (list of species; biology; distribution; localities in Slovakia and Transcarpathian Ukraine; bibliography); BORCHMANN (1936): 539, pl. 8, fig. 63e,f, pl. 9, fig. 70 (monograph; distribution); SAINTE- CLAIRE DEVILLE (1937): 315 (list of species; distribution in France); BORCHERT (1938): 84 (list of species; distribution); TRÄGER (1937): 7 (list; locality in Hungary); HUDEČEK (1940): 184 (biological data; locality in Moravia, Czech Republic); ROUBAL (1941): 270, 276 (biological note); PITTIONI (1943): 129 (list of species; locality in Austria); HORION (1951): 349 (list of species; distribution in Central Europe); BORCHERT (1951): 161 (list of species; from type locality only); PFEFFER (1955): 53 (biological data; in Lagriidae ); HORION (1956): 173 (list of species; biology; distribution; localities in Europe; bibliography); KASZAB (1957): 73 (characters; biological data; distribution; not in Hungary – but see also KOVÁCS et al. (2000)), fig. 53; BALTHASAR (1957): 444 (key; biological data); HLISNIKOVSKÝ (1958): 213 (collecting circumstances; localities in Moravia and Bulgaria); MEDVEDEV (1965):351 (key of species); KASZAB (1969): 215 (key; distribution); ABDULLAH (1974): 20 (comments), 22 (key), 24 (catalogue); FRANZ (1974): 299 (list of species; distribution; localities in northern Austria; in Lagriidae ); MAMAEV (1976): 642 (larva; description); IABLOKOFF- KHNZORIAN (1985): 223 (revision; characters; biology; distribution), fig. 12(1–6); LUCHT (1987): 226 (list of species; distribution in Central Europe); BURAKOWSKI et al. (1987): 159 (catalogue; biological data; distribution; records from Silesia probably refer to Moravian Silesia in the Czech Republic; unknown from Poland); KOCH (1989): 329 (distribution; biology); RENNER (1991): 135 (list; locality in Westphalia, Germany); JELÍNEK (1993): 116 (list of species; distribution in former Czechoslovakia); ANGELINI et al. (1995): 19 (list of species; distribution in Italy); ALEXAND- ROVICH (1995): 72 (list of species; locality in Belarus; in Cononotidae ); ALEXANDROVICH et al. (1996): 44 (catalogue; Belarus); KLAUSNITZER (1996): 316 –317 (larva; characters; biological data), figs. 1–3; KÖHLER & KLAUSNITZER (1998): 126 (list of species; distribution in Germany); SZALÓKI (1999): 197 (locality in Hungary; in Pyrochroidae ); KOVÁCS et al. (2000): 201 ( Agnathus -bibliography for Hungary; localities in Hungary; in Pyrochroidae ); JÁSZAY (2001): 139 (list of species; locality in Slovakia), 193, 203 (conservation); JÄCH et al. (2002): 211 (faunistics; biological data; distribution; localities in Austria, Czech Republic and European part of Turkey; in Pyrochroidae ); BOUYON & VAN MEER (2004): 87 (new observations; distribution; localities in France; in Pyrochroidae ); HŮRKA (2005): 211 (list of species; in Pyrochroidae ); JELÍNEK (2005): 524 (red list; in Agnathidae ); NARDI (2007): web page (list of species; distribution: Italian mainland only(!); dating of species name 1825); THÉRY (2007): 165 (locality in France; in Pyrochroidae ); POLLOCK & YOUNG (2008): 417 (catalogue; general distribution); NIKITSKY et al. (2008): 318 (list of species; localities in southern European Russia; in Agnathidae ).

Geographic distribution ( Fig. 8 )

POLOCK & YOUNG (2008: 417) listed A. decoratus from the following countries: Austria, Bosnia Herzegovina, Belarus, Croatia, Czech Republic, France, Georgia, Germany, Italy, Poland, Romania, Slovakia, Russia: South European Territory, Switzerland, Ukraine and Asian Turkey. Occurrence in Bulgaria (published by ROUBAL 1936) and in Slovenia is not mentioned. The record from ‘Steiermark... bei St. Leonhard’ ( BRANCSIK 1871, subsequently HORION 1956 etc.) refers to nowadays Slovenia (Lenart v Slovenskih Goricah).A record from the Asian part of Turkey is probably based on misinterpretation of data from JÄCH et al. (2002), who recorded the species from the European part of Turkey (see below).

