Globivalvulina novamexicana, Vachard, Krainer & Lucas, 2015

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 32-34

Globivalvulina novamexicana n. sp.

Figures 20.1-20.4 , 27.2, 27.5-27.8, 27.12-27.19 , 28.1-28.3, 28.7 , 29.6, 29.9 , 30.7 , 31.1 , 32.4, 32.5

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1972 Globivalvulina bulloides (Brady) ; Toomey, p. 295, pl. 2, figs. 19, 20, 21, 22, 23.

v. 1993 Globivalvulina ex gr. bulloides (Brady) ; Vachard, Oviedo, Flores de Dios, Malpica, Brunner, Guerrero and Buitrón, pl. 6, fig. 8; pl. 8, figs. 7, 8, 9.

2000 Globivalvulina ex gr. bulloides (Brady) ; Groves, pl. 2, figs. 25, 26, 27, 28, 29, 30, 31, 32.

v. 2003 Globivalvulina spp. (partim); Krainer, Vachard and Lucas, pl. 8, figs. 1, 11, 15, 16?

v. 2009 Globivalvulina ex gr. mosquensis Reitlinger ; Krainer, Lucas and Vachard, pl. 4, figs. 6, 7.

v. 2013b Globivalvulina cf. G. mosquensis Reitlinger ; Lucas, Krainer and Voigt, fig. 8G.

v. 2013 b Globivalvulina cf. donbassica Potievskaya ; Vachard, Krainer and Lucas, p. 348 (no illustration).

Etymology. Latinized name indicating a New Mex-

ican origin.

Holotype. NMMNH P-67561; Figure 25.1 ; sample

MLY 6-17.

Paratypes. NMMNH P-67562, 67563; Figures

21.15, 22.2.

Material. 70 specimens.

Locus Typicus. NMMNH locality 8899; San Andres Formation, McLeod Hills in the southern Caballo Mountains (New Mexico).

Stratum Typicum. Middle? Kungurian (late Leonardian).

Diagnosis. Test medium-sized, often ovoid, with septa curved backward, funnel wide and shallow, and thin microgranular wall.

Description. Test subspherical to nautiloid, moderate in size (D = (rarely 0.23)- 0.28-0.55 mm; w = (rarely 0.24-0.25)- 0.30-0.62 mm; w/D = 0.89-1.20). Slightly trochospiral axis of biseriality. Dorsal side convex. Sutures absent or very weak. Lateral sides strongly convex to a few compressed on the flanks. Proloculus medium to large (0.05-0.06 mm), spherical. Whorl: 1-1.5. Peripheral margin round. Lateral sides of spire inflated. Umbilicus absent. Chambers: 6-7 (rarely 8) pairs, subtriangular, and with h = (rarely 0.07-0.09) 0.11-0.20 mm. Septa almost perpendicular to the wall. Ventral side flattened to slightly depressed. Aperture in basal slit. Oral valvula weakly developed to absent. Wall bilayered, with an inner layer relatively developed, especially on the septa; s = 0.02-0.04 mm with a maximum of 0.01 mm for the inner layer.

Comparisons. This taxon is similar to G. donbassica Potievskaya, 1962 but differs by slightly larger dimensions, chambers less rounded in transverse section, and a basal part more planar in axial section. This species also resembles Globivalvulina biserialis Cushman and Waters 1928b sensu St Jean, 1957 (p. 36-37, plate 3, figures 3a-c, 4-5) but differs in the shape of the axial section. It also resembles G. mosquensis Reitlinger, 1950 but differs in its larger dimensions (D, w, h) for the same number of chambers and in the great difference of age ( G. mosquensis is initially a Moscovian species). The morphology is relatively similar to that of Globivalvulina bulloides ; the main difference is the wall microstructure (a possible but controversial generic character; see Brenckle, 2005; Vachard et al., 2006).

