Hylarana galamensis (Duméril & Bibron, 1841)

Segniagbeto, Gabriel Hoinsoudé, Ohler, Annemarie, Rödel, Mark-Oliver, Luiselli, Luca & Dubois, Alain, 2024, Amphibians of Togo: taxonomy, distribution and conservation status, Zoosystema 46 (25), pp. 631-670 : 660

Hylarana galamensis (Duméril & Bibron, 1841)

( Fig. 13D )

MATERIAL EXAMINED. — Togo • 12 ♂, 1 ♀; Kloto ; MNHN-RA-1979.7130-7142 • 1 ♂; Nangbéto ; MNHN-RA-2006.2161 • 1 ♂; N’Gambi ; MNHN-RA-2006.2162 • 6 ♂; Toglekope ; MNHN-RA-2006.2159-2164 .

DESCRIPTION. — Large frog (SVL 72-81 mm ♂, 63-75.5 mm ♀), with an elongate, robust body. Snout rounded. Head roughly as long as broad (HW 29-38% SVL; HL 33-45% SVL). Tympanum distinct (TYD 7-9% SVL). Hind legs moderately long (TL 40-50% SVL). Webbing small leaving three phalanges free; webbing formula: I 2 – 2 ½ II 1 – 3 III 1 – 3 IV 3 - 1 ½ V. Toe tips pointed. Skin of head and anterior part of back smooth; posterior part of the back and flanks with smooth glandular warts; chest and belly smooth.

COLOURATION. — The back and sides are olive-gray with creamy white dorsolateral folds. The white band on upper lips continues as a white glandular ridge ending in the inguinal region. The flanks show gray and whitish spots of irregular shapes. The ventral side is white.

SEXUAL DIMORPHISM. — The male of the species is distinguished by two vocal sacs on either side of the throat and the presence of the bulging humeral glands.

HABITATS AND DISTRIBUTION. — This species occurs in savannahs, but can also be found in the forest zone. It has been observed in all ecological zones of the country. It is more frequent in the Fazao and Bassar regions and along the Zio and Haho basins in the south. The presence of the species in Togo has been reported by Bourgat (1979: 604), in the monograph on biodiversity in Togo ( PNAE 2002), Rödel & Agyei (2003) and Leaché et al. (2006).

Figures

FIG. 13. — Representatives of anuran species from Togo in life: A, Ptychadena pumilio (Boulenger, 1920) (Mango); B, Ptychadena tellinii (Peracca, 1904) (Koui); C, Aubria subsigillata (Duméril,1856) (Kovié);D, Hylarana galamensis (Duméril & Bibron,1841) (Avévé);E, Hylarana parva (Griesbaum, Jongsma, Penner,Kouamé, Doumbia, Gonwouo, Hillers, Glos, Blackburn & RÖdel, 2023) (Diguengue); F, Geotrypetes seraphini (Duméril, 1859) (Agou).