Cinetodus Ogilby, 1898

Marceniuk, Alexandre Pires, Oliveira, Claudio & Ferraris Jr, Carl J., 2024, A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data, Zoological Journal of the Linnean Society 200 (2), pp. 426-476 : 466

Cinetodus Ogilby, 1898

( Figs 1–3 , 42 )

Type species: Arius froggatti Ramsay & Ogilby, 1886 .

Diagnosis

Mesethmoid median portion very narrow (2, 2> 0); extrascapular subtriangular (37, 1> 2); epioccipital contacting a small narrow area of diagonal crest associated with neural spine of fourth vertebra (44, 1> 0); vomer anterior margin very pronounced and acute (50, 0> 1); basioccipital lateral process absent with anterior and posterior portions equally developed (83, 1> 0); anteroventral portion of opercle subtrapezoidal, very long (127, 1> 0); metapterygoid one and one-half times longer than deep in perpendicular section (135, 1> 2); posterolateral processes of urohyal short (160, 1> 0); third basibranchial chalice shaped (168, 0> 1); upper (pharyngeal) tooth plate round, as wide as long (195, 1> 0); dorsal processes of upper (pharyngeal) tooth plate very short or absent (197, 1> 0); median crest associated with neural spine of fourth vertebra very high (202, 0> 1); Müllerian ramus robust (207, 0> 1); ventral surfaces of parapophyses of fifth and sixth vertebrae conspicuously concave (212, 0> 1); adipose fin long (222, 2> 1).

Ambiguousoptimization: Mesethmoidmedialnotchnarrowanddeep (1, 0> 1); lateral horn of lateral ethmoid compressed and spatulate (11, 0> 1); premaxilla very wide and short, as long as wide (120, 1> 3); posterior process of cleithrum very long (224, 0> 2).

Included species

Cinetodus froggatti Ramsay & Ogilby, 1886 .

Habitat and distribution: Fresh and brackish waters, southern New Guinea and northern Australia ( Fig. 38 ).

Remarks

The results of the total-evidence analysis support the recognition of Cinetodus as previously defined by morphological and molecular data ( Marceniuk and Menezes 2007, Betancur–R. 2009, Marceniuk et al. 2012), with a new hypothesis of relationships.

Figures

Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 38. Representative skulls, in dorsal view, and distribution of the subtribe Doiichthyina.A, Brustiarius nox, AUM 47488. B, Paracinetodus carinatus, AUM 47550.C, Cochlefelis spatula, AUM 50296. D, Nedystoma dayi, AUM 47500. E, Aceroichthys dioctes, AUM 47507. F, Pachyula crassilabris, AUM 47509. G, Pararius proximus, AUM 47486.H, Potamosilurus velutinus, AUM 47489.* marine, ** brackish waters, *** freshwater.

Figure 42. Cinetodus froggatti, USNM 217080, Papua New Guinea and northern Australia, type species of the genus.