Bachmarima, Constant & Pham, 2025

Constant, Jérôme & Pham, Hong Thai, 2025, Issid planthoppers from Bach Ma and Phong Dien in Central Vietnam. (III) Tribe Sarimini (Hemiptera: Fulgoromorpha: Issidae), European Journal of Taxonomy 1025, pp. 1-109 : 6-9

Genus Bachmarima gen. nov.

urn:lsid:zoobank.org:act:

Type species

Bachmarima expansa gen. et sp. nov. by present designation.

Diagnosis

The genus Bachmarima gen. nov. can be differentiated from the other genera of Sarimini by the following combination of characters: (1) the vertex more than twice as wide as long in dorsal view; (2) the frons with complete median carina and obsolete peridiscal carinae, weakly distinct only in dorsal portion of frons; (3) the tegmina elongate, about 2.2 × as long as wide, with distinct lateral hump slightly before basal ⅓ hiding costal margin in dorsal aspect, and with rather narrow but distinct epipleuron; (4) the vein ScP of the tegmen rather short, curved and not extending beyond midlength of tegmen; (5) the first fork of MP and the first fork of CuA at about the same level, slightly beyond halflength of tegmen; (6) the anal tube elongate, dorsoventrally flattened, and sublanceolate, rather narrow in dorsal view; (7) the massive gonostyli, with capitulum elongate, strongly projecting dorsad and with poorly distinct neck, and with anterodorsal margin distinctly rounded; (8) the aedeagus with a single pair of elongate, subapical, lateroventral processes, and with the dorsal lobe of the periandrium expanded lateroventrally in basal portion.

Differential diagnosis

The most similar genera are Dactylissus Gnezdilov & Bourgoin, 2014 , Neosarima Yang, 1994 , Sarimissus Wang, Zhang & Bourgoin, 2019 , Sarimites Meng, Qin & Wang, 2020 , Sinesarima Yang, 1994 and Yangissus Chen, Zhang & Chang, 2014 . However, Bachmarima gen. nov. can be separated from all of these genera by showing a complete median carina on the frons, extending from the dorsal margin to the frontoclypeal suture, while it does not reach the latter in the other genera.

Additionally, Bachmarima gen. nov. can be separated (1) from Dactylissus (see illustrations in Gnezdilov et al. 2014a: figs 18–21) by the more elongate anal tube, about 2.8 × as long as wide in dorsal view (2.5 × in Dactylissus ) and the strongly elongate capitulum of the gonostyli (rather short in Dactylissus ); (2) from Neosarima (see illustrations in Chan & Yang 1994: figs 43–44) by the less elongate tegmina, about 2.2 × as long as wide (2.5 × in Neosarima ), the more elongate anal tube, about 2.8 × as long as wide in dorsal view (2.5 × in Neosarima ) and the strongly elongate capitulum of the gonostyli (rather short in Neosarima ); (3) from Sarimissus (see illustrations in Wang et al. 2019: figs 1–11) by the frons nearly 1.2 × as wide as long in midline (1.0 time in Sarimissus ), the vein ScP+RA of tegmen rather short, curved towards RP (longer and subparallel to RP in Sarimissus ) and the capitulum of the gonostyli in lateral view tapering towards pointed apex (parallel-sided with rounded apex in Sarimissus ); (4) from Sarimites (see illustrations in Zhang et al. 2020: figs 173–174) by the strongly elongate capitulum of the gonostyli (short in Sarimites ) and by the lateroventral processes of the aedeagus arising subapically (in middle portion of aedaegus in Sarimites ); (5) from Sinesarima (see illustrations in Chan & Yang 1994: figs 40–42) by the less elongate tegmina, about 2.2 × as long as wide (2.5 × in Sinesarima ), the more transverse vertex, about 2.2 × as wide as long in midline (1.7 × in Sinesarima ), by the strongly elongate capitulum of the gonostyli (short and somewhat foliate in Sinesarima ) and by the lateroventral processes of the aedeagus arising subapically (in middle portion of aedaegus in Sinesarima ); (6) from Yangissus (see illustrations in Chen et al. 2014: figs 2–82) by the less elongate tegmina, about 2.2 × as long as wide (2.9 × in Yangissus ), the more elongate anal tube, about 2.8 × as long as wide in dorsal view (2.4 × in Yangissus ), the strongly elongate capitulum of the gonostyli (short in Yangissus ) and the periandrium without processes (dorsal lobe of periandrium with elaborate processes in Yangissus ).

Etymology

The genus name is formed by the combination of ‘Bach Ma’, the national park where the genus was first discovered, and ‘ Sarima ’, the type genus of the tribe Sarimini . Gender feminine.

Description

Medium sized (around 5.0–6.0 mm), very convex, moderately elongate, rather robust-bodied.

COLOUR. Mostly brown with paler markings on frons, often with symmetrical, zigzaged markings of white wax on tegmina.

