Phanaeus cambeforti Arnaud, 1982

Cárdenas-Bautista, Johann Stephens, Parada-Alfonso, Jenny Andrea & Carvajal‑Cogollo, Juan E., 2020, Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia, Check List 16 (4), pp. 821-839 : 826

Phanaeus cambeforti Arnaud, 1982

Materials examined. COLOMBIA • 1, 14.52 mm (mean); Meta, Villavicencio, Vereda Buenavista, “El Porvenir” Farm ; 04°08′41.97″N, 073°40′47.92″W; 1177 m a.s.l.; 12 Feb. 2019; Cárdenas Johann and Parada Jenny leg.; high secondary forest; Bvbt1; UPTC-In-00032 . • 1♂, 14.52 mm (mean); Meta, Villavicencio, Vereda El Carmen, “La Manigua” Farm ; 04°08′09.54″N, 073°40′16.92″W; 918 m a.s.l.; 27 Dec. 2018; Cárdenas Johann and Parada Jenny leg.; high secondary forest; Crbt8; UPTC-In-00033 . • 2, 14.52 mm (mean); Meta, Villavicencio, Vereda El Carmen, “La Loma” Farm ; 04°08′30.69″N, 073°40′00.06″W; 729 m a.s.l.; 15 Feb. 2019; Cárdenas Johann and Parada Jenny leg.; wooded pasture; Crpat1; UPTC-In-00034 .

Identification. This species can be distinguished from other Phanaeus species by the pronotal punctures, while these were not strong, they usually are distinct both on the sides and on the disc; the pronotal central micro spots were often difficult to detect (×40) and were absent in some specimens; the basal pronotal fossae was lacking or represented only by small punctures (×10); the dorsum was olive green or olive brown with metallic highlights on the pronotum and the pygidium was green ( Edmonds and Zidek 2012).

Distribution. For the Orinoco region there are records from 200–600 m a.s.l. ( Medina and Pulido 2009). In the study area, the species was found between 700–950 m a.s.l. Phanaeus cambeforti was rare species (58 individuals) and was collected both during the dry season and in the rainy season with a greater habitat preference for areas with vegetation cover such as high secondary forest (43 individuals), riparian forest relics (four individuals), unlike open areas such as wooded pasture where only 11 individuals were recorded.