Occurrence. Kungurian (late Leonardian) of New Mexico (Yeso Group, Los Vallos Formation, Torres Member, Massacre Gap in the southern Fra Cristobal Mountains: samples MG(2) 10-1, MG 10-3, MG 10-6, MG 10-6a, MG 10-7, MG 10-8, MG 10- 14, MG 10-17, MG 10-17a, MG 10-18, MG 10-19a, MG 12-1, MG 12-4; San Andres: samples MLY 3-2, MLY 3-6, MLY 3-7a, MLY 3-8, MLY 3-10, MLY 4-1, MLY 4-2, MLY 4-4, MLY 4-6, MLY 4-9, MLY 4-13, MLY 4-17, MLY(2) 5-2, MLY(2) 5-2a, MLY 5-7c, MLY 5-20a, MLY(2) 5-21, MLY 5-22, MLY(2) 5-23,

MLY 6-1, MLY(2) 6-1, MLY(2) 6-2, MLY 6-16, MLY 6-17, MLY(2) 6-23, MLY(2) 6-25, MLY 6-29a, MLY 8-8, SAR 7-3, SAR(2) 8-4, SAR 8-5, SAR 8a-5a, SAR 8a-9, SAR 8a-12, SAR 8a-20, SAR 8a-21, SAR 8a-24, SAR 12-2, SAR(2) 12-4, SAR(2) 13-3, SAR(2) 13-4, SAR(2) 13-5, SAR(2) 13-8, SAR(2) 13-11, SAR 13-12, SAR(2) 13a-3/3a, SAR 20-8).

Figures

FIGURE 20. 1-4. Globivalvulina novamexicana n. sp. 1. Subaxial section. Yeso Group. Sample MG 10-18. 2. Subaxial section. Yeso Group. Sample MG 10-19a. 3. Oblique section. Yeso Group. Sample MG 12-1. 4. Recrystallized subaxial section. Yeso Group. Sample MG 12-4. 5-8, 10-11. Hemigordiellina aff. simplex (Harlton, 1928). 5. Random section. Yeso Group. Sample MG 10-23. 6. Random section. Yeso Group. Sample MG 12-7. 7. Random section. Yeso Group. Sample MG 12-11. 8. Random section. Yeso Group. Sample MG 12-12. 10. Random section. Yeso Group. Sample MG 12-13. 11. Random section. Yeso Group. Sample MG 12-14. 9. Eotuberitina reitlingerae Miklukho-Maklay, 1958. Axial section with basal disk. Yeso Group. Sample MG 12-12a. 12. Geinitzina indepressa Cherdyntsev, 1914. Subaxial section. Yeso Group. Sample MG 12-13a. 13, 15. Boueina? crassundia n. gen. n. sp. 13. Oblique section. Yeso Group. Sample MG 10-1. 15. Tangential section. Yeso Group. Sample MG 10-10. 14. Palaeonubecularia sp. Subaxial section. Yeso Group. Sample MG 10-5. 16. Gymnocodium? bacillum n. sp. Axial section. Yeso Group. Sample MG 10-11. 17, 19. Gymnocodium? johnsonii n. sp. 17. Two transverse sections. Yeso Group. Sample MG 10-13. 19. Subaxial section. Yeso Group. Sample MG 12-4a. 18. Undetermined Chlorophyta. Subtransverse section through the laterals. Yeso Group. Sample MG 10-19. Scale bars 1-12 equal 0.10 mm; 13-19 equal 0.50 mm.

FIGURE 27. 1, 4, 9, 20. Globivalvulina retroseptata n. sp. 1. Subaxial section. San Andres Fm. Sample MLY 8-2a. 4. Subtransverse section. San Andres Fm. Sample MLY 5-23. 9. Subaxial section. San Andres Fm. Sample MLY 5-3. 20. Transverse section. Yeso Group. Sample MG 10-9. 2, 5-8, 12-19. Globivalvulina novamexicana n. sp. 2. Subtransverse section. San Andres Fm. Sample MLY 4-13. 5. Oblique subaxial section. San Andres Fm. Sample MLY 4- 9. 6. Oblique section. San Andres Fm. Sample MLY4-1. 7. Axial section. San Andres Fm. Sample MLY 4-2. 8. Subaxial section. San Andres Fm. Sample MLY 4-6. 12. Axial section. San Andres Fm. Sample MLY 3-8. 13. Subtransverse section. Yeso Group. Sample MG 10-8 (already illustrated in Lucas et al., 2013b, figure 8G). 14. Subaxial section. Yeso Group. Sample MG 10-14. 15. Oblique subtransverse section. Yeso Group. Sample MG 10-17a. 16. Subtransverse section. Yeso Group. Sample MG 10-17. 17. Subaxial section. Yeso Group. Sample MG 10-7. 18. Subtransverse section. Yeso Group. Sample MG10 6. 19. Subaxial section. Yeso Group. Sample MG 10-6a. 3, 10- 11. Globivalvulina parapiciformis n. sp. 3. Axial section. San Andres Fm. Sample MLY 6-16. 10. Axial section. San Andres. Sample MLY 5-20a. 11. Axial section. San Andres Fm. Sample MLY 5-7c. Scale bars equal 0.10 mm.