HEAD. Vertex distinctly broader than long in midline (about 2.2 ×), weakly concave with obsolete median carina; anterior margin forming a widely obtuse angle, posterior one rather deeply concave; all margins moderately carinate. Frons weakly convex, narrowly visible from above, nearly 1.2 × as wide as long in midline, smooth with distinct complete median carina, and obsolete peridiscal carina marked by paler colour; maximum breadth slightly under level of antennae; dorsal margin weakly concave. Anteroventral angle of genae slightly projecting anteriad. Ocelli present, under eye. Clypeus triangular, convex, smooth, not keeled or carinate. Labium with last segment longer than broad, shorter than penultimate. Antennae with scape short, ring-shaped, and bulbous pedicel.

THORAX. Pronotum subtriangular, projecting anteriorly in a blunt straight angle, about ⅔ as long in midline, as mesonotum; smooth with anterior margin carinate and pair of impressed points on each side of paler median line; lateral fields very narrow behind eyes; paranotal lobes without tubercles /carinae, and with posteroventral angle rounded. Mesonotum subtriangular with posterolateral margins slightly incurved, smooth, weakly convex with shallow depression before scutellum; incomplete, rather distinct, sublateral carinae.

TEGMINA. Distinctly convex, elongate, about 2.2 × as long as wide, with longitudinal veins elevated; costal margin forming a broadly rounded angle at basal 4 /5; apical margin rounded; distinct lateral hump including vein ScP+RA slightly before basal ⅓, hiding costal margin in dorsal aspect; costal margin hidden by vein RP in distal 2 /5, in dorsal view; rather narrow but distinct epipleuron; clavus closed, reaching 4 /5 of tegmen length. Venation: ScP+R rather short; ScP+RA rather short, curved towards RP but not fused, and not extending beyond midlength of tegmen; RP unforked, long and sinuate; first fork of MP around midlength of tegmen, MP1 with three terminales; first fork of CuA at about same level, slightly beyond halflength of tegmen; Pcu and A1 fused at about ⅔ of clavus length, Pcu+A1 reaching apex of clavus; cross-veins more numerous and more strongly marked along costal margin and in distal half of tegmen.

HIND WINGS. Well developed, with three distinct lobes ( Sarimini type) more or less equal in width; mostly brown. Venation: ScP+R and CuA furcate; MP simple, sinuate; second branch of CuA fused distally with CuP; Pcu and A1 fused on basal half, Pcu unforked and A2 simple; one transverse vein between second branch of ScP+R and MP, and between MP and first branch of CuA.

LEGS. Moderately elongate and slender, with pro- and mesofemora and pro- and mesotibiae slightly flattened dorsoventrally, tibiae more slender than corresponding femora; posteroventral margin of pro- and mesofemora with row of minute teeth in distal portion; pro- and mesotarsi rather elongate. Metatibiae with two lateral spines in distal half and seven apical spines. Metatarsi rather short with first segment about as long as combined length of remaining segments. First metatarsomere with two latero-apical and 7 intermediate spines arranged in arc. Metatibiotarsal formula: (2) 7/ 9/ 2.

Male terminalia

Pygofer short, about 2.4 × as high as long at midheight in lateral view, with posterior margin broadly rounded in lateral view; in caudal view suboval, 1.3 × as high as wide. Gonostyli massive, moderately convex, with posterior margin roundly projecting caudad in lateral view, and anterodorsal margin distinctly rounded; capitulum elongate, strongly projecting dorsad and with poorly distinct neck, curved anterodorsad and evenly tapering towards apex in lateral view, with basilateral laminate process directed lateroventrad in caudal view. Anal tube elongate, dorsoventrally flattened, and sublanceolate, rather narrow, about 2.8 × as long as wide in dorsal view and with anal opening in basal ¼; in lateral view, weakly downcurved. Aedeagus symmetrical, curved posterodorsad in lateral view. Ventral lobe of periandrium laminate, spatulate. Dorsal lobe of periandrium wide, abruptly tapering in distal portion towards more or less truncate apex, and expanded lateroventrally in basal portion, forming lamina sometimes more or less covering distal portion of lateroventral processes of aedeagus; laterodorsal processes of periandrium arising ventrally from basal portion of dorsal lobe, pointed and curved towards the posterior and with lateral tooth. Aedeagus (sensu stricto) surpassing dorsal and ventral lobes of periandrium, bifid, and with robust lateroventral process arising subapically and curved ventrocephalad. Connective well developed, corpus connective long, regularly curved in lateral view, tectiductus well developed, conical with anteroventral apodemes and wide anterior foramen.

Distribution

Vietnam: Thừa Thiên-Huế and Quang Tri provinces.

Species included

Bachmarima expansa gen. et sp. nov.

Bachmarima recta gen. et sp. nov.

Bachmarima valida gen. et sp. nov.