Published records. AUSTRIA: ‘Österreich’, early 19 th century [as ‘ Agnathus ornatus Megerle von Mühlfeld’] ( GERMAR 1818; LAFERTÉ- SÉNECTÈRE 1849; SEIDLITZ 1898). KÄRNTEN: Völkermarkt Bezirk, Gallizien, Vellach river bank, 28.vii.2001, R. Schuh leg. and det., 5 spec., RSCW [1 spec. in CHCV, C. Holzschuh pers. comm.] ( JÄCH et al. 2002). NIEDERÖSTERREICH: Wien env., Dr. C. Hampe leg., repeatedly ( REDTENBACHER 1849, 1858, 1874); Wien- XIV Bezirk, Kellerwiese bei Purkersdorf, 250 m a.s.l., R. Reithoffer leg., 1 spec., ex coll. M. Curti [destroyed, NLHW] ( PITTIONI 1943). STEIERMARK: Spielfeld, ca. 1900, Weber leg., 1 spec., SLJG ( HORION 1956). BELARUS: HRODNO: Grodnenskii reg.: Novoselki village env., Avgustovskii canal bank, 21.v.1990, O. R. Alexandrovich leg., 3 spec., ZIN, ZMUB ( ALEXANDROVICH 1995). BOSNIA HERZEGOVINA: ‘Bosnien’, 1 spec., DEI ( HORION 1956). BULGARIA: ‘Blg.’ ( ROUBAL 1936). BLAGOEVGRAD: SW of Gotse Delchev [as ‘Nevrokop’] in Alibotush Mts., J. Hlisnikovský leg. ( HLISNIKOVSKÝ 1958). BURGAS: Strandzha planina Mts., Malko Tarnovo env., J. Hlisnikovský leg. ( HLISNIKOVSKÝ 1958). CROATIA: ‘Kroatien’, 1 spec., NHMB in coll. G. Frey [‘Croat.’, V.K. revid.] ( HORION 1956); near Križevci; Podravina reg.: near Čukovec and Ludbreg ( SCHLOSSER KLEKOVSKI 1878: 571). CZECH REPUBLIC: MORAVIA: Paskov env., Olešná brook-Ostravice river valley [6275d], 17.iv.1869, on Quercus stump, E. Reitter leg., more spec. ( REITTER 1870); Olešná brook near Paskov [6275d], ~1871, Alnus stump, E. Reitter leg., 32 spec. ( REITTER 1908, 1911); Litovel env. [62–6368], ~1920, on Alnus stump in flooded brook, J. Slavíček leg. ( HUDEČEK 1940); ‘Morava’, formerly more abundant ( FLEISCHER 1930); Moravia (without exact data), 1990’s, more spec. ( JÄCH et al. 2002; see also ‘Unpublished records’); Hynkov env. [6369a], 24.vi.2006, 1 spec. ( NAKLÁDAL 2008); Litovel [6368b], 2.vii.2006, gravel bank – on a poplar log ( Populus sp. ) above the water level of the Morava river, 5 spec. ( NAKLÁDAL 2008). SILESIA: Ostravice (1.5 km NW), Ostravice river, ‘Peřeje’ [now ‘PP Koryto řeky Ostravice’ nature reserve, 49°33′04.50″N 18°22′59.30″E] [6476c], 400 m a.s.l., July-August 1953, J. Hlisnikovský leg., 2 JJ, 3 ♀♀ ( HLISNIKOVSKÝ 1958). FRANCE: ‘Gallia’ ( DEJEAN 1834, 1836). ALLIER: Saint-Pourçain-sur-Sioule ( DESBROCHERS DES LOGES 1900); Broût-Vernet, H. du Buysson leg., 3 spec., NHMB in coll. G. Frey [all V.K. revid., one with ‘8.x.07’] ( HORION 1956); Saint Pourçain, Sioule river, 20.viii.2003, 15 spec. ( BOUYON & VAN MEER 2004). ALLIER and PUY- DE- DÔME: Sioule and Allier rivers ( BUYSSON 1910). GERS: Adour river ( DELHERM DE LARCENNE 1877; sec. rec. DESBROCHERS