Fastosarion mcdonaldi Stanisic, 1993

Hyman, Isabel T. & Köhler, Frank, 2019, Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae), Contributions to Zoology 88 (4), pp. 351-451 : 430

Fastosarion mcdonaldi Stanisic, 1993

Figs. 18 , 19H , 20G , 22G–H , 28

Eungarion mcdonaldi Stanisic, 1993: 28 , figs. 1a–b, 2, 3a–f, 4a–c; Hyman & Ponder, 2010: 56, figs. 6L–M, 13K, 14E–F, 18C, 19C, 21C; Stanisic et al., 2010: 316–317, 333; Stanisic, 2018: fig. 4F.

Material examined

Types: Holotype: QM MO43309 ( Dalrymple Heights , Eungella National Park, 1000 m, ME QLD).

Paratypes: QM MO43310 .

Non-type material: See table 1 .

Diagnosis

External morphology: Shell (fig. 19H) small (11.3–11.6), orange amber, 3.2–3.5 whorls, flattened, thin, base uncalcified, membraneous. Shell sculpture very faint on protoconch, obsolete on teleoconch. Body (fig. 20G) 28–32 mm, reddish brown. Shell lappets and mantle lobes large with pigmented pustules, lappets joined on both sides by a narrow collar.

Genital anatomy: Genitalia (figs. 22G–H, 28) with medium length vagina, very swollen; bursa copulatrix moderately short, duct distinct, bursa elongate oval. Penis long, swollen proximally, internally with 2–4 longitudinal pilasters, internal wall pustulose; 60% of penis contained in penial tunica. Epiphallus approx. equal length to penis, enters penis through small rounded verge; epiphallic caecum of moderate length; flagellum short, slender. Spermatophore simple.

Remarks

Fastosarion mcdonaldi is restricted to the Clarke Ranges in Mideastern Qld, usually found above 900 m altitude (fig. 18). Several other semislugs are found in this region, including F. aquavitae , F. comerfordae , F. ephelis and F. tuljun . Fastosarion mcdonaldi can be distinguished from F. aquavitae , F. comerfordae and F. ephelis by its more flattened, reduced shell and smaller size, and from F. tuljun by its larger size, pustulose shell lappets and more reduced shell with a membraneous base.

Figures

FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun.

FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm.

FIGURE 20 Fastosarion live images (Clade B). A-D.Fastosarion alyssa, A:AM C.512537, Mt Etna Caves; B:AM C.512480, Mt Ossa; C:AM C.512579, The Hummock. D: NMV F230748,Brandy Creek (Type locality of F.peterbrocki; A.Moussalli, NMV). E. Fastosarion ephelis (J. Stanisic).F.Fastosarion hannianus (QM image).G. Fastosarion mcdonaldi, Mt Dalrymple (QM image). H. Fastosarion rowani, AM C.512541,Dan Dan NP.

FIGURE 22 Spermatophores of Clade B species. A-B. F.alyssa, MO77158.C-D.F.deensis, MO85834.E-F. F.hannianus, MO72277.G-H.F.mcdonaldi, MO50434.I-J. F.minerva, MO78358.

FIGURE 28 Genitalia of F.mcdonaldi, MO504354,Eungella NP. A. Reproductive system.Arrow shows the junction of the penis and epiphallus. B. Spermatophore.C.Penis interior. Scale bars 1 mm (B–C), 2 mm (A).