Hemipimelodus Bleeker, 1857

Marceniuk, Alexandre Pires, Oliveira, Claudio & Ferraris Jr, Carl J., 2024, A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data, Zoological Journal of the Linnean Society 200 (2), pp. 426-476 : 458

Hemipimelodus Bleeker, 1857

( Figs 1–3 , 31 )

ZooBank registration: urn:lsid:zoobank.org:act:84D53B59-4ABB-4F76-B370-F9A510BB29F2 .

Type species: Pimelodus borneensis Bleeker, 1851 .

Diagnosis

Posterior branches of mesethmoid narrow (6, 0> 1); posterior branches of mesethmoid parallel along entire extension (7, 0> 1); fenestra delimited by lateral ethmoid and frontal very large (17, 1> 2); frontal mesial laminar projection absent (23, 1> 0); posterior cranial fontanel very wide and long (27, 1> 3); epiphyseal bar transversely elongate and longitudinally narrow (29, 0> 1); medial groove of cranium absent (30, 0> 1); fenestra delimited by parieto-supraoccipital, pterotic and sphenotic present (34, 0> 1); ventral crest of parieto-supraoccipital process well developed through entire extension of process (48, 0> 1); accessory tooth plates absent (58, 1> 0); optic foramen very reduced (67, 1> 2); ventral tip of subvertebral process spatulate (80, 2> 3); premaxilla narrow and very long, its length two to three times in width (120, 1> 2); ventral crest of hyomandibula absent (142, 0> 1); dorsal crest of urohyal projected anteriorly (156, 0> 1); anterior process of first hypobranchial inconspicuous (173, 1> 0); second hypobranchial transversely elongate, its mesial face acute (176, 0> 1); distal portion of uncinate process of third epibranchial acute (183, 1> 0); posterior margins of fourth epibranchial slightly convex, one-fourth as wide as long (187, 0> 1); median crest associated with neural spine of third vertebra very high (201, 0> 1); accessory crest connecting transverse and median crests associated with neural spine of fourth vertebra present (204, 0> 1); general aspect of superficial ventral ossification regularly arched (211, 0> 1); posterior process of cleithrum very short (224, 1> 0).

Ambiguous optimization: Frontal as main component of bony bridge formed by lateral ethmoid and frontal (15, 0> 2); third basibranchial hourglass shaped (168, 1> 0); transverse crest associated with neural spine of fourth vertebra low (200, 1> 0).

Included species

Hemipimelodus borneensis (Bleeker, 1851) .

Habitat and distribution: Freshwater, South and Southeast Asia ( Fig. 24 ).

Remarks

See Remarks of Cephalocassis .

Figures

Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support.

Figure 24. Representative skulls, in dorsal view, and distribution of the subtribe Ariina.A, Arius maculatus, AUM 44875. B, Cephalocassis melanochir, CAS 49426. C, Cryptarius truncatus, AUM 50243. D, Hexanematichthys sagor, AUM 50242. E, Kyataphisa nenga, AUM 46280. F, Osteogeneiosus militaris, AUM 44880. G, Pseudosciades sona, AUM 44861. * marine, ** brackish water, *** freshwater.

Figure 31. Hemipimelodus macrocephalus Bleeker, 1858, junior synonym of Hemipimelodus borneensis Bleeker, 1851, BMNH 1863.12.4.78, Sundaland, Malaysia to Vietnam, type species of the genus.