DES LOGES 1900). GIRONDE: Belin-Beliet, La Leyre river, 20.ix.2003, 15 spec., 9.xi.2003, 6 spec. ( BOUYON & VAN MEER 2004). LOIRET: Saint-Benoît-sur Loire, 27.vii.2007, 2 spec. ( THÉRY 2007). LOT: Cabreret, Célé river, 27.xi.1991, 1 spec.; Vayrac, Dordogne river, 1 spec.; Martel, 1 spec., all D. Delpy leg. ( BOUYON & VAN MEER 2004). MAINE- ET- LOIRE: Sainte-Gemmes-sur-Loire, J. Gallois leg. ( ABOT 1928). RHÔNE: d’Izeron brook near Lyon, September 1843, more spec., C. Rey leg. (LAFERTÉ- SÉNECTÈRE 1849, MULSANT & REY 1866); Vallon d’Oullins, 15. v. and 3.vii.1845, E. Foudras leg. ( FALCOZ 1912). TARN: Agout river, Castres, Beaudecamy, 24.viii., on Alnus, H. Galibert leg., 9 spec. ( GALIBERT 1932). GERMANY: BAYERN: Donaustauf (E of Regensburg), Donau river bank under Walhalla monument, 4.vi.1949, H. Wichmann leg., 1 spec., ( HORION 1956). BRANDENBURG: Berlin, by flood, 2 spec. (LAFERTÉ- SÉNECTÈRE 1849); Oberspreewald [ca 60 km SE of Berlin], in flood debris ( QUEDENFELDT 1888). SACHSEN- ANHALT: Halle [type locality], on river at flight [?Saale river], 7.v.[?1818], 1 spec. ( GERMAR 1818, LAFERTÉ- SÉNECTÈRE 1849). WESTFALEN: Telgte env. [near Münster], 1.vi.1972, C. Rövekamp leg. ( RENNER 1991). GEORGIA: Borzhomi, June, O. Schneider leg., 1 spec. ( SCHNEIDER & LEDER 1878); Tbilisi ( JAKOBSON 1915). HUNGARY: ‘Ungarn’, Miller leg., 4 spec., ZSMC ( HORION 1956). SZABOLCS- SZATMÁR- BEREG: Tiszacsécse: Kis-mező, FU33, 11.v.2000, T. Kovács leg. ( KOVÁCS et al. 2000). VAS: Szentgotthárd: Zsida valley, from Alnus stump ( TRÄGER 1937, SZALÓKI 1999). ITALY: Upper Italy ( BERTOLINI 1874). EMILIA ROMAGNA: near Piacenza: at Po river ( BAUDI DI SELVE 1877); Po river near Piacenza, many spec., Meda leg. ( SEIDLITZ 1898); Piacenza, Meda leg., 2 spec., NHMB in coll. G. Frey [V.K. revid.] ( HORION 1956). LAZIO: ( PORTA 1934). LOMBARDIA: Pavia ( BAUDI DI SELVE 1877); Milano, 4 spec., ZSMC ( HORION 1956). PIEMONTE: at Po and Ticino rivers ( BAUDI DI SELVE 1889). TOSCANA: ( BERTOLINI 1874); Pisa ( BAUDI DI SELVE 1877). POLAND: ‘Silesia’ (by SCHILSKY (1888, 1908); and subsequently KUHNT (1912); Łomnicki (1913)); ‘Silesia’ sensu SCHILSKY (1888, 1908) is not Polish Silesia – unknown in Poland (BURAKOVSKI et al. 1987). ROMANIA: ARAD: Sebiş [as ‘Borossebes’], near Crişul Alb [= Fehér-Kőrös] river ( KUTHY 1897). CARAŞ- SEVERIN: Băile Herculane, near Cerna river, 1932, K. Dorn leg., more spec. ( HORION 1956). HUNEDOARA: Haţeg [as ‘Hátszeg’] ( BIELZ 1887; KUTHY 1897; PETRI 1912). SATU MARE: Tăşnad [as ‘Tasnád’] ( KUTHY 1897). EUROPEAN RUSSIA: NW Caucasus Mts.: ADYGEYA: Maikop reg., Novoprokhladnoe env., 1958 [no collector data], 1 spec. ( NIKITSKY et al. 2008). KRASNODAR