Scutellera nepalensis amethystina ( Germar, 1839 ) Rédei & Tsai, 2022

Rédei, Dávid & Tsai, Jing-Fu, 2022, A revision of Scutellera (Hemiptera: Heteroptera: Scutelleridae), Zootaxa 5092 (1), pp. 1-40 : 15-16

Scutellera nepalensis amethystina ( Germar, 1839) , new status

( Figs. 11–13, 15, 16 )

Cimex amethystinus Lichtenstein, 1796: 103 . For unnamed species in Stoll (1787: 138, pl. XXXVI fig. 251). Syntype (s): unknown locality; ZMUH? Unavailable, published in a work placed on the Official Index of Rejected and Invalid Works in Zoological Nomenclature (Opinion 1820) ( ICZN 1995).

Calliphara amethystina Germar, 1839: 124 . Syntype (s): [ Indonesia:] Java; ZMHB! Junior secondary homonym and junior subjective synonym of Cimex amethystinus Lichtenstein, 1796 . Synonymized with fasciata by Dallas (1851: 19).

Callidea lanius Stål, 1854: 231 . Syntype (s): [ Indonesia:] Java; NHRS! Synonymized with S. amethystina ( Germar, 1839) by Vollenhoven (1863: 12). Confirmed subjective synonym.

Scutellera pilosa: Vollenhoven (1863: 13) . Unavailable name ( ICZN 1999, Art. 11.6).

Philia amethystina: Schiødte (1843: 286) (diagnostic characters).

Calliphara amethystina: Herrich-Schäffer (1853: 30) (listed, distribution).

Scutellera amethystina: Vollenhoven (1863: 12) (redescription, records, distribution, variability), Mayr (1866: 22) (record), ( Walker 1868: 507) (listed, distribution).

Scutellera lanius: Stål (1856: 51) (redescription, distribution), Dohrn (1859: 2) (catalogue, distribution), Walker (1867: 15) (records).

Scutellera fasciata var. lanius: Stål (1873: 14) (diagnosis, distribution) Lethierry & Severin (1893: 21) (catalogue), Breddin (1901: 9) (distribution), Schouteden (1904: 23) (catalogue).

Scutellera amethystina var. lanius: Kirkaldy (1909: 303) (catalogue).

Scutellera pilosa: Krikken et al. (1981: 252) (listed).

Diagnosis. Differs from the nominotypical subspecies in the purple ground colour of the body.

Distribution. This subspecies is widely distributed in the Malay Archipelago, and extends to Wallacea (Lombok Is., Sulawesi, Ambon Is.; a specimen from an unspecified locality of the Sunda Islands was seen as well) ( Fig. 72 ). The record from Malacca ( Walker 1867) might pertain to this subspecies or it is possibly erroneous.

INDONESIA. Sumatra!; Borneo: Pelaihari !; Java!; Bali Is.!; Lombok Is.!; Sulawesi!; Ambon Is.!; Sunda Islands : unspecified locality !

Type material examined. Calliphara amethystina Germar, 1839 . Lectotype: ♂, “2”, “ Java Hag” [hw, badly darkened], “Var \ villosa Hagenb” [hw, badly darkened], “ pilosa De Haan \ Java De Haan” [hw yellow sq], “ Java \ Hagenb.” [yellow, hw], “amethystina \ Stoll. 251” [hw], “ Scutellera \ nobilis F. \ var.” [hw]; pinned, distiflagelli of both antennae and right mid leg lacking ( ZMHB) ( Figs. 11–14 ) . Paralectotypes (1 ♂ 2 ♀♀): “Cat. No 2” [hw], “ Java \ Hagenb.” [yellow, hw], “ Scutellera \ nobilis F. \ var.” [hw] (1 ♂ 2 ♀♀ ZMHB) .

Callidea lanius Stål, 1854 . Lectotype (present designation): ♀, “ Java ” [hw], “lanius \ Stål.” [hw], “Type.”, “ Typus ” [red, with pr black frame], “NHRS-GULI \ 000044793”; pinned, right distiflagellum, right fore tarsus, left mid leg, left hind tarsus lacking ( NHRS) ( Figs. 15–17 ) . Paralectotype: ♀, “ Java ”, “ Stål ”, “Type.”, “ Paratypus ” [red, with pr black frame], “NHRS-GULI \ 000044794”; pinned, left distiflagellum, right flagellum, right fore and mid legs, and tibia and tarsus of right hind leg lacking ( NHRS) .