FIGURE 28. 1-3, 7. Globivalvulina retroseptata n. sp. 1. Transverse section. San Andres Fm. Sample SAR 13-5. 2. Transverse section. San Andres Fm. Sample SAR 13-5b. 3. Transverse section. San Andres Fm. Sample SAR 13- 5c. 7. Transverse section. San Andres Fm. Sample SAR 8-5. 4-6, 8-12, 14-16. Globivalvulina praegraeca n. sp. 4. Transverse section. San Andres Fm. Sample SAR 13-7. 5. Transverse section. San Andres Fm. Sample SAR 8-2. 6. Subtransverse section. San Andres Fm. Sample SAR 8-3. 8. Subaxial section. San Andres Fm. Sample SAR 8-7a. 9. Subaxial section. San Andres Fm. Sample SAR 8a-2. 10. Subaxial section. San Andres Fm. Sample SAR 8a-5. 11. Axial section. San Andres Fm. Sample SAR 8a-5a. 12. Transverse section. San Andres Fm. Sample SAR 8a-18. 14. Subtransverse section. San Andres Fm. Sample SAR 8a-9. 15. Oblique subtransverse section. San Andres Fm. Sample SAR 8a-10. 16. Subtransverse section. San Andres Fm. Sample SAR 8a-12. 13. Globivalvulina parapiciformis n. sp. Transverse section. San Andres Fm. Sample SAR 8a-8a. Scale bars equal 0.10 mm.

FIGURE 29. 1-4, 7. Globivalvulina retroseptata n. sp. 1. Transverse section. San Andres Fm. Sample SAR 19-2. 2. Transverse section. San Andres Fm. Sample SAR 19-3. 3. Transverse section. San Andres Fm. Sample SAR 19-4. 4. Subtransverse section with developed oral valvula. San Andres Fm. Sample SAR 20-1. 7. Teratogenic specimen. San Andres Fm. Sample SAR 3-1. 5, 8, 10-13. Globivalvulina praegraeca n. sp. 5. Transverse section. San Andres Fm. Sample SAR 20-3. 8. Axial section. San Andres Fm. Sample SAR 3-4. 10. Subaxial section. San Andres Fm. Sample SAR 8-1. 11-12. Two transverse sections. San Andres Fm. Sample SAR 7-4. 13. Subaxial section. San Andres Fm. Sample SAR 7-5. 6, 9. Globivalvulina novamexicana n. sp. 6. Subtransverse section. San Andres Fm. Sample SAR 20-4. 9. Axial section. San Andres Fm. Sample SAR 7-3. Scale bars equal 0.10 mm.

FIGURE 30. 1, 4-6, 8-10, 12, 14-15. Globivalvulina praegraeca n. sp. 1. Subtransverse section. San Andres Fm. Sample SAR 8a-13a. 4. Transverse section. San Andres Fm. Sample SAR 9-1. 6. Transverse section. San Andres Fm. Sample SAR 13-10. 8. Subaxial section. San Andres Fm. Sample SAR(2) 13-2. 9. Subtransverse section. San Andres Fm. Sample SAR(2) 13-6. 10. Transverse section. San Andres Fm. SAR 13-9. 12. Axial section showing the proloculus and the first whorl. San Andres Fm. Sample SAR(2) 8-21. 14. Transverse section. San Andres Fm. Sample MLY(2) 4-9. 15. Transverse section. San Andres Fm. Sample SAR 8-18. 2-3, 11, 13, 16. Globivalvulina retroseptata n. sp. 2. Subtransverse section. San Andres Fm. Sample SAR 8a-19. 3. Transverse section. San Andres Fm. Sample SAR 8a-24. 11. Transverse section. San Andres Fm. Sample SAR 13a-7. 13. Transverse section. San Andres Fm. Sample MLY(2) 6-13 (already illustrated by Brose et al., 2013, figure 6.7). 16. Axial section. San Andres Fm. Sample SAR(2) 12-6. 5. Globivalvulina parapiciformis n. sp. Subtransverse section. San Andres Fm. Sample SAR 13-9. 7. Globivalvulina novamexicana n. sp. Transverse section. San Andres Fm. Sample SAR 13-12. Scale bars equal 0.10 mm.

