Compsophorini HEINRICH

A. M, 2009, Illustrated key to the tribes of subfamilia Ichneumoninae and genera of the tribe Platylabini of world fauna (Hymenoptera, Ichneumonidae), Linzer biologische Beiträge 41 (2), pp. 1317-1608 : 1350-1351

Tribe Compsophorini HEINRICH

Compsophorina HEINRICH 1962 - Syn. Nearct. Ichn. Stenop.: 638.

Compsophorini HEINRICH 1968 - Ichn. Stenop. of Africa 2: 297-299.

Type genus: Compsophorus SAUSSURE.

Compsophorini : HEINRICH 1975 - Ann. Zool. Wars. 32 (20): 470-471.

D i s t r i b u t i o n: Ethiopian (mainly) and Oriental Regions.

I n t r o d u c t i o n:

The representatives of tribe are characterized by moderately long bristle-shaped flagellum of females, thick head with genae long from front and temples from above. Surface of head and clypeus form single undifferentiated surface, not divided by sutures or impressions, like Listrodromini . Structure of strong short and very broad mandibles, with strong, almost equal teeth, divided by broad and deep gape and absence of lunulae on tergites of abdomen are shown to similarity with them. They are differed from Listrodromini by smooth claws, quadrangular areolet, peculiar morphology of propodeum and semiamblypygous abdomen of females. Mesonotum is with different unusually rough rugosity. Propodeum is gradually roundly slanted from base to apex and area superomedia begins practically from postscutellum. Abdomen of overwhelming number of species is broadly oval with strongly sclerotized second and third tergites. Flagellum of males without tyloides. There are species from small to very big, some species rust-colored, majority with metallic-blue or metallic-green basic coloration, abdomen apically often with white spots.

M o r p h o l o g y:

F l a g e l l u m: Of females slender, bristle-shaped, not widened or slightly widened beyond middle, attenuated toward apex. Flagellum of males without tyloides, moderately nodose. Of females moderately long, shorter than the front wing.

H e a d: Stout with broad temples and long genae; fields of face and clypeus not differentiated, without sutures and impressions. Occiput deeply impressed to ocelli; temples long; clypeus without expressed lateral corners; mandibles massive, short, very broad with large practically equal teeth situated in the same plane and divided by broad gape; clypeal foveae presented by only just visible pore; labrum not protrude, or only just visible protrude from under clypeus; ocelli small.

T h o r a x: Collar of pronotum with tendency to form developed in different degree, directed backwards lamellae, having middle notch of different form and expressed in different degree; sometimes, pronotal ridge transversally crenulated. Mesonotum almost always rather flat, short, only just longer than width; notauli in form of broad crenulated impression, developed all round of mesonotum; middle lobe, sometimes medially in front depressed and thus bipartite; external lobes of mesonotum often carinated by carina, parallel to external margin of mesonotum at some distance from it; surface of mesonotum with specific sculpture, extremely sharply wrinkled; every lobe of mesonotum usually with smooth and shining longitudinal band; subalarum thin, sharp; speculum smooth shining; area of mesopleural fovea slightly impressed or wide or in form of longitudinal impression; sternauli absent; scutellum always laterally carinated and elevated above postscutellum, or convex, or pyramidal, of females as rule, less convex than of males, sometimes convex of females and pyramidal of associated males, sometimes with big apico-lateral projections (males Eccoptosagellus). Propodeum from lateral rather short, gradually slanting from base to apex; areas of propodeum more or less expressed, at least area superomedia and lateral carinae of vertical part distinct; basal furrow of propodeum narrow, if present; usually area superomedia in middle borders with postscutellum; spiracles large, slit-shaped. Surface of propodeum, usually roughly wrinkled, area superomedia, however, polished of most species; areae superoexternae often regularly longitudinally wrinkled.

L e g s: Moderately long, femora rather thin (slender); claws not pectinate; hind coxae of females, with tendency to develop of projections or carinae, varying in size and form at internal side of theirs apices; hind tarsi of one group of species thickened and shortened, with flattened from outside metatarsus.

W i n g s: Areolet mainly quadrangular or with very narrow base; nervulus postfurcal, rarely interstitial; radius distinctly sinuous; wings usually hyaline, often uniformly and deeply darkened, of some species with deeply darkened transversal bands.

A b d o m e n:Offemales,semiamblypygous, longish-oval to stumpy and broadly-oval (of Pyramidamblys almost parallel sided and amblypygous, apically blunted). Petiolus of first tergite thin, at base not flattened, gradually broadened to postpetiolus, last usually transversal, without distinct median field, usually laterally scatterly punctured and in middle more or less considerably striated or longitudinally-wrinkled. Gastrocoeli always deep and transverse, triangle, with distinct thyridia, interspace between them narrow, striated or with longitudinal wrinkles; second and third tergites strongly sclerotized and very roughly sculptured, separated from each other by constrictions; surface of second and third tergites strongly sclerotized and very roughly sculptured, striated or wrinklypunctured, subsequent tergites rather smooth.

C o l o r a t i o n: In the tribe dark metallic blue, or in some cases, metallic green basic color of abdomen predominate, usually in combination with rusty-red head or head and thorax and often with white anal marks; white rings on front tergites only in genera Compsophorus and Oxyjoppa; ferruginous basic color of abdomen, or some tergites, often ordinary of all tribe.