Syzeuctus frontator, Riedel, 2021

Riedel, Matthias, 2021, The genus Syzeuctus FÖRSTER in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species, Linzer biologische Beiträge 52 (2), pp. 1013-1098 : 1040-1042

Syzeuctus frontator nov.sp. ( Figs 5 , 14 , 34, 70, 113)

H o l o t y p e: (♀) " Turkey, N of Sivas, 8.7.2000 leg. M. Halada" (Linz).

P a r a t y p e s: (2♀♀ ♂) " Syria occ. 95 Bloudan 16.5. K. Denes jun. lgt." (Linz); (♀) "TR, Sirkans, Oltu, Erzurum, 25.6.96" (Munich).

S t u d i e d m a t e r i a l: 4♀♀, 1♂; Syria * and Turkey *.

D e s c r i p t i o n: ♀: Body length 10-12 mm. Antenna with 43-45 flagellomeres; 1 st flagellomere 4.1-4.2× longer than wide and 0.56× as long as eye; preapical flagellomere 0.9× as long as wide. Temple strongly and roundly narrowed behind eye, dorsally c. 0.6× as long as eye. OED 1.8× and OOD 1.9× ocellar diameter. Vertex smooth, with very scattered punctures. Frons moderately impressed medially, with dense punctures. Face densely punctate, inner eye margins slightly convergent ventrally. Clypeus c. 1.6× wider than long, with very few setiferous punctures, apical margin rounded. MI 0.55-0.65. Mandible with two teeth, lower tooth smaller than upper one. Genal carina reaching mandibular base.

Mesoscutum and mesopleuron with rather dense punctures. Frontoventral edge of pronotum moderately curved; lateral pronotal groove smooth. Speculum partly smooth. Epicnemial carina low, with ± distinct dentiform lamella on transversal part. Propodeum coarsely and densely punctate, with strong apical transversal carina. Hind femur 5.9× longer than wide. Hind metatarsus 0.65× as long as hind femur. 2 nd hind tarsomere 1.4× longer than 3 rd one. Hind claw with long teeth. Areolet stalked, 2 nd recurrent vein in its middle. Nervulus postfurcal by 1-2× its width. Hairs of basal cell slightly shorter than or as long as their intervals. Hind wing with 9 distal hamuli.

Metasoma slender. 1 st tergite 2.0-2.1× longer than wide, almost smooth and strongly shining. 2 nd tergite 1.1-1.35× longer than wide, with fine very scattered punctures, shining. 3 rd tergite 1.05-1.3× longer than wide, with fine dense but superficial punctures, shining. Spiracles of 1 st to 3 rd tergites slightly protruding laterally. Ovipositor 1.1-1.2× longer than fore wing, 1.25-1.45× longer than metasoma, 2.2-2.7× longer than hind tibia.

Color: Black. Clypeus, sometimes narrow stripe facial orbit, strongly widened spot on frontal orbit, narrow stripe on outer orbit, wing base, spot on fore coxa and trochanter, sometimes spots on mid and hind trochanters and trochantelli yellow. Mesosoma black or (in Syrian specimens) with yellow spots on upper pronotal margin, anterolateral on mesoscutum, apical spot on scutellum and postscutellum and central spot on propodeum. Fore and mid legs yellow, sometimes fore femur with brown stripe dorsally. Hind femur black, with yellow apical margin or yellow dorsal stripe; hind tibia blackish internally, ± yellowish externally. Mid and hind tarsi ± blackish. Metasoma completely black or (in Syrian specimens) 1 st to 4 th tergites with narrow apcial yellow bands. Wings hyaline or with weak brown apical spot (in Syrian specimens), pterostigma dark brown.

♂: Body length 9 mm. Antenna with 43 flagellomeres; 1 st flagellomere 4.5× longer than wide and 0.67× as long as eye. Temple moderately and roundly narrowed behind eye, dorsally c. 0.6× as long as eye. OED 1.4× ocellar diameter. Face densely punctate, inner eye margins slightly convergent ventrally. Clypeus c. 1.6× wider than long, with very few setiferous punctures, apical margin rounded. MI 0.65. Genal carina reaching mandibular base.

Frontoventral edge of pronotum moderately curved; lateral pronotal groove smooth. Speculum partly smooth. Epicnemial carina low, with ± distinct dentiform lamella on transversal part. Propodeum with strong apical transversal carina. Hind femur 6.2× longer than wide. 2 nd hind tarsomere 1.4× longer than 3 rd one. Hind claw with long teeth. Areolet stalked; 2 nd recurrent vein in its middle. Nervulus postfurcal by 1-2× its width.

Metasoma slender. 1 st tergite 2.2× longer than wide, almost smooth and strongly shining. 2 nd tergite 1.35× longer than wide, with fine very scattered punctures, shining. 3 rd tergite 1.2× longer than wide, with fine superficial punctures, shining.

Color: Black. Mandible except teeth, clypeus, face, malar space, complete orbits (frontal orbit strongly widened) yellow. Mesosoma black; collare, side of pronotum largely, hook-shaped antero-lateral spots on mesoscutum (confluent centrally), scutellum except antero-median black spot, postscutellum, tegula, spot on subtegular ridge, large stripe on mesopleuron, two spots on metapleuron and large triangular spot on posterior half of propodeum yellow. Metasoma black; 1 st tergite subbasally and 1 st to 4 th tergites apically with narrow yellow bands. Fore and mid legs yellow; mid tarsus brown in distal 2/3. Hind leg black; hind coxa yellow dorsally; hind femur with dorsal yellow stripe; hind tibia yellow externally. Fore wing with weak brown apical spot, pterostigma dark brown.

R e m a r k: This new taxon differs from S. vigil (TOSQUINET) by: Epicnemial carina

with dentiform lamella on transversal part, basal tergites slenderer and with scattered fine punctures, hind femur slenderer.

Figures

Figs 1-6: Habitus of ♀♀ from dorsal: (1) Syzeuctus altaicus nov.sp. HT (holotype), (2) S. flagellator nov.sp. HT, (3) S. hyalinus nov.sp. HT, (4) S. flavator nov.sp. HT (♂), (5) S. frontator nov.sp. HT, (6) S. maroccanus nov.sp. HT.

Figs 12-15: Habitus of ♀♀ from lateral: (12) Syzeuctus altaicus nov.sp. HT, (13) S. flagellator nov.sp. HT, (14) S. frontator nov.sp. HT, (15) S. robustus TELENGA LT (lectotype).