Scaphisoma sumpichi, Löbl, 2018

Löbl, Ivan, 2018, A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae), Acta Entomologica Musei Nationalis Pragae 58 (1), pp. 151-165 : 158-159

Scaphisoma sumpichi sp. nov.

( Figs 31–34 , 52 )

Type locality. Indonesia, South Sulawesi, Gowa District, 6 km E of Malino, Gunung Bawakaraeng [Mt.] Area, near Lembanna Base camp, ca. 05°15.4 ′ S, 119°54.5 ′ E.

Type material. HOLOTYPE: ( NMPC), INDONESIA, S SULAWESI: Gowa Distr. / 6 km E of Malino, Gn.Bawakaraeng Area / border of gardens and mixed forest / (dominant Pinus) nr Lembanna Base camp / 05°15.4 ′ S, 119°54.5 ′ E, 1520 m / J.Hájek & J.Šumpich leg., 11-13.ii.2015. PARATYPES: 3 3 ♀♀ ( NMPC, MHNG), with the same data as the holotype.

Description. Length 1.76–1.95 mm, width 1.14–1.25 mm. Head, mouthparts, basal antennomeres, hypomera, legs and apical abdominal segments ochreous; antennomeres V to XI brown; pronotum reddish-brown to black, in middle sometimes darker than on sides; elytra dark brown to black along basal margin, on adsutural areas, along apical margins, and each with large dark brown to black discal spot situated between basal fourth and apical third of disc, joint to dark basal area along sutural stria; area surrounding discal spot ochreous ( Fig. 52 ); ventral side of mesothorax and metathorax dark brown to blackish, abdominal ventrites 1 to 5 reddish-brown. Antennae long, length/width ratios of antennomeres as: III 15/9: IV 35/8: V 44/8: VI 43/10: VII 50/15: VIII 42/10: IX 47/14: X 45/14: XI 60/15. Pronotum and elytra not microsculptured. Pronotum strongly narrowed anteriad, with regularly arcuate lateral margins, lateral margins carinae concealed in dorsal view; punctation on disc very fine, dense, hardly visible at magnification 30×, consisting of not well delimited punctures, puncture intervals much larger than puncture diameters, punctation along lateral carinae indistinct. Tip of scutellum exposed. Elytra moderately narrowed apically, lateral margins almost regularly curved, lateral margin carinae hardly visible in dorsal view, apical margin weakly rounded, inner apical angles situated in same level as outer angles, apical serration inconspicuous, sutural margin not raised, adsutural areas flat, narrow, each with single puncture row. Sutural striae parallel, curved near pronotal lobe, not extended laterad of pronotal lobe. Punctures along lateral margins very fine. Elytral disc with punctures fine, almost evenly large, not well delimited, larger than those on pronotum, puncture intervals mostly about two to four times as large as puncture diameters. Hind wings fully developed. Hypomera smooth, not microsculptured. Mesepimera as long as intervals between them and mesocoxae and four times as long as wide. Metaventrite with strigulate microsculpture; centre of metaventrite convex, with two shallow apical impressions, lacking mesal stria, punctation all over sparse and very fine; apical intercoxal margin slightly concave. Lateral parts of metaventrite sparsely and very finely punctate, antecoxal puncture rows absent. Submesocoxal lines convex, distinctly punctate; submesocoxal areas 0.03 mm, about as long as seventh of shortest intervals between them and metacoxae. Metanepisterna flat, hardly narrowed anteriad, with slightly rounded angles. Tibiae straight. Abdomen with strigulate microsculpture and extremely fine punctation; ventrite I with submetacoxal lines convex, distinctly punctate; submetacoxal areas each 0.05 mm, as long as fourth of shortest intervals between them and apical margin of ventrite.

Male characters. Protarsomeres I–III strongly widened, narrower than apices of protibiae; mesotarsomeres I and II distinctly widened. Abdominal ventrite VI with angular apical margin. Tergite IX lacking macrosetae, stalk evenly narrow, oblique laterally ( Fig. 33 ). Aedeagus ( Figs 31, 32 ) 0.81–0.95 mm long, symmetrical. Median lobe with apical process much shorter than basal bulb, strongly inflexed, tapering and with ventral side slightly curved in lateral view, hardly narrowed and with obtuse tip in dorsal view; dorsal valve sclerotized. Articular process indistinct. Parameres wide at bases, narrowed toward apical thirds, apical section widened in dorsal view, slightly inflexed and narrowed in lateral view, inner margin membranous. Internal sac tubular, sinuate, bearing robust rod and spine-like structures.

Female characters. Protarsomeres narrow. Gonocoxite bent, evenly narrow ( Fig. 34 ).

Differential diagnosis. This species also possesses aedeagal characters suggesting relationships with S. bugi and its allied species (see differential diagnoses under S. ogawai sp. nov., S. flavolineatum sp. nov. and S. pellax sp. nov.). It may be readily distinguished from these species by the colour pattern of the elytra, and from S. ogawai sp. nov. and S. pellax sp. nov. by the metaventrite lacking a median stria. This new species is unique in having the internal sac with a single, robust, sinuate rod surrounded by dense spine-like structures.

Etymology. The species is named after one of its collectors, Jan Šumpich, lepidopterist at the National Museum, Prague. The name is a noun in the genitive case.

Distribution. Indonesia, South Sulawesi.

Figures

Figs 20–34. 20–24 – Scaphisoma ogawai sp. nov.: 20 – gonocoxite; 21 – male tergite 9; 22, 23 – aedeagus in dorsal and lateral views; 24 – paramere in ventral view. 25–27 – S. flavolineatum sp. nov.: 25 – male tergite 9; 26, 27 – aedeagus in dorsal and lateral views. 28–30 – S. pellax sp. nov.: 28, 29 – aedeagus in dorsal and lateral views; 30 – male tergite 9. 31–34 – S. sumpichi sp. nov.: 31, 32 – aedeagus in dorsal and lateral views; 33 – male tergite 9; 34 – gonocoxite. Scale bars: 0.1 mm (Figs 20, 28–29, 34); 0.2 mm (Figs 21–27, 30–32); 0.3 mm (Fig. 33).

Figs 50–53. Habitus of Scaphisoma. 50, 51 – S. ancora sp. nov. (50 – male, 51 – female); 52 – S. sumpichi sp. nov., male; 53 – S. versicoloreum sp. nov., male.