Sclerophrys regularis (Reuss, 1833)

Segniagbeto, Gabriel Hoinsoudé, Ohler, Annemarie, Rödel, Mark-Oliver, Luiselli, Luca & Dubois, Alain, 2024, Amphibians of Togo: taxonomy, distribution and conservation status, Zoosystema 46 (25), pp. 631-670 : 639

Sclerophrys regularis (Reuss, 1833)

( Fig. 4B )

MATERIAL EXAMINED. — Togo • 2 ♂, 2 ♀; Lomé ; MNHN-RA-1995.7155, MNHN-RA-1995.7165-7167 • 3 ♀; Assoukoko ; MNHN-RA-1995.4399-4401 • 3 ♀; Kara; MNHN-RA-2006.2177, MNHN-RA-2006.2180, MNHN-RA-2006.2188 • 1 ♀; Kpalimé ; MNHN-RA-2006.2181 .

DESCRIPTION. — Large toad (SVL 76-78 mm ♂, 63-92 mm ♀), with robust body. Snout rounded. Head wider than long (HW 36- 42% SVL; HL 29-36% SVL). Tympanum distinct (TYD 7-8% SVL). Tibia short (TL 37-42% SVL). Webbing rudimentary, leaving four phalanges free; webbing formula: I 1 ½ – 2 ½ II 1 ½ – 3 III 2 – 4 IV 4 – 2 V. Skin with distinct warts on the dorsal parts (back, flanks, thighs, legs); belly granular. Parotoid glands large, flat, smooth, separated from the eyelids.

COLOURATION. — The dorsal colouration is gray, light brown, brown, with more or less dark spots. The flanks, tympanum and loreal region are brown, light brown or beige. The dorsal side of forearm, thigh, leg and feet is of light brown with darker stripes. Chest and belly are dark gray.

SEXUAL DIMORPHISM. — Males have a subgular vocal sac, indicated by a blackish colouration. It is often more robust and its forelegs are stronger than those of females. Males have well developed nuptial pads on their thumbs.

HABITATS AND DISTRIBUTION. — Sclerophrys regularis is the most common amphibian species in Togo, present in all regions, and widely distributed throughout Africa ( Frost 1985; Rödel 2000; Channing & Rödel 2019). Its presence in Togo has been reported by Bourgat (1979), Salami-Cadoux (1979), Pujol & Exbrayat (1987, 2002), Hillers et al. (2009) and Segniagbeto et al. (2022).

Figures

FIG. 4. — Representatives of anuran species from Togo in life: A, Sclerophrys maculata (Hallowell, 1854) (Kalaré, Fazao Malfakassa); B, Sclerophrys regularis (Reuss, 1833) (Adjarala); C, Sclerophrys togoensis (Ahl, 1924) (Diguendue); D, Conraua derooi Hulselmans, 1972 (Yikpa-Dzigbe); E, Hoplobatrachus occipitalis (Günther, 1859) (Akloa); F, Hemisus marmoratus (Peters, 1854) (Kpalimé).