Hyperolius guttulatus Günther, 1859

Segniagbeto, Gabriel Hoinsoudé, Ohler, Annemarie, Rödel, Mark-Oliver, Luiselli, Luca & Dubois, Alain, 2024, Amphibians of Togo: taxonomy, distribution and conservation status, Zoosystema 46 (25), pp. 631-670 : 646

Hyperolius guttulatus Günther, 1859

MATERIAL EXAMINED. — Togo • 10 ♂, 3 ♀; Kloto ; MNHN-RA-1993.6106, MNHN-RA-1993.6107, MNHN-RA-1993.6110-6118, MNHN-RA-1993.6108, MNHN-RA-1993.6109 .

DESCRIPTION. — Medium sized tree frog (larger than most other Togolese tree frogs) (SVL 32.3-37.2 mm ♂, 41.6-43.5 mm ♀), with moderately slender but fairly robust body, wider anteriorly than posteriorly. Head slightly longer than broad (HW 29-34% SVL, HL 33-36% SVL). Snout rounded.Tympanum distinct (TYD 2-5% SVL). Tibia short (TL 44-53% SVL). Webbing large, leaving up to two phalanges free; webbing formula: I ½ – 1 ½ II ½ – 1 ½ III ½ – 1 IV 1 ½ – ½ V. Skin smooth on the snout and strongly shagreen on the back and flanks; belly and chest granular; some warts at the corner of the mouth.

COLOURATION. — The dorsal colouration of females is extremely variable, but fades in alcohol. Males are mostly uniformly greenish. For colour descriptions and figures see SchiØtz (1963, 1967, 1999), Rödel (2000), Amiet (2012), Kouamé et al. (2015) and Channing & Rödel (2019). This species differs from other large and robust Hyperolius species by a dark canthal stripe.

SEXUAL DIMORPHISM. — Males have a subgular, bluish-green vocal sac with a roundish gland. Females are distinctly larger.

HABITATS AND DISTRIBUTION. — Hyperolius guttulatus is essentially found in swampy forests. Our localities represent the typical habitat of the species according to SchiØtz (1967), Amiet (1975, 2012) and Rödel (2000).