DRYOPIDAE

Hubbard, Todd, 2017, Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes, The Coleopterists Bulletin 71 (3), pp. 471-505 : 472-475

DRYOPIDAE

Helichus basalis LeConte. Appanoose : Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Clarke: South Whitebreast Cr., Weldon, TMM, 8/17/15; Davis: Fox R., Moulton, CTH, 7/17/06; Henry: Little Potomac Cr., Mt. Pleasant, CTH, 8/7/ 15; Jefferson: North Walnut Cr., Pleasant Plain, CTH, 8/14/15; Keokuk: Twin Grove Cr., Ollie, CTH, 6/2/15; Lee: Lick Cr., Shimek State Forest, CTH, 9/10/10; Lucas: Wolf Cr., Chariton, CTH, 8/ 10/05; Monroe: Sage Cr., Stephens Forest, CTH, 8/ 8/14; Poweshiek: Bear Cr., Brooklyn, CTH, 9/25/ 02; Ringgold: Middle Fk. Grand R., Mt. Ayr, KLS, 7/14/04; Taylor: Honey Cr., Bedford, KBK, 8/18/ 04; Van Buren: Coppers Cr., Keosauqua, CTH, 10/ 16/09; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Wayne: Wolf Cr., Millerton, CTH, 8/10/ 05. Distribution: Eastern USA, extending west to Iowa in the north and Texas in the south. Iowa forms the northwestern border in the USA with distribution restricted to southern Iowa. This species’ habitat preference is under rocky substrate at the edge of streams. In Iowa, H. basalis tends to be collected more frequently in smaller streams. New state record.

Helichus fastigiatus (Say) . Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Black Hawk: Black Hawk Cr., Popp County Access, CTH, 8/22/12; Cerro Gordo: Bailey Cr., Thornton, TMM, 8/30/07; Clayton: Volga R., Elkport, MWB, 9/15/08; Clinton: Brophy Cr., McCausland, CTH, 9/30/13; Davis: Soap Cr., Eldon, CTH, 10/4/10; Decatur: Little R., Leon, CTH, 9/23/02; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Harrison: Soldier R., Pisgah, KLS, 9/21/01; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/12/08; Ida: Battle Cr., Battle Cr., TFW, 9/11/02; Jefferson: North Walnut Cr., Pleasant Plain, CTH, 8/14/15; Johnson: Iowa City ( Wickham 1911) Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Lick Cr., Farmington, CTH, 7/14/03; Linn: Wapsipinicon R., Central City, MWB, 9/27/06; Lucas: Whitebreast Cr., Lacona, TMM, 8/5/09; Mahaska: Muchakinock Cr., Eddyville, MWB, 9/2/11; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Ringgold: Lotts Cr., Lamoni, CTH, 8/26/96; Van Buren: Chequest Cr., Pittsburg, CTH, 7/21/11; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05. Distribution: Eastern USA, extending west to Iowa in the north and Texas in the south. Iowa forms the northwestern border. Habitat preference is under rocky substrate in small to large streams.

Helichus lithophilus (Germar) . Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Upper Iowa R., Dorchester, MWB, 9/11/01; Appanoose: Cooper Cr., Centerville, CTH, 8/11/05; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: Wapsipinicon R., Dunkerton, CTH, 8/2/06; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: East Branch Wapsipinicon R., Tripoli, CTH, 8/27/97; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Butler: Coldwater Cr., Greene, CTH, 8/29/96; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/ 13/07; Cass: West Nodaway R., Grant, CTH, 8/ 19/98; Cedar: Cedar R., West Branch, CTH, 9/ 12/06; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 9/22/15; Chickasaw: Wapsipinicon R., Ionia, CTH, 10/ 12/00; Clarke: South Whitebreast Cr., Weldon, CTH, 9/28/95; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 10/9/10; Clayton: Volga R., Elkport, MWB, 9/15/08; Clinton: Brophy Cr., McCausland, CTH, 8/11/04; Crawford: Big Cr., Denison, TMM, 10/1/13; Dallas: Mosquito Cr., Panora, CTH, 7/15/96; Davis: Fox R., Moulton, CTH, 6/29/06; Decatur: Long Cr., Van Wert,JML, 10/9/01; Delaware: South Fk. Maquoketa R., Dundee, CTH, 7/12/05; Des Moines: Skunk R., Burlington, MWB, 9/7/06;

Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Fayette: Nutting Cr., Clermont, CTH, 7/26/06; Floyd: Cedar R., Charles City, MWB, 9/9/08; Franklin: Maynes Cr., Hampton, CTH, 8/18/08; Fremont: Nishnabotna R, Hamburg, TMM, 10/ 1/13; Greene: Buttrick Cr., Grand Junction, CTH, 8/30/95; Grundy: Middle Fk. South Beaver Cr., Wellsburg, CTH, 8/14/01; Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hamilton: White Fox Cr., Webster City, CTH, 8/22/95; Hardin: Iowa R., Eldora, CTH, 10/11/ 05; Harrison: Mosquito Cr., Persia, TFW, 10/12/ 00; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Riceville, CTH, 7/10/06; Humboldt: West Fk. Des Moines R, Humboldt, TMM, 8/2/12; Ida: Maple R., Galva, CTH, 8/6/ 97; Iowa: Big Bear Cr., Victor, CTH, 9/20/99; Jackson: Maquoketa R., Canton, CTH, 9/7/05; Jasper: Indian Cr., Mingo, TMM, 9/30/03; Jefferson: Cedar Cr., Stockport, CTH, 7/20/ 05; Johnson: Iowa R., Iowa City, CTH, 9/22/ 05; Jones: Maquoketa R., Monticello, CTH, 9/6/ 05; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: East Fk. Des Moines R., Seneca, TMM, 8/29/11; Lee: Skunk R., Augusta, CTH, 10/6/15; Linn: Buffalo Cr., Central City, CTH, 9/ 2/08; Louisa: Iowa R., Columbus Junction, MWB, 9/10/08; Lucas: Wolf Cr., Chariton, CTH, 8/10/05; Madison: North Branch North R., Winterset, CTH, 6/20/95; Mahaska: North Skunk R., Rose Hill, CTH, 7/10/96; Marion: North Cedar Cr., Bussey, CTH, 8/25/98; Marshall: South Minerva Cr., Clemons, TMM, 8/8/07; Mills: West Nishnabotna R., Malvern, TMM, 9/4/12; Mitchell: Wapsipinicon R., McIntyre, CTH, 8/27/08; Monona: Maple R., Mapleton, TMM, 9/5/12; Montgomery: Pilot Branch, Stennett, CTH, 9/9/96; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/14; Page: East Nodaway R., Hawleyville, CTH, 10/20/98; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/98; Pottawattamie: Keg Cr., McClelland, JML, 9/4/02; Poweshiek: Little Bear Cr., Brooklyn, CTH, 8/18/05; Ringgold: Lotts Cr., Lamoni, CTH, 8/26/96; Sac: North Raccoon R., Auburn, KBK, 9/13/05; Scott: Lost Cr., Princeton, CTH, 8/30/04; Shelby: East Branch West Nishnabotna R., Avoca, TMM, 8/ 16/07; Story: South Skunk R., Ames, CTH, 9/ 15/95; Tama: Richland Cr., Haven, CTH, 9/8/95; Taylor: West Branch 102 R., New Market, CTH, 10/10/95; Van Buren: Des Moines R., Farmington, CTH, 10/10/06; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Warren: North R., Norwalk, KLS, 10/8/04; Washington: English R., Riverside, MWB, 10/15/99; Wayne: Wolf Cr., Millerton, CTH, 8/10/05; Webster:

Lizard Cr., Clare, CTH, 9/11/96; Winneshiek: Yellow R., Postville, CTH, 7/25/07. Distribution: Eastern USA, extending west to Minnesota in the north, Nebraska, and Texas in the south. Habitat preference is woody debris and rocky substrate in small to large streams .

