Helophorus, Fabricius, 1775

Bilton, David T., 2023, Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (2), pp. 249-264 : 262

publication ID

https://doi.org/ 10.37520/aemnp.2023.014

publication LSID

lsid:zoobank.org:pub:D982BDAC-CB44-41BE-A35C-92D630F96BF7

DOI

https://doi.org/10.5281/zenodo.10166075

persistent identifier

https://treatment.plazi.org/id/03F6746E-FFD4-B62F-FF45-F88BFA7FB431

treatment provided by

Felipe

scientific name

Helophorus
status

 

Key to the Afrotropical species of Helophorus View in CoL

1 Antennae 9-segmented. ............................................ 2

– Antennae 8-segmented. ............................................ 4

2 Pronotal internal and middle intervals smooth, granulation reduced to pores and setae over most of surface ( Figs 1D View Fig & 3C View Fig ); median lobe of aedeagus with struts as long as tube ( Fig. 3G View Fig ); habitus relatively robust ( Fig. 1D View Fig ), body length 3.45–4.0 mm. .......................... ................................... H. cooperi Orchymont, 1947 View in CoL

– Pronotal internal and middle intervals with evident granulation over most of surface ( Figs 1A & G View Fig , 3A & F View Fig ); median lobe of aedeagus with struts distinctly shorter than tube. ...................................................... 3

3 Granulation on pronotal internal and middle intervals weaker ( Figs 1A View Fig , 3A View Fig ); prontoal submedian groove more strongly angled medially ( Fig. 3A View Fig ); parameres relatively straight, with evenly pointed tips ( Figs 2A–B View Fig ); body size smaller, 2.60–3.50 mm; aedeagus smaller, 0.45 mm. ... H. aethiopicus Régimbart, 1907 View in CoL

– Granulation on pronotal internal and middle intervals stronger ( Figs 1G View Fig , 3F View Fig ); pronotal submedian groove more weakly angled medially ( Fig. 3F View Fig ); paramere tips more broadly rounded ( Fig. 3L View Fig ); body size larger, 3.15–3.85 mm; aedeagus larger, 0.55 mm. .................. ................................................ H. simiensis sp. nov.

4 Aedeagus very large, 0.78–0.85 mm, parameres with acuminate tips ( Fig. 2K View Fig ); pronotal intervals raised and granulate ( Figs 1F View Fig , 3E View Fig ); body robust ( Fig. 1F View Fig ) and relatively large, 3.25–4.55 mm; species from East African Rift. ................................ H. nyandaruaensis sp. nov.

– Aedeagus smaller, maximum length 0.66 mm, paramere tips various, but not strongly acuminate ( Figs 2C–F, H–J View Fig ); pronotal intervals flatter ( Figs 1B–C, E View Fig , 3B, E View Fig , 4C–F View Fig ). ....................................................................... 5

5 Pronotal submedian groove broader, granulation on middle and internal intervals weaker ( Figs 1B–C View Fig , 3B View Fig , 4C, E View Fig ); walls of pronotal grooves straighter and sharper ( Figs 4C, E View Fig ); punctures of elytral striae larger and intervals narrower and more arched apically ( Fig. 4G View Fig ); aedeagus smaller, 0.50–0.55 mm, parameres with relatively narrow apices ( Figs 2C–F View Fig ). ............................... ...................... H. aethiops J. Balfour-Browne, 1954 View in CoL

– Pronotal submedian groove narrower, granulation on middle and internal intervals stronger ( Figs 1E View Fig , 3D View Fig , 4D, F View Fig ); walls of pronotal grooves more rounded and indistinct ( Figs 4D, F View Fig ); punctures of elytral striae smaller and intervals broader and flatter apically ( Fig. 4H View Fig ); aedeagus larger, 0.58–0.66 mm, parameres usually with relatively broad apices ( Figs 2H–J View Fig ). ................ ...................................... H. brumopluvialis sp. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Helophoridae

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