Dichotomius (Selenocopris) danieli Kohlmann and Solís, 1997

Valois, Marcely C., Silva, Fernando A. B. & Vaz-de-Mello, Fernando Z., 2022, A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae), Journal of Natural History 56 (1 - 4), pp. 119-147 : 131-132

Dichotomius (Selenocopris) danieli Kohlmann and Solís, 1997

( Figures 2 (a–f), 6))

Dichotomius danieli Kohlmann and Solís 1997: 345 , 362; Krajcik 2006: 53; 2012: 91; Solís and Kohlmann 2012: 6; Vaz-de-Mello and Nunes 2016: 299.

Diagnosis

Dichotomius danieli , D. rodrigoi and D. favi are the only known species of the globulus group recorded in Costa Rica. As pre-apical teeth are absent on the dorsal surface of paramera, D. danieli is suggested to be more closely related to D. favi than to D. rodrigoi . The following characters can be used to distinguish D. danieli from D. favi and other species in the globulus group: surface of head covered with dense ocellate punctures; pronotal punctures equally spaced from each other; anteromedian region of pronotum with a feeble central impression; elytral striae wide, crenulate, punctures almost as wide as striae; elytral interstriae opaque with indistinct smooth dots; paramera rectangular, apex rounded in dorsal view ( Figure 2 (d)).

Redescription (male)

Body length: 9.5 mm. Dorsum black, barely shiny ( Figure 2 (a)). Head: clypeal edge bidentate, teeth acute, separated by a deep V-shaped emargination. Clypeogenal junction rounded. Surface with different-sized ocellate punctures; punctures forming transverse rows at anterior clypeal region. Clypeogenal suture present. Cephalic process produced into a small central tubercle ( Figure 2 (a)). Antennal club with light brown tumescence. Thorax: pronotum approximately twice wider than long. Pronotal disc convex, anteromedian region with a shallow central excavation; surface with ocellate punctures as dense as those on anterior angles ( Figure 2 (a)). Surface of hypomera with sparse, coarse setiferous punctures; setae short, and usually absent in some punctures. Prosternum filled with ocellate punctures. Mesoventrite filled with ocellate punctures usually bearing short, orange setae. Metaventrite surface with ocellate punctures, except on disc; punctures on anterior region usually bearing short setae. Meso-metaventral suture distinct. Mid-longitudinal sulcus feebly marked posteriorly. Lateral lobes of metaventrite with ocellate punctures larger than those on anterior region. Elytra: slightly convex in lateral view. Humeral callus distinct. Striae thin, with well-defined punctures spaced by their diameter. Interstriae I–VII slight convex, barely shiny, with shagreened microsculpture; medial region with distinct smooth dots. Legs: protibiae with three distinct teeth externally. Protibial spur curved at apical fifth. Metatibial spur deeply emarginate apically. Ventral surface of the profemora covered with coarse setiferous punctures along its entire extension, setae short. Ventral surface of the mesofemora with ocellate punctures at apical half. Ventral surface of the metafemora with ocellate punctures concentrated on apical half; basal half with sparse, scattered punctures. Meso and metafemora bearing a thin longitudinal groove along the entire extension of posterior margin, wider on apical half. Abdomen: ventrites covered with ocellate punctures laterally, punctures medially disposed in a single row along anterior edge. Pygidium twice wider than long, disc strongly convex apically. Surface of pygidium with ocellate punctures concentrated along anterior half, and spaced by half their diameter. Apex of the pygidium margined. Tegmen: in dorsal view, paramera rectangular, one-third longer than its width medially, apex truncate; inner edges of paramera curved outward along its whole length ( Figure 2 (d)). Dorsal surface of paramera smooth ( Figure 2 (d)). Apical half noticeably narrower than base in lateral view ( Figure 2 (f)).Ventral sclerotised membrane between paramera thinner and curved at basal half; rounded laterally ( Figure 2 (e)). Female: differs from male by clypeal teeth acute and separated by a deep V-shaped emargination, and sixth abdominal ventrite 3 times longer than fifth.

Type material examined

Holotype of Dichotomius danieli ♂ [ MNCR]: ‘Est. Pitilla, 9 km S. Santa Cecilia, Prov./Guana, COSTA RICA, 700 m. Ago 1994 /C. Moranga, L N 330200_380200 # 3198’ // ‘ Dichotomius danieli /Kohlmann and Solís’/‘ COSTA RICA INBIO/[barcode] CRI002/029748’ // ‘HOLOTYPO/ Dichotomius danieli /Kohlmann y Solís’. Examined via photographs ( Figure 2 (c)).

Figures

Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels.