Boueina, F. Toula, 1884

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 14-16

Boueina ? crassundia n. sp.

Figures 11.1-11.4, 11.6, 11.7 , 13.1, 13.2, 13.4, 13.5, 13.8, 13.10 , 14.1-14.4, 14.10 , 15.1, 15.2, 15.4, 15.7, 15.9 , 16.1, 16.2, 16.6, 16.8 , 18.6-18.11 , 20.13, 20.15

zoobank.org/ F99005B2-1230-43E7-992C-18910EB11B9A

v. 2013 Paraboueina; Brose, Lucas and Krainer, fig. 6.8.

v. 2013 b Boueina ? sp. 2; Vachard, Krainer and Lucas, p. 348 (no illustration).

Etymology. Latin crassundia: large intestine; because of the broad medullar siphons.

Holotype. NMMNH P-67506; Figure 11.1 ; sample MLY (2) 6-7a.

Paratypes. NMMNH P-67494, 67495, 67496, 67507, 67508; Figure 11.2-4, 11.6-7 .

Material. 200 specimens.

Locus Typicus. NMMNH locality 8892; San Andres Formation, McLeod Hills (New Mexico).

Stratum Typicum. Middle? Kungurian (late Leonardian).

Diagnosis. Boueina ? with broad siphons in both medullar and cortical zones.

Description. The thallus is cylindrical (L = 1.00- 1.75 mm; w = 0.45-0.75 mm). Medullar zone with medium-sized, tangled, longitudinal threads exhibiting long segments of tubes with a diameter of 0.05-0.09 mm. Cortical zone with relatively wide (0.03-0.04 mm), oblique, and one time branching, radial or oblique threads. The subdermal zone shows some small deltoid terminations (0.03-0.05 mm).

Comparison. Boueina ? crassundia n. sp. differs from B.? tubulata n. sp. by broader siphons, even where the other dimensions are the same. This character is not diagenetic because there are no transitional measurements.

Occurrence. Early?-middle? Kungurian (late Leonardian) of New Mexico (Yeso Group, Los Vallos Formation, Torres Member: samples MG 10-1, MG 10-10, MG(2) 12-6, MG(2) 12-11, MG(2) 12- 15; San Andres Formation, McLeod Hills (lower part); samples MLY 5-1, MLY 5-2, MLY 5-6, MLY(2) 5-6, MLY 5-7b, MLY(2) 5-9, MLY(2) 5-11, MLY(2) 5- 14, MLY(2) 5-16, MLY 5-18, MLY(2) 5-18, MLY(2) 5-19, MLY(2) 5-19d, MLY 5-20, MLY 5-26, MLY 5- 27, MLY 5-28, MLY(2) 5a-1, MLY(2) 5a-3, MLY(2) 5a-4, MLY(2) 5a-10, MLY(2) 5a-11, MLY(2) 5a-12, MLY(2) 5a-13, MLY(2) 5a-19, MLY(2) 5a-19a, MLY(2) 5a-21, MLY(2) 5a-23, MLY(2) 5a-25, MLY(2) 6-7, MLY(2) 6-7a, MLY 6-8, MLY(2) 6-8, MLY(2) 6-8a, MLY 6-10, MLY(2) 6-11, MLY 6-13, MLY 6-14, MLY(2) 6-15, MLY(2) 6-16, MLY(2) 6-17, MLY(2) 6-21, MLY(2) 6-27, MLY(2) 6-28, MLY(2) 6- 30, MLY 8-3)).

Figures

FIGURE 11. 1-4, 6-7. Boueina? crassundia n. sp. 1. NMMNH P-67506, holotype. Oblique section showing all the types of siphons. San Andres Fm. Sample MLY(2) 6-7a. 2. NMMNH P-67494, paratype. Axial section. San Andres Fm. Sample MLY(2) 5a-13. 3. NMMNH P-67495, paratype. Axial section with medullar siphons. San Andres Fm. Sample MLY(2) 5a-11. 4. NMMNH P-67507, paratype. Oblique section with cortical siphons. San Andres Fm. Sample MLY(2) 6-27. 6. NMMNH P-67507, paratype. Oblique sections with cortical siphons (already illustrated in Brose et al., 2013, figure 6.8). San Andres Fm. Sample MLY(2) 5-9. 7. NMMNH P-67496, paratype. Oblique section with cortical siphons. San Andres Fm. Sample MLY(2) 6-28. 5. Undetermined Chlorophyta. NMMNH P-67516, oblique sections with siphons or L2 and L1 laterals (bottom, left). San Andres Fm. Sample MLY(2) 6-10. 8-9. Gymnocodium? johnsonii n. sp. 8. NMMNH P-67511, paratype. Axial section with well preserved medullar zone. Yeso Group. Sample MG(2) 12-3. 9. NMMNH P-67512, paratype. Axial section more recrystallized than Figure 11.8. Specimen already illustrated in Lucas et al. (2013b, figure 8J). Yeso Group. Sample MG(2) 12-5. 10. Tubiphytes epimonellaeformis n. sp. NMMNH P-67518, a paratype encrusting a productoid spine (left); another paratype (right); and an isolated brachiopod spine (centre). San Andres Fm. Sample SAR 12-3.