Additional specimens examined. INDONESIA. Sumatra: Deli [Serdang], leg. H. Fruhstorfer (1 ♂ NHMW) , same but leg. L. Martin (1 ♀ ZMHB) , Tandjong [= Tanjung] Morawa, leg. B. Hagen (1 ♀ RMNH) , Sumatra, coll. Noualhier 1898 (1 ♀ MNHN) , Kepahiang , v. Lansberg (1 ♂ RMNH) ; Borneo: Pleihari [= Pelaihari], 1866, leg. Semmelink (1 ♀ RMNH) , Borneo (2 ♂♂ NHMW) , same, coll. Muller (1 ♀ RMNH) ; Java: Batavia [= Jakarta], coll. P. Serre 1904 (2 ♂♂ MNHN) , same locality, Novara Exp. (1 ♂ 2 ♀♀ NHMW) , same locality, 11.v.1881, leg. Moskowi (1 ♀ NHMW) , same locality, Jagor, 7100 (1 ♀ ZMHB) , Baai v. Batavia [= Jakarta Bay ], 1928, leg. W.C. v. Heurn (2 ♂♂ RMNH) , Buitenzorg [= Bogor], leg. A. Ward, B.M.1901-313 (1 ♀ BMNH) , same locality, leg. v. d. Hoeven (1 ♂ RMNH) , same locality, [18]98, leg. Fleischer (2 ♂♂ 1 ♀ ZMHB) , same locality, 1918, leg. W. Roepke, coll. D. MacGillavry (5 ♂♂ ZMAN > RMNH) , same locality, 16.viii.1926, on fruit of Phyllanthus acidus (1 ♀ RMNH) , same locality, 250 m, 17.iv.1923, No. 612, leg. L.G.E. Kalshoven (1 ♂ RMNH) , Boitenzorg [= Buitenzorg, = Bogor] (3 ♀♀ ZMHB) Sukabumi , 2000 ft., 1893, leg. H. Fruhstorfer, coll. Noualhier 1898 (1 ♀ MNHN) , Preanger [= Parahyangan], Regentschappen [= Regencies], 1899, coll. M. Bartels (1 ♂ 1 ♀ ZMAN > RMNH) , Preanger [= Parahyangan], Ardja-Sari [= Arjasari], coll. D. MacGillavry (1 ♀ ZMAN > RMNH) , Pengalengan , 4000 ft., 1893, leg. H. Fruhstorfer (1 ♀ NHMW) , Palabuan , coll. Noualhier 1898 (1 ♂ MNHN) , Noesa [= Nusa] Kambangan , x.1917, coll. D. MacGillavry (1 ♀ RMNH) , Kediri , coll. W.L. Distant, B.M.1911-383 (1 ♀ BMNH) , Malang (1 ♂ HNHM) , Java (3 ♀♀ BMNH, 1 ♂ 2 ♀♀ RMNH) , same, coll. Muller (1 ♂ 3 ♀♀ RMNH) , same but coll. v. Hasselt (1 ♂ RMNH) , same but coll. Hagenb [eck] (2 ♂♂ 2 ♀♀ ZMHB) , same but coll. Breddin (2 ♀♀ SDEI) , same but coll. Noualhier 1898 (1 ♂ 1 ♀ MNHN) , same but coll. Signoret (1 ♂ 1 ♀ NHMW) , same but coll. W.L. Distant, B.M.1911-383 (2 ♀♀ BMNH) , same but leg. Xántus (1 ♂ HNHM) , same locality, 1868, leg. Warsberg (1 ♀ NHMW) , same locality, ii.1917, coll. D. MacGillavry (1 ♂ ZMAN > RMNH) , same locality, 1924–1927, leg. W.C. v. Heurn (3 ♂♂ 6 ♀♀ RMNH) , same locality, No. 257, leg. L.G.E. Kalshoven (1 ♀ RMNH) ; Bali Is. : 24.ii.1998 (1 ♂ 1 ♀ NSMT) ; Lombok Is. : i.1988, native collector (1 ♀ NSMT) ; Sulawesi: Celebes [= Sulawesi] (1 ♀ BMNH) , same but leg. Wallace, coll. Saunders, 65-13 (1 ♀ BMNH) ; Ambon Is.: Amb [oina] [= Ambon Is.], leg. Wallace, coll. Saunders, 65-13 (1 ♂ BMNH) ; Sunda Islands : coll. Gedult [18]88 (1 ♀ NHMW) .

Figures

FIGURES 11–17. Lectotypes of Calliphara amethystina Germar, 1839 and Scutellera lanius Stål, 1854 and their labels. Fig. 11, C. amethystina, dorsal view; Fig. 12, same, lateral view; Fig. 13, same, ventral view; Fig. 14, same, labels; Fig. 15, S. lanius, dorsal view; Fig. 16, same, ventral view; Fig. 17, labels. Scales in mm. © ZMHB (Figs. 11–14), © NHRS, courtesy of G. Lindberg (Figs. 15–17).

FIGURES 71–72. Geographical distribution of Scutellera spp. Fig. 71: S. perplexa (Westwood, 1837). Fig. 72: Circles = S. nepalensis nepalensis (Westwood, 1837); squares = S. nepalensis amethystina (Germar, 1839); triangles = S. spilogastra (Walker, 1867). Solid markers represent specimens examined during the present study, empty markers unverified literature data.