Civetissus, Constant & Pham, 2025

Constant, Jérôme & Pham, Hong Thai, 2025, Issid planthoppers from Bach Ma and Phong Dien in Central Vietnam. (III) Tribe Sarimini (Hemiptera: Fulgoromorpha: Issidae), European Journal of Taxonomy 1025, pp. 1-109 : 48-50

Genus Civetissus gen. nov.

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Type species

Civetissus pagumoides gen. et sp. nov. by present designation.

Diagnosis

The genus Civetissus gen. nov. can be differentiated from the other genera of Sarimini by the following combination of characters: (1) the head strongly projecting anteriorly in a pointed process, with vertex longer than wide; (2) the frons about as long in midline, as wide, without median and peridiscal carinae; (3) the tegmina elongate, about 2.4 × as long as wide, with distinct lateral hump slightly around basal ¼ hiding costal margin in dorsal aspect, and with distinct epipleuron; (4) the vein ScP of the tegmen long, weakly curved and reaching margin of tegmen around distal 1 /5 of tegmen length; (5) the first fork of MP at about 2 / 5 of tegmen length, more basal than the first fork of CuA at about 3 / 5; M1 forked at about ¾ of tegmen; (6) the anal tube elongate, dorsoventrally flattened, widening from base towards anal opening, and tapering in distal portion in dorsal view; (7) the gonostyli with anterodorsal margin angularly rounded, and with capitulum rather short and curved anterodorsad; (8) the aedeagus (sensu stricto) with a single pair of elongate, shaft-like, lateroventral processes arising at about distal ⅓, and the laterodorsal process of the periandrium bearing an elongate shaft directed cephalad.

Differential diagnosis

The most similar genus is Pseudocoruncanius Meng, Qin & Wang, 2020 . However, Civetissus gen. nov. can be separated by its head strongly projecting anteriorly in a pointed process, with vertex longer than wide, and frons about as long in midline, as wide, while Pseudocoruncanius shows a broadly rounded anterior margin of the vertex/ dorsal margin of frons, a strongly transverse vertex, about twice as wide as long in midline, and a strongly transverse frons, about 1.5 × as wide as long in midline (see illustrations in Constant & Pham 2024a: fig. 35); the peridiscal carina of the frons is weak but distinct in Pseudocoruncanius , while it is obsolete in Civetissus , and on the tegmen, the fork of MP1 is very distal in Pseudocoruncanius ( 5 / 6 of tegmen length), while it is distinctly more basal in Civetissus (¾ of tegmen length).

Etymology

The genus name is formed by the combination of ‘civette’ (French) meaning ‘civet’, and Issus Brullé, 1832 , the type genus of the family Issidae . Several names were derived from ‘civette’ like the genus Civettictis Pocock, 1915 or the species C. civetta (Schreber, 1776) , as well the the common name ‘civet’, used for the members of the family Viverridae ( Mammalia, Carnivora ). The new genus name refers to the pointed head and markings which are reminiscent of some members of the family Viverridae ; the gender is masculine.

Description

Rather large ( 6.3–7.5 mm), very convex, rather elongate with pointed head.

COLOUR. Mostly brown or reddish brown with paler markings on head and thorax, and black rings on pro- and mesotibiae.

HEAD. Vertex distinctly projecting anteriad, slightly longer in midline, than broad, more or less flat with obsolete median carina; anterior margin angular with narrowly rounded apex, posterior one distinctly concave; all margins weakly carinate. Frons convex, oblique in lateral view, narrowly visible from above, slightly wider than long in midline, smooth without median carina and with peridiscal carina obsolete; maximum breadth slightly above level of antennae; dorsal margin strongly projecting anteriad in subtriangular process. Anteroventral angle of genae distinctly projecting anteriad, anterior margin slightly carinate under eye; narrow groove between eye and margin stopping under ocellus. Clypeus triangular, convex, smooth, not keeled or carinate. Labium with last segment longer than broad, shorter than penultimate. Antennae with scape short, ring-shaped, and subcylindrical; barrel-shaped pedicel.