FIGURE 13. 1, 4-5, 8, 10. Boueina? crassundia n. sp. 1. Tangential section (right) with Gymnocodium? johnsonii n. sp. (left) and crinoids (centre top and bottom). San Andres Fm. Sample MLY 5-26. 4. Oblique subaxial section showing the siphons of the medullar zone. San Andres Fm. Sample MLY 5-6. 5. Oblique section showing the siphons of the medullar zone. San Andres Fm. Sample MLY 5-18. 8. Axial section with cortical and medullar zones. San Andres Fm. Sample MLY 5-8. 10. Axial section with a well-developed medullar zone but a poorly preserved cortical zone. San Andres Fm. Sample MLY 5-20. 2, 9. Undetermined Chlorophyta. 2. Fragment of a longitudinal section and microfacies (bivalve, top left; Palaeonubecularia sp. (top right); Eotuberitina reitlingerae Miklukho-Maklay, 1958 (right, centre); crinoid (bottom left); Boueina? crassundia n. sp. (bottom right)). San Andres Fm. Sample MLY 5-2. 9. Detail of Figure 13.2; with Eotuberitina reitlingerae Miklukho-Maklay, 1958 (top right). San Andres Fm. Sample MLY 5-2a. 3, 6-7. Boueina? tubulata n. sp. 3. Subaxial section showing the ramified siphons of the medullar zone. San Andres Fm. Sample MLY 5-14. 6. Oblique subaxial section showing some ramifications of the siphons of the medullar zone. San Andres Fm. Sample MLY 5-7b. 7. Axial section with typical cortical and medullar siphons. San Andres Fm. Sample MLY 5-10.

FIGURE 14. 1-4, 10. Boueina? crassundia n. sp. 1. Axial section through dilated deltoid terminations. San Andres Fm. Sample MLY(2) 6-16. 2. Oblique section through the siphons. San Andres Fm. Sample MLY(2) 5a-25. 3. Tangential sections of the siphons. Yeso Group. Sample MG(2) 12-5. 4. Subaxial sections of cortical siphons. San Andres Fm. Sample MLY(2) 6-7. 10. Subaxial section showing medullar and cortical siphons. San Andres Fm. Sample MLY(2) 5a- 19a. 5, 8-9. Gymnocodium? bacillum n. sp. 5. Tangential section. San Andres Fm. Sample SAR(2) 8-9. 8. Axial section. San Andres Fm. Sample SAR(2) 8-20. 9. Axial section. San Andres Fm. Sample SAR(2) 8-14. 6-7, 11. Gymnocodium? johnsonii n. sp. 6. Axial section. Yeso Group. Sample MG(2) 12-13. 7. Subaxial section without calcified medullar zone; the internal sediment contains an Eotuberitina reitlingerae Miklukho-Maklay, 1958. San Andres Fm. Sample SAR(2) 8-17. 11. Axial section completely recrystallized. San Andres Fm. Sample MLY(2) 6-14.

FIGURE 15. 1-2, 4, 7, 9. Boueina? crassundia n. sp. 1. Oblique subaxial section. San Andres Fm. Sample MLY 5-1. 2. Oblique section. San Andres Fm. Sample MLY 5-27. 4. Axial section. San Andres Fm. Sample MLY 5-28. 7. Atypical axial section with excessively dilated siphons of medullar zones. San Andres Fm. Sample MLY 6-8. 9. Two oblique sections with typical dilated siphons of medullar zones. San Andres Fm. Sample MLY 6-10. 3, 5-8, 10-11. Gymnocodium? johnsonii n. sp. 3. Tangential section. San Andres Fm. Sample MLY 6-6. 5. Transverse section. San Andres Fm. Sample MLY 6-11. 6. Subaxial section. San Andres Fm. Sample MLY 6-7. 7. Oblique section. San Andres Fm. Sample MLY 6-8. 8. Oblique subaxial section. San Andres Fm. Sample MLY 6-15. 10. Axial section. San Andres Fm. Sample MLY 6-12. 11. Axial section. San Andres Fm. Sample MLY 6-25.

