Cathoropsini, Marceniuk & Oliveira & Ferraris Jr, 2024

Marceniuk, Alexandre Pires, Oliveira, Claudio & Ferraris Jr, Carl J., 2024, A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data, Zoological Journal of the Linnean Society 200 (2), pp. 426-476 : 437

Cathoropsini tribe nov.

MP 64, BI 1, ML 85

( Figs 1–3 , 9 )

ZooBank registration: urn:lsid:zoobank.org:act:429CBD26-08F8-4207-BB19-13F4260D7CFB .

Type genus: Cathorops Jordan & Gilbert, 1883.

Diagnosis

Mesethmoid median portion very narrow (2,?> 0), state 1 in Notarius biffi , Notarius kessleri , and Notarius neogranatensis , and state 2 in Notarius cookei , Notarius grandicassis , and Notarius troschelii ; bony bridge formed by lateral ethmoid and frontal cylindrical and thin (16, 0> 1); otolith longer than wide (73, 0> 1); subvertebral process long and narrow (79, 0> 1); transcapular process forming a right angle in relation to body axis 86 (0> 1), state 0 in Notarius cookei and Notarius grandicassis ; exoccipital bony crest parallel to vertebral column and mesially folded (92, 1> 2), state 3 in Notarius .

Ambiguous optimization: Frontal mesial laminar projection present (23, 1> 0); posterior cranial fontanel formed by frontals and parieto-supraoccipital (25, 1> 0); pterotic mesial border with parieto-supraoccipital longer than anterior border with sphenotic (36, 0> 1), reversed in Notarius ; vomer lateral processes very narrow (53, 0> 1), reversed in Aspistor ; otolith anteromesial process inconspicuous (72, 1> 0), homoplastic in Galeichthys ; dorsal crest of urohyal projected anteriorly (156, 0> 1), reversed in Notarius rugispinis , Notarius planiceps , and Notarius phrygiatus ; 14 or fewer ribs (213, 1> 0), reversed in Notarius ; seventh vertebra free from ventral superficial ossification (214, 0> 1).

Included genera Notarius Gill, 1863

Cathorops Jordan & Gilbert, 1883.

Remarks

BI analyses and ML analyses did not recognize Aspistor as a valid genus as was reported by Marceniuk and Menezes (2007) and Marceniuk et al. (2012) on morphological evidence (see Notarius Remarks).

Figures

Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 9. Representative skulls, in dorsal view, and distribution of Cathoropsini tribe nov. A, Cathorops hypophthalmus, USNM 293275. B, Notarius grandicassis, MPEG 30647. * marine, ** brackish waters, *** freshwater.