THORAX. Pronotum transverse with anterior margin rounded and posterior margin slightly bisinuate, about 0.55 × as long in midline, as mesonotum; smooth with anterior margin finely carinate and pair of impressed points on each side of median line; lateral fields very narrow behind eyes; paranotal lobes without tubercles/ carinae, and with posteroventral angle angularly rounded. Mesonotum subtriangular with posterolateral margins slightly incurved, smooth, weakly convex with shallow depression before scutellum; no distinct carinae.

TEGMINA. Convex, elongate, about 2.4 × as long as wide, with longitudinal veins elevated and cross-veins weakly elevated; costal margin forming a broadly rounded angle at basal 2 /5; apical margin obliquely rounded; distinct lateral hump including vein ScP+RA around basal ¼, hiding costal margin in dorsal aspect; costal margin more or less hidden by vein RP in distal ¾, in dorsal view; distinct epipleuron; clavus closed, reaching about ¾ of tegmen length. Venation: ScP+R rather short; ScP+RA long, reaching external margin of tegmen around distal 1 /5 of tegmen length; RP unforked, long and weakly curved; first fork of MP at about 2 /5 of tegmen length, more basal than the first fork of CuA at about 3 /5; M1 with two terminales, forked at about ¾ of tegmen; Pcu and A1 fused slightly beyond halflength of clavus, Pcu+A1 reaching apex of clavus; cross-veins more numerous and more strongly marked in distal half of tegmen.

HIND WINGS. Well developed, with three distinct lobes ( Sarimini type) more or less equal in width; mostly brown. Venation: ScP+R and CuA furcate; MP simple, sinuate; second branch of CuA fused distally with CuP; Pcu and A1 fused on basal half, Pcu unforked and A2 simple; one transverse vein between second branch of ScP+R and MP, and (more basally) between MP and first branch of CuA.

LEGS. Rather short, slender, with pro- and mesofemora and pro- and mesotibiae slightly flattened dorsoventrally, tibiae more slender than corresponding femora; posteroventral margin of pro- and mesofemora with row of minute teeth; pro- and mesotarsi elongate and slender. Metatibiae with two lateral spines in distal half and eight apical spines. Metatarsi moderately elongate with first segment about as long as combined length of remaining segments. First metatarsomere with two latero-apical and seven intermediate spines arranged in arc ventrally. Metatibiotarsal formula: (2) 8 / 9/ 2.

MALE TERMINALIA. Pygofer short, about 2.8 × as high as long at midheight in lateral view; in caudal view suboval, 1.4 × as high as wide. Gonostyli rather large, moderately convex, with posterior margin roundly projecting caudad in lateral view, and anterodorsal margin distinctly angularly rounded; capitulum rather short and curved anterodorsad in lateral view, curved mesad in caudal view, with distinct neck and lateral hook, distally laminate antero-posteriorly. Anal tube elongate, dorsoventrally flattened, widening from base to around anal opening and tapering in distal portion in dorsal view; in lateral view, slightly downcurved. Aedeagus elongate, symmetrical, moderately curved posterodorsad in lateral view; base of periandrium dorsally with laminate projection directed caudad. Ventral lobe of periandrium laminate, constricted in middle portion, and spatulate in distal portion. Dorsal lobe of periandrium elongate, notched apically and with strong median carina (fin-shaped); elongate shaft arising apically and strongly recurved cephalad in basal portion. Aedeagus (sensu stricto) bifid from basal portion and more or less truncate apically, with pair of elongate, shaft-like, lateroventral processes arising at about distal ⅔ of length. Connective well developed, corpus connective long, straight in lateral view, tectiductus well developed, conical with anteroventral apodemes and wide anterior foramen.

Distribution

Vietnam: Thừa Thiên-Huế Province.

Species included

Civetissus pagumoides gen. et sp. nov.