Helichus striatus LeConte. Adair : Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Little Paint Cr., Harpers Ferry, CTH, 9/11/07; Audobon: Unnamed trib. to East Nishnabotna R., Audobon, TMM, 8/23/11; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Boone: Unnamed trib. to Little Beaver Cr., Bouton, JML, 10/4/06; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/ 23/11; Buchanan: Lime Cr., Brandon, CTH, 5/ 15/96; Butler: Dry Cr., Clarksville, CTH, 5/29/ 07; Calhoun: Unnamed trib. to Purgatory Cr., Lohrville, KLS, 6/25/08; Cass: Sevenmile Cr., Grant, KLS, 7/26/06; Cedar: Sugar Cr., Tipton, CTH, 8/23/01; Cerro Gordo: Bailey Cr., Thornton, TMM, 8/30/07; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 5/15/12; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 8/26/14; Clarke: South Whitebreast Cr., Weldon, TMM, 9/29/10; Clay: Willow Cr., Royal, CTH, 9/13/99; Clayton: North Cedar Cr., Clayton, CTH, 8/22/94; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Crawford: Big Cr., Denison, CTH, 8/12/98; Dallas: Little Beaver Cr., Woodward, JML, 7/24/07; Decatur: Weldon R., Woodland, TMM, 8/8/13; Delaware: Fenchell Cr., Strawberry Point, CTH, 9/17/07; Dubuque: Hickory Cr., New Vienna, CTH, 8/11/ 99; Fayette: Nutting Cr., Clermont, CTH, 7/26/ 06; Floyd: Little Cedar R., Colwell, CTH, 8/20/ 02; Franklin: Bailey Cr., Sheffield, CTH, 7/24/ 03; Greene: Buttrick Cr., Grand Junction, TMM, 7/30/14; Guthrie: South Raccoon R., Guthrie Center, KLS, 6/12/03; Hamilton: Long Dick Cr., Ellsworth, TMM, 10/6/08; Hancock: East Branch Iowa R., Goodell, TMM, 10/7/10; Hardin: Tipton Cr., Buckeye, CTH, 7/30/99; Harrison: Potato Cr., Persia, KLS, 7/12/05; Howard: Crane Cr., Saratoga, CTH, 10/10/00; Ida: Maple R., Galva, CTH, 8/6/97; Jackson: Storybook Hollow, Bellevue, CTH, 8/29/07; Jasper: North Skunk R., Kellogg, CTH, 8/7/ 01; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Jones: Bear Cr., Wyoming, CTH, 7/13/ 04; Keokuk: Troublesome Cr., South English, CTH, 7/7/03; Kossuth: Black Cat Cr., Algona, MWB, 8/25/03; Linn: Wapsipinicon R., Central City, CTH, 9/27/06; Louisa: Honey Cr., Conesville, CTH, 9/2/98; Lucas: Wolf Cr., Chariton, TMM, 9/14/15; Madison: North Branch North

R., Winterset, CTH, 5/15/97; Mahaska: Buck

Cr., Barnes City, MWB, 8/21/00; Marion: Whitebreast Cr., Knoxville, TMM , 8/27/12; Marshall: South Minerva Cr. , Clemons, 8/8/ 07 ; Mills: Mud Cr., Hastings, JML, 8/2/05 ; Mitchell: Wapsipinicon R ., McIntyre , MWB, 8/27/08 ; Monona: Jordan Cr., Soldier, TMM , 7/ 5/06; Monroe: Sage Cr., Stephens Forest , CTH , 8/8/14; Montgomery: Walnut Cr., Red Oak, CTH , 10/14/96; Muscatine: Mad Cr., Muscatine, CTH , 6/30/04; O’ Brien: Waterman Cr., Sutherland, JML, 8/15/01 ; Page: East Nishnabotna R ., Shenandoah , TMM , 9/18/08; Plymouth: Perry Cr., Hinton, KLS, 7/20/04 ; Polk: Camp Cr., Mitchellville, TMM , 7/20/05; Pottawattamie: Mosquito Cr., Underwood, TMM , 8/17/04; Poweshiek: Walnut Cr., Belle Plaine, CTH , 9/ 17/08; Ringgold: Lotts Cr., Lamoni, TMM , 7/18/ 03; Scott: Lost Cr., Princeton, CTH , 8/30/04; Shelby: West Nishnabotna R ., Irwin , TMM , 8/ 25/03; Sioux: Dry Cr., Hawarden, JML, 7/12/06 ; Story: Ballard Cr., Cambridge, KJS, 7/18/07 ; Tama: Richland Cr., Haven, CTH , 9/8/95; Warren: North R ., Norwalk , KLS, 10/8/04 ; Washington: English R ., Riverside , CTH , 9/ 24/02; Wayne: Dick Cr., Corydon, TMM , 9/ 11/08; Webster: Allen Cr., Stratford, JML, 8/ 15/07 ; Winneshiek: East Pine Cr. , Burr Oak, MWB, 9/29/08 ; Woodbury: West Fk. Little Sioux R ., Bronson , JML, 8/9/05 ; Wright: Drainage Ditch #49, Eagle Grove, JML, 9/3/02. Distribution: Transcontinental in the USA. Habitat preference for H. striatus is on debris and under rocky substrate in small to large streams .