Amplirhagada ponderi, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 231

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
DOI	https://doi.org/10.5281/zenodo.5239017
persistent identifier	https://treatment.plazi.org/id/62F53C96-5F51-4572-BE68-8E11E73F9986
taxon LSID	lsid:zoobank.org:act:62F53C96-5F51-4572-BE68-8E11E73F9986
treatment provided by	Felipe
scientific name	Amplirhagada ponderi
status	sp. nov.

Treatment

Amplirhagada ponderi View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia, northwestern Kimberley, Bonaparte Archipelago, Admiralty Gulf , Kingsmill Island , 14°09'24"S 125°46'16"E; KC-067 (leg. V. Kessner & A. Longbottom, 19 July 1988) GoogleMaps .

Type material. Holotype WAM S34605 View Materials (Pl.1.10) . Paratype WAM S41489 View Materials (1 preserved specimen), FMNH 219268 (1 preserved specimen).

Etymology. Named in honour of Winston F. Ponder, senior fellow of the Australian Museum, in recognition of his achievements in malacological research.

Shell (Pl. 1.10). Broadly conical, with low spire; thin (translucent) to solid. Periphery slightly angulate; upper and basal sectors of whorls rounded. Umbilicus open, narrowly winding, c. 40–60 percent concealed by columellar reflection. Background colour yellowish brown to horn; with diffuse, thin, light brown sub-sutural and mid-whorl bands being most conspicuous on last whorl; ventral colour whitish horn; outer lip colour same as shell; inner lip colour horn to translucent. Protoconch and teleoconch smooth except for growth lines. Angle of aperture 45°; outer lip rounded, sharp to moderately thick, expanded, slightly reflected; basal node of lip weak to moderately developed; palatal node absent. Parietal wall of inner lip inconspicuous.

Pallial morphology. Pallial cavity moderately deep, extending 3 ⁄ 4 whorl. Blackish mottled pigmentation on mantle. Kidney extending half of pallial cavity.

Genital morphology ( Figs. 18–19 View Figure 18 View Figure 19 ). Penis bent; much longer than anterior part of oviduct. Vas deferens forms complex coiling before entering penis. Penial retractor muscle stubby, much shorter than penis. Penial verge medium sized to long (1 ⁄ 8 – 1 ⁄ 3 of penial chamber), slender to spatulate with pointed tip. Penial wall pustules very small, arranged in rows over entire length of inner penial wall. Main stimulatory pilaster well-differentiated, narrow and quite long comprising apical to median portion of penial chamber; corrugated by ridges of undifferentiated pustules that support little hooks. Vas deferens entering penial sheath in upper third. Vagina relatively long, tubular, posteriorly slightly inflated. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct moderately thick with smooth longitudinal pilasters inside. Spermathecal head elongately inflated, connected with oviduct by connective tissue, entirely smooth inside; wall delicate. Length of free oviduct less than half of anterior part of oviduct, zig-zagfolded underneath entrance to spermoviduct. Posterior part of genitalia unknown.

Radular morphology ( Fig. 20A–C View Figure 20 ). Rectangular. Tooth formula C+16+3+18. Average number of rows of teeth 165 (n = 1) with 35.6 rows per mm (n = 1). Central teeth with sharply pointed, elongate to triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, elongate to triangular mesocones, not exceeding base of tooth; ectocones well developed, endocones vestigial. Marginal teeth with elongate to triangular mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.

Comparative remarks. The studied museum material was labelled as “ Amplirhagada sp. 73” by Solem. The shell of this species is similar to those of other species from the Montesquieu Islands, such as A. mckenziei , A. montesquieuana or A. puescheli (for comparison with the latter two see below), and is not readily differentiated from the former in shell characters alone. The inner penial wall of A. ponderi , however, is very characteristic. It differs from A. mckenziei most markedly by its much longer and narrow main pilaster, which extends almost to the anterior end of the penial chamber. The penis of A. ponderi is proportionally longer. For comparison with other species from the Montesquieu Islands see below. Phylogenetically, A. ponderi is more closely related to A. varia , A. solemiana , and A. indistincta . From all these it differs by possessing a well-developed, corrugated main pilaster.