FIGURE 31. 1. Globivalvulina novamexicana n. sp. Axial section. San Andres Fm. Sample SAR 8-4. 2-8. Globivalvulina praegraeca n. sp. 2. Subtransverse section. San Andres Fm. Sample SAR 8-6. 3. Axial section. San Andres Fm. Sample SAR 8-7. 4. Subaxial section. San Andres Fm. Sample SAR 8a-10a. 5. Paratype. Oblique section. San Andres Fm. Sample SAR 8a-20. 6. Subaxial section. San Andres Fm. Sample SAR 8a-21. 7. Subaxial section. San Andres Fm. Sample SAR 12-2. 8. Subaxial section. San Andres Fm. Sample SAR 13-1. 9-12. Globivalvulina retroseptata n. sp. 9. Oblique subaxial section. San Andres Fm. Sample SAR 13a-7. 10. Subaxial section. San Andres Fm. Sample SAR 13a-1. 11. Oblique subaxial section. San Andres Fm. Sample SAR 13a-8. 12. Axial section. San Andres Fm. Sample SAR 13a-9. Scale bars equal 0.10 mm.

FIGURE 32. 1-3, 6. Globivalvulina retroseptata n. sp. 1. Subaxial section. San Andres Fm. Sample SAR 13a-10. 2. Oblique subaxial section. San Andres Fm. Sample SAR 13a-11. 3. Subaxial section. San Andres Fm. Sample SAR 20-6. 6. Axial section. San Andres Fm. Sample SAR (A-1§). 4-5. Globivalvulina novamexicana n. sp. 4. Oblique subaxial section. San Andres Fm. Sample SAR 20-8. 5. Subaxial section. San Andres Fm. Sample SAR 20-9. 7-10. Hemigordius? sp. Four sections. San Andres Fm Sample SAR 1. Scale bars equal 0.10 mm.

FIGURE 25. 1-2. Globivalvulina novamexicana n. sp. 1. NMMNH P-67561, holotype. Subaxial section. San Andres Fm. Sample MLY 6-17. 2. NMMNH P-67563, paratype. Subaxial section. San Andres Fm. Sample MLY 6-29a. 3. Globivalvulina parapiciformis n. sp. NMMNH P-67552, two paratypes. Two oblique sections. San Andres Fm. Sample MLY 6-11a. 4. Nestellorella? sp. Subaxial section. San Andres Fm. Sample MLY 1-1. 5. Protonodosaria? sp. Subaxial section. San Andres Fm. Sample MLY 3-5. 6-7, 10, 17-19. Nodosinelloides longa (Lipina, 1949). 6. Axial section. San Andres Fm. Sample SAR 8a-4a. 7. Subaxial section. San Andres Fm. Sample SAR 8a-8. 10. Axial section. San Andres Fm. Sample SAR 8a-17 (already illustrated by Brose et al., 2013, figure 6.5). 17. Axial section. San Andres Fm. Sample SAR 11a-6. 18. Axial section. San Andres Fm. Sample SAR 11a-9. 19. Axial section. San Andres Fm. Sample SAR 12-4. 8-9, 12-14. Geinitzina indepressa Cherdyntsev, 1914. 8. Axial section. San Andres Fm. Sample SAR 8a-11. 9. Axial section. San Andres Fm. Sample SAR 8a-14. 12. Subaxial section. San Andres Fm. Sample SAR 8a-7. 13. Subaxial section. San Andres Fm. Sample SAR 11-13. 14. Oblique subaxial section. San Andres Fm. Sample SAR 11a-10. 11, 15, 20?, 21. Nodosinelloides netjaschewi (Cherdyntsev, 1914). 11. Axial section. San Andres Fm. Sample SAR 11-17. 15. Axial section. San Andres Fm. Sample SAR 11-19/19a. 20. Axial section. San Andres Fm. Sample SAR 20.2. 21. Axial section. San Andres Fm. Sample SAR 13a-2. 16. Nodosariata indet. cf. Dentalina cf. ninae Grozdilova sensu Chuvashov et al., 1990. Axial section. San Andres Fm. Sample SAR 11a-4. Scale bars equal 0.10 mm.