REGION: Krasnaya Polyana env., Mzymta river valley, June 1996, on Alnus, B. M. Mamaev leg., larvae – more spec. ( MAMAEV 1976); Ubinskaya village, Ubinskoe lesnichestvo [forest district], viii.1971, at light, B. N. Nikitsky leg., 1 spec., ZMUM ( NIKITSKY et al. 2008). SLOVAKIA: Košice [as ‘Kaschau’] [72–7393] ( RÓZSAY 1868 – sec. rec. ROUBAL 1936); Medzilaborce [as ‘Mezőlaborcz’] [6797a] ( KUTHY 1897); Nová Sedlica, Zbojský brook valley [6901ac], 15.vi.1998, on Alnus, T. Jászay leg., 1 spec., SMBC ( JÁSZAY 2001). SLOVENIA: Lenart v Slovenskih Goricah, on Prunus padus L., 1 spec., J. N.Spitzy leg.[‘Die Käfer der Steiermark.’, ‘Auf Prunus Padus bei St. Leonhard 1 Ex. (Sp.)’] ( BRANCSIK 1871). SWITZERLAND: BASEL: Basel env., L. Imhoff leg., 1 spec. ( STIERLIN & GAUTARD 1871; STIERLIN 1890). BERN: Laupen ( HORION 1956). TICINO: Lugano, Stabile leg. ( STIERLIN & GAUTARD 1871, STIERLIN 1890). EUROPEAN TURKEY: İSTANBUL: Istranca [= Binkılıç], M. Jäch leg., 1 spec., NHMW ( JÄCH et al. 2002). UKRAINE: TRANSCARPATHIAN UKRAINE: Shopurka river [near Velikiy Buchkov], July 1924, on Fagus, C. Purkyně leg., more spec. ( PURKYNĚ 1926); Kosova rika river, 660 m a.s.l., August, on Alnus incana L., A. Pfeffer leg. ( ROUBAL 1934, 1936).

Unpublished records. BELARUS: HRODNO: Grodnenskii reg., Novoselki village env., Avgustovskii canal, 21.v.1990, O. R. Alexandrovich leg. (1 spec., ZIN, B. A. Korotayev and M. G. Volkovitsh pers. comm.). BOSNIA HERZEGOVINA: ‘Bosnia’ (1 spec., NMPC). BULGARIA: BLAGOEVGRAD: Pirin Mts. , August 1957, A. Pfeffer leg. (1 spec., NMPC). BURGAS: Strandzha planina Mts., Malko Tarnovo, 17.v.1939, J. Hlisnikowski leg. (1 spec., NMPC) [see also HLISNIKOVSKÝ 1958] ; Strandzha planina Mts. , July 1934, A. Pfeffer leg. (4 spec., NHMB in coll. J. Gottwald; 12 spec., NMPC). CZECH REPUBLIC: ‘ Moravia Reitter’ (1 spec., NMPC). MORAVIA- C: Chřiby hills, Divoky near Těšánky [6769d] (1 spec., MMBC in coll. E. Jagemann) ; ‘ CHKO Litovelské Pomoraví’ Protected Landscape Area, 10 km NW of Olomouc: Hynkov (ca 1 km E) [6369a], Morava river bank: January 1991, by sifting, V. Vyhnálek leg. (1 spec., ZKCM; 2 spec., VVCO) ; 23.iii.–3.iv.1991, by sifting, overwintering in bark crevices under moss at base of a poplar trunk ( Populus sp. ), M. Bednařík, R. Fornůsek, I. Martinů & V. Vyhnálek leg. (ca 80 spec., MBCO, VVCO, etc., M. Bednařík pers. comm.; also labelled as: ‘ Střeň’, ‘ Střeň / Litovel env.’ or ‘ Štěpánov, bř[eh]. Moravy [i.e. Morava river bank]’) ; 28.iii.1991, R. Fornůsek leg. (1 spec., HNHM, O. Merkl pers. comm.) ; 30.iii.1991, V. Vyhnálek leg. (1 spec., PZCW, P. Zabransky pers. comm.; 1 spec., ZKCM) ; 3.iv.1991, M. Bednařík leg. (1 