FIGURE 16. 1-2, 6, 8. Boueina? crassundia n. sp. 1. Oblique section. San Andres Fm. Sample MLY 6-13. 2. Subaxial section. San Andres Fm. Sample MLY 6-14. 6. Tangential section. San Andres Fm. Sample MLY 6-30. 8. Atypical oblique section. San Andres Fm. Sample MLY 8-3. 3-5. Undetermined Chlorophyta. 3. Two sections (centre) with Gymnocodium? johnsonii n. sp. (centre bottom and top) and Boueina? tubulata n. sp. (top right). San Andres Fm. Sample MLY 6-20. 4. At least six fragments. San Andres Fm. Sample MLY 6-21. 5. Tangential section. San Andres Fm. Sample MLY 8-5. 7, 9. Gymnocodium? johnsonii n. gen. n. sp. 7. Various sections associated with Globivalulina parapiciformis n. sp. (bottom, left) and G. novamexicana n. sp. (centre). San Andres Fm. Sample MLY 6-32. 9. Subaxial section. San Andres Fm. Sample MLY 6-27.

FIGURE 18. 1-5. Boueina? tubulata n. sp. 1. Axial section. San Andres Fm. Sample MLY(2) 5a-26. 2. Axial section. San Andres Fm. Sample MLY(2) 5a-9. 3. Subaxial section. San Andres Fm. Sample MLY(2) 5a-2. 4. Oblique section. San Andres Fm. Sample MLY 5-18. 5. Subaxial section. San Andres Fm. Sample MLY(2) 5-19. 6-11. Boueina? crassundia n. sp. 6. Axial section. San Andres Fm. Sample MLY(2) 5-9. 7. Axial section. San Andres Fm. Sample MLY(2) 5-19d. 8. Oblique section. San Andres Fm. Sample MLY(2) 5-11. 9. Oblique subaxial section. San Andres Fm. Sample MLY(2) 5-16. 10. Longitudinal section. San Andres Fm. Sample MLY(2) 5-14. 11. Oblique section (left). San Andres Fm. Sample MLY(2) 5a-12. 11. Gymnocodium? johnsonii n. sp. An axial section through the cortical zone (right). San Andres Fm. Sample MLY(2) 5a-12. 12. Tubiphytes epimonellaeformis n. sp. Specimen similar to Latitubiphytes Vachard, Krainer and Lucas, 2012 exhibiting a neosparitization of Miliolata, especially Hemigordiopsis, Baisalina or Pseudobaisalina (see Gargouri and Vachard, 1988; Vdovenko et al., 1993). San Andres Fm. Sample SAR(2) 12-15.

FIGURE 20. 1-4. Globivalvulina novamexicana n. sp. 1. Subaxial section. Yeso Group. Sample MG 10-18. 2. Subaxial section. Yeso Group. Sample MG 10-19a. 3. Oblique section. Yeso Group. Sample MG 12-1. 4. Recrystallized subaxial section. Yeso Group. Sample MG 12-4. 5-8, 10-11. Hemigordiellina aff. simplex (Harlton, 1928). 5. Random section. Yeso Group. Sample MG 10-23. 6. Random section. Yeso Group. Sample MG 12-7. 7. Random section. Yeso Group. Sample MG 12-11. 8. Random section. Yeso Group. Sample MG 12-12. 10. Random section. Yeso Group. Sample MG 12-13. 11. Random section. Yeso Group. Sample MG 12-14. 9. Eotuberitina reitlingerae Miklukho-Maklay, 1958. Axial section with basal disk. Yeso Group. Sample MG 12-12a. 12. Geinitzina indepressa Cherdyntsev, 1914. Subaxial section. Yeso Group. Sample MG 12-13a. 13, 15. Boueina? crassundia n. gen. n. sp. 13. Oblique section. Yeso Group. Sample MG 10-1. 15. Tangential section. Yeso Group. Sample MG 10-10. 14. Palaeonubecularia sp. Subaxial section. Yeso Group. Sample MG 10-5. 16. Gymnocodium? bacillum n. sp. Axial section. Yeso Group. Sample MG 10-11. 17, 19. Gymnocodium? johnsonii n. sp. 17. Two transverse sections. Yeso Group. Sample MG 10-13. 19. Subaxial section. Yeso Group. Sample MG 12-4a. 18. Undetermined Chlorophyta. Subtransverse section through the laterals. Yeso Group. Sample MG 10-19. Scale bars 1-12 equal 0.10 mm; 13-19 equal 0.50 mm.