spec., NMPC; 1 spec., PZCW, P. Zabransky pers. comm.) ; Hynkov (0.8 km NE) [6369a], ‘NPR Ramena řeky Moravy’ near ‘PR Kenický’ Nature Reserve , 49°40′24,7″N 17°10′52,4″E, 19.viii.2008, M. Horsák, M. Straka & A. Lacina leg. (8 spec., JSBC, MHBC, MMBC, MSOC, MTBC, P. Kment pers. comm.; 4 spec. NMPC) ; Litovel-Víska, ca. 20 m from Mlýnecký brook, on a white wall below a street light, 22.iv.2004, O. Nakládal leg. and det. (1 spec., ONCL). MORAVIA- S: near Bzenec-Přívoz [7069d]: ‘ Černé jezero’ forest, by sifting mosses at feet of Juglans nigra and Quercus sp. , 18.x.2003, M. Boukal leg. and det. (1 spec., MBCP) ; ditto, by sifting rotten wood, detritus and moss at foot of an oak ( Quercus ), 17.ii.2007, M. Boukal leg. and det.(1 spec., MBCP) ; near Bzenec-Přívoz [7069d], meander of Morava river: 25.xi.2003, J. Ch.Vávra leg. and det. (many spec., JVCO) ; 6.xii.2003, J. Ch. Vávra and T. Sitek leg., J. Ch.Vávra det. (many spec., JVCO, TSCO; 1 spec., ZKCM) ; Strážnice (3.5 km NW), ‘PP Osypané břehy’ nature reserve, meander of Morava river [7069d], 48°55′N 17°16′30″E, 170 m a.s.l., 5.ii.2009, V. Kubáň leg. (1 J, 2 ♀♀, NMPC) ; Břeclav (S env.): ‘ Pohansko’ [7267c], meadow near a path, sweeping, 1.viii.2007, R. Fornůsek leg. and det. (1 spec., not collected, M. Boukal, pers. comm.). SILESIA: Beskydy Mts. : Lysá hora massif, Kobylík brook [6476d], ca. 500 m a.s.l., ~1960, several spec., J. Hlisnikovský leg. (Petr Nohel, pers. comm.) ; Krásná (3 km S), near Řepčonka, Mt. Trávný (W slope), near the confluence of Česnekový potok brook and Mohelnice river [6476b], ca. 600 m a.s.l., 3.ix.1967, on Alnus, P. Nohel leg. and det. (1 dead spec., PNCP). FRANCE: ‘ Frankreich’ (2 spec., NHMB in coll. general) ; ‘ Gallia’ (1 spec., MMBC). ALLIER: Broût-Vernet , on Alnus, H. du Buysson leg.: 7.x.1906 (1 spec., NMPC), 3.x.1907 (4 spec., NMPC; 1 spec., SNMC, I. Rychlík pers. comm.), 18.x.1908 (2 spec., NMPC), without date (4 spec., HNHM, O. Merkl pers. comm.; 1 spec., NHMB in coll. J. Gottwald; 2 spec., NMPC) ; ‘ Allier’ (2 spec., NMPC) ; Montluçon , ii.1888, abbé Chanrion leg. (1 spec., CHCV, C. Holzschuh pers. comm.). ITALY: ‘ Italia, Dohrn’ (1 spec., ZIN, B. A. Korotayev and M. G. Volkovitsh pers. comm.) ; ‘ Italia, Rosenhauer’ (1 spec., NMPC). EUROPEAN RUSSIA: ‘ Caucas. ’ [or GEORGIA] (1 spec., ZIN, B. A. Korotayev and M. G. Volkovitsh pers. comm.). NW Caucasus Mts. : KABARDINO- BALKARIA: Urvanskij reg. [Urvan’, 10 km E of Nal’chik], Chernaya rechka [Black river], 22.iii.1986, under bark of poplar ( Populus ), Kravets leg. (1 spec., MKCY, M. Yu. Kalashian pers. comm.). KRASNODAR: Black Sea coast, 15 km SE of Novorossiisk, Kabardinka (1 spec., ZIN, B. A. Korotayev and M. G.Volkovitsh pers. comm.) ; 40 km NEE of Sochi, Krasnaya Polyana, A.Yu.Solodovnikov leg. (1 spec., ZIN, B. Korotyaev pers. comm.). SLOVAKIA: Kamenica nad Hronom, Hron river bank [8178c], in stump, 9.v.1978, L. Bocák leg. and det. (1 spec., LMBC) ; Giraltovce [6895c], 1933, V. Depta leg. (1 spec., MMBC; 1 spec., NMPC; 1 spec., SNMC, I. Rychlík pers. comm.) ; Remetské Hámre [7199ac], 21.vi.1965, Z. Tesař leg. (1 spec., SMOC, J. Roháček pers. comm.) ; Ulič, ca 50 m from the confluence of brooks Ulička and Zbojský [7000b], at dusk, sandy bank, flying, July 1995, T. Kopecký leg. and det. (1 spec., TKCH). SWITZERLAND: ‘ Swisse-Tessin, Lugano, Stierlin. ’ (1 spec., MHNG, coll. Switzerland, G. Cuccodoro pers. comm.). UKRAINE: TRANSCARPATHIAN UKRAINE: ‘ Carp. or.’ (2 spec., NMPC) ; ‘ Podk [arpatská]. Rus [= Transcarpathian Ukraine]’ (1 spec., NMPC) ; Šopurka river , [vii.] 1924, C. Purkyně leg. (1 spec., SNMC, I. Rychlík pers. comm.) ; Chernaya Tisa river valley east of Mt. Goverla [as ‘ Howerla’], C. Purkyně leg. (2 spec., NMPC) ; Kvasy [as ‘Kuzy’], July 1928, A. Pfeffer leg. (2 spec., NMPC) ; Kevelovo [as ‘ Apšinec’], C. Purkyně leg. (1 spec., MMBC; 1 spec., NHMB in coll. J. Gottwald; 1 spec., NMPC) .

Known distribution. Austria (Kärnten, Niederösterreich, Steiermark); Belarus (Hrodno) ; Bosnia Herzegovina; Bulgaria (Blagoevgrad, Burgas); Croatia (in the north) ; Czech Republic (Moravia, Silesia); France (Allier, Gers, Gironde, Isère, Loiret, Lot, Maine-et-Loire, Puy-de- Dôme, Rhône, Tarn) ; Germany (Bayern, Brandenburg, Sachsen-Anhalt, Westfalen) ; Georgia (Borzhomi, Tbilisi) ; Hungary (Szabolcs-Szatmár-Bereg, Vas) ; Italy (Emilia-Romagna, Lazio, Lombardia, Piemonte, Toscana);? Poland ; Romania (Arad, Caraş- Severin, Hunedoara, Satu Mare) ; European part of Russia (Adygeya, Kabardino-Balkaria, Krasnodar region) ; Slovakia (southern and eastern part); Slovenia (northeastern part); Switzerland (Basel, Bern, Ticino) ; European part of Turkey (İstanbul) ; Ukraine (Transcarpathian Ukraine).

Figures

Figs. 1–7. 1–4 – Agnathus secundus sp. nov.: 1 – holotype, habitus, 5.2 mm; 2 – antenna; 3 – aedeagus; 4 – pronotum. 5–7 – A. decoratus (Germar, 1818): 5 – habitus, specimen from Czech Republic (Moravia, Strážnice env.), 4.8 mm; 6 – antenna; 7 – aedeagus.

Fig. 8. Distribution of Agnathus decoratus (Germar, 1818).

Figs. 9–10. 9 – Czech Republic, Moravia, Litovelské Pomoraví Protected Landscape Area – locality with common occurrence of Agnathus decoratus (Germar, 1818). Photo by E. Svobodová.10 – China, northeastern Yunnan province, Hengduan Mts., Baima Massif, ~3900 m a.s.l. – locality of paratypes of A. secundus sp. nov. Photo